Printer Friendly

Select Coleoptera of the Ravenna Training and Logistics Site, Ohio, 1999 (1).

ABSTRACT. This is the first report of a three-year survey of the Coleoptera at the Ravenna Training and Logistics Site (RTLS) near Ravenna, OH. The goal of this study was to conduct a survey of the beetles at the RTLS in order to establish a benchmark of the species found at this site. This is the first study of its kind at the Training and Logistics Site. More than 11,000 specimens were collected during the activity period, April--November 1999. A total of 282 different species in 51 families were collected and identified. Collection methods included flight intercept traps, Lindgren[R] funnel traps, aerial traps, pit fall traps, aggregation pheromone traps, Japanese beetle traps, aquatic traps, beating sheets, UV light traps, hand collection, Berlese[R] funnels, and sweep nets. Several new state and regional records have been established.

INTRODUCTION

The goal of this survey was to collect and identify as many beetle species as possible within the range of habitats at the Ravenna Training and Logistics Site (RTLS) in Northeast Ohio (Fig. 1). A great deal of general and specific collecting at various habitats was done in order to get an idea of the species represented and their relative abundance. This benchmark data will be of interest to those studying invertebrates within the RTLS and also to others in Northeast Ohio and neighboring states where many of these species have not been previously reported. Several of the families, such as Carabidae, Nitidulidae, Scolytidae and aquatic beetle families, have not been identified and will be treated in subsequent publications.

[FIGURE 1 OMITTED]

AREA OF STUDY

The three major streams in the Training and Logistics Site are Hinkley Creek, Sand Creek, and South Fork of Eagle Creek The topography is an irregular plain and tableland with a gentle slope, lying on a glacial plain underlain by sandstone and siltstone. The two predominant forest types are beech maple and oak hickory forest (Tertuliani 1999). In ravines, the associated species are American basswood, American elm, red maple, eastern hemlock, white ash, black cherry, white pine, and northern red oak. In riparian areas, American elm, black ash, beech, and red maple are dominant species, and silver maple, pin oak, swamp white oak, sycamore, tupelo, and cottonwood are less abundant (Tertuliani 1999). Braun (1989) classified woody vegetation in this region as beech maple forest.

Four permanent collecting sites were chosen within the RTLS. These sites were sampled on a weekly basis covering various habitats and were spaced to access the four quadrants of the property. Collections were begun in early April and continued until there was little or no beetle activity in November.

Site 1 was located on the far west end of the RTLS in a level mesic woodland with semi-dense scrub under growth. This site consisted of a mixture of young to medium aged deciduous trees, providing a moderately open habitat.

Site 2 was near the headwaters of Hinkley Creek. The site was reached by descending a gentle slope forested primarily with medium to large beech, tulip, and maple trees. Here the canopy was continuous and the undergrowth sparse. During the spring the understory was carpeted with perennial wildflowers (jack-in-the-pulpit, violets, and trillium).

Site 3 was on a level forested area of mixed deciduous trees toward the eastern border of Portage and Trumbull counties. The undergrowth here was minimal due to the heavy forest canopy and deep leaf litter.

Site 4 was open, exposed, dry, and located near the southern boundary of the compound. This area had few mature trees, was composed of low scrub undergrowth, and provided a perfect habitat for beetles preferring open areas with abundant sunlight.

Table 1 lists locations of all sites where collections were made utilizing various sampling techniques such as beating sheets. Sweep-net collections were made along the side of the roads and in open fields. In addition, we set up pitfall traps and used hand nets, searched under logs and in leaf litter in areas other than the four permanent sites. Also, we received assistance from other researchers who collected both terrestrial and airborne species of beetles via various methods including light traps. These additional specimens greatly increased both the total number specimens collected and species diversity.

MATERIALS AND METHODS

The traps used at each of the primary sites were: 2-liter plastic bottles, window traps, pitfall traps, aerial pheromone traps, Lindgren funnel traps, and Japanese beetle traps.

Malt and brown sugar traps were made of 2-liter plastic bottles, suspended from a tree branch about 2.25 m above ground. They contained approximately 250 ml of solution with a 10 cm circular hole in opposite sides to allow beetle access. The malt recipe was 0.25 1 malt syrup per 0.75 1 of water, and the brown sugar solution was 0.5 kg brown sugar per 4.0 1 of water.

The window traps were single sheets of Plexiglas[R] 0.7 [m.sup.2] mounted on two 1.0 m long wooden boards driven into the ground. These were similar to the design of Peck and Davies (1980). These traps were equipped with a PVC[R] water trough on each side, and the troughs were filled about half full of soapy water.

Pitfall traps were baited with whole wheat bread dough wrapped in fiberglass netting inside a 0.5 1 glass Mason[R] jar with wire mesh lid. The aerial wheat dough traps were anchored to a 2.0 m wooden pole at the height of approximately 1.0 m (Williams and others 1995).

Aerial aggregation pheromone traps were suspended from tree branches. The pheromone-impregnated caps were hung on a string inside the Mason[R] jar with the same type of wire mesh lid. The aggregation pheromones were nitidulid attractants developed for several species of sap beetles.

Japanese beetle traps were hung on 2.0 m metal poles. Two Lindgren[R] funnel traps were used; one baited with a Chalcoprax[R] lure and the other with 3 exotic bark beetle lures. These traps were secured to 1.5 m metal poles. The lures in the 2 Lindgren[R] traps were replaced on June 29 and again on August 18.

An area we called the "stumps" was surveyed weekly. It was located near Site 1 at the western end of the RTLS. Recent logging left many exposed freshly cut maple and cherry stumps, which attracted sap beetles, rove beetles, and other ground dwelling beetles. In order to make the stumps more attractive we placed wooden wedges on top of them which provided a cover for the beetles and made it more likely they would stay in situ.

We also collected beetles from deer carcasses in the Training Grounds. Carrion beetles, nitidulids, histerids, clerids, and rove beetles were encountered in the carrion. By mid-June, scavenger activity made it necessary to wrap chicken wire around the aerial, malt, and brown sugar traps to keep unidentified animals (probably raccoons) from removing the bait.

During the summer months, Megan Michael processed several samples of leaf litter through Berlese[R] funnels at the Ohio Department of Natural Resources, Natural Areas and Preserves Division. These provided an additional source of terrestrial beetles.

Window traps were maintained until October 27. The Lindgren[R] traps were maintained until November 4, while the 2-liter plastic bottle traps were run until November 10. The aerial, ground, and aggregation pheromone traps were removed on November 17. The UV light traps used by Michael Gilligan were removed in the beginning of October.

The location of each specimen collected at RTLS was designated by site number, date, and coordinates, along with the collection method. Most specimens were pinned or placed on insect points. Labels with all pertinent data were placed with the specimens and separate determination labels were added once the specimens were identified. In most cases these determinations were to the species level.

RESULTS AND DISCUSSION

The 1999 collecting season was extremely dry resulting in several of the ponds drying up and apparent plant stress when compared to adjacent areas which had received more precipitation. Although we do not have data for comparison, we speculate that the effects of the drought reduced the numbers of beetles in some habitats, especially those living in the leaf litter, riparian, and woodland areas. However, the light traps yielded hundreds of beetle specimens from woodland and water habitats. At various times our collecting traps were invaded by other insects that were attracted to the baits. Ants, moths, yellow jackets, spiders, slugs, bald face hornets, and European hornets were among the unwelcome insects captured in our traps.

One of the benefits of this research project was the Ravenna complex itself. It is a unique place to do field work in a variety of habitats. We were so intrigued by the diversity of beetles found at the RTLS that we decided to design a poster featuring some of the beetles we collected for posterity and educational purposes. The poster will be one of the legacies of this project.

We did not find any federally listed endangered, threatened, or candidate species, or any state listed endangered, threatened, or potentially threatened species. However, we did find three new state records (NSR) all of which belonged to the Family Staphylinidae. Specialist Alfred Newton contributed the following comment on the staphylinid species Erichsonius cinerascens: "this species was earlier recorded from North America, then thought to not occur here in the most recent revision of the genus, but this specimen is definitely cinerascens and I think the species is actually widespread in North America but confused with another species."

Abundance ratings were used to describe a particular species population as follows beginning with the highest populations: A = abundant, C = common, U = uncommon, R = rare, and SI = special interest. The abundance ratings were solicited from specialists in the different families. This ranking is very subjective and should not be given much weight. For the most part the specialists were familiar with the relative abundance of their group of interest in the Midwest. The insects in Table 2 represent the majority of beetles collected in our 1999 studies.
TABLE 1
Sampling sites in the Ravenna Arsenal, 1999.

 Township Site-Location Latitude Longitude

1 Charlestown Site 1 S of Newton Falls 41 11 20.3 81 11 03.8
 Rd., opposite Franks
 Pond.

2 Charlestown Site 2 Near W end of 41 11 46.1 81 06 03.1
 Magazine Rd. where
 Hinkley Creek crosses.

3 Windham Site 3 About 100m N of 41 12 59.4 81 01 17.7
 Ramsdell Rd. on W side
 of Snow Rd.

4 Paris Site 4 S of Newton Falls 41 11 22.2 81 03 16.6
 Rd. at Maintenance area
 near Paris-Windham

5 Charlestown Area on E. side of small 41 11 12.9 81 05 57.8
 road off of Fuze and
 Booster Spur Rd.

6 Braceville Pine stand just N of 41 12 12.1 81 00 14.2
 Butts Kistler Rd. at
 Portage Co. line.

7 Charlestown Hinkley Creek at S 41 09 46.7 81 07 58.7
 Boundary Rd.

8 Charlestown McCormick Rd 1/2 way 41 10 37.4 81 09 07.7
 between Rt. 80 and
 B Block.

9 Charlestown Area near bridge over 41 11 30.5 81 07 20.9
 Sand Cr. on Newton
 Falls Rd., W of
 Slagle Rd.

10 Charlestown Quarry pond in block A 41 10 54.3 81 10 34.7
 just N of McCormick Rd.
 on Rd. 5A.

11 Charlestown Rt. 80 Hatchery pond 161 41 11 35.7 81 08 54.9
 to N of Newton Falls Rd.
 on W side of Rt. 80.

12 Charlestown Rt. 80 Trout Pond 322 m 41 11 33.2 81 08 31.7
 N of Newton Falls Rd.,
 E of Rt. 80.

13 Paris South Service Rd., S of 41 11 35.6 81 00 23.8
 Criggy's Pond, near SE
 corner of arsenal.

14 Charlestown Near trib. to W Br. 41 10 30.2 81 09 37.1
 Mahoning R. at SW corner
 of arsenal toward oil
 pumps.

15 Paris S Patrol Rd. near 41 11 43.1 81 00 33.3
 intersection with B&O
 NYE Rd. at E end of
 Portage Co.

16 Charlestown Area off of Rt. 80 near 41 10 11.1 81 08 49.4
 Gate 22 just N of S
 Patrol Rd.

17 Charlestown Trib. to Hinkley Cr. just 41 11 34.9 81 08 50.0
 S of Hatchery Ponds on
 Rt. 80

18 Charlestown W Patrol Rd. 1.6 km S of 41 11 05.9 81 11 32.9
 the NW corner of the
 arsenal.

19 Paris Big Cobb's Pond off of 41 11 55.9 81 02 51.9
 Paris Windham Rd., 644 m
 S of Remalia Rd.

20 Windham Beaver flooding Trib. to 41 12 30.9 81 02 55.6
 Sand Creek on Paris Wind.
 Rd. 200m N of Remalia

21 Paris E of S Service Rd 762 m 41 11 44.0 81 00 10.3
 S of Criggy's Pond, E
 border of Portage Co.

22 Windham Ed's Pond 322 m W of 41 13 09.1 81 03 22.1
 Paris Windham Rd., 250 m
 N of Smalley Rd.

23 Windham N Patrol Rd. Pond 161 m 41 13 28.5 81 05 42.9
 S of N Patrol Rd. on road
 leading into E Block.

24 Paris Pine stand on S Service 41 10 21.5 81 04 38.9
 Rd opposite of S Service
 Rd. Pond.

25 Paris S Service Rd. Pond, 41 10 25.4 81 04 45.0
 halfway between George
 Rd. and Wilcox Wayland
 Rd.

26 Paris Sand Creek near mouth, 41 13 34.7 81 02 06.1
 halfway between Smalley
 & N Patrol Rd.

27 Windham Sand Creek at Smalley Rd. 41 13 14.3 81 02 24.5
 bridge.

28 Windham Boy Scout Pond 41 13 37.0 81 04 44.0
 impoundment on S Fk.
 Eagle Creek off of
 Wadsworth Rd.

29 Windham S Fk. Eagle Cr. at wooden 41 13 34.9 81 02 49.0
 bridge on N Patrol Rd.

30 Charlestown Big beaver swamp on W end 41 12 01.7 81 09 01.1
 of Bundling Rd.

31 Charlestown Franks Pond, on N side of 41 11 32.7 81 11 03.5
 Newton Falls Rd. E. of
 W boundary.

32 Paris Pine stand on S side of 41 11 23.9 81 04 15.5
 Newton Falls Rd. 161 m
 from Wilcox-Wayland Rd.

33 Charlestown S side of Northline Rd. 41 12 16.6 81 08 05.1
 near beaver impoundment
 & heron rookery.

34 Charlestown S Patrol Rd where trib. 41 11 05.7 81 10 04.9
 to Hinkley Creek crosses,
 near Demolition pond
 area.

35 Paris Pauls Ponds area, 250 m 41 11 05.6 81 03 49.0
 S of Newton Falls Rd.

36 Paris Remalia Rd. near Load 41 12 09.6 81 02 14.0
 Line 2, just before
 entrance to Load
 Line 2 Rd.

37 Paris George Rd pond just S of 41 11 23.0 81 05 01.0
 intersection with Newton
 Falls Rd.

38 Charlestown Rt. 80 & N Patrol Rd. 41 12 18.6 81 08 52.2

39 Paris Fence behind garage 41 10 15.0 81 05 29.2
 building N of entrance
 gate, W side of George Rd

40 Braceville Woods in UTES property. 41 12 39.6 80 59 46.0

41 Charlestown Sand Creek @ Newton Falls 41 11 31.2 81 06 19.0
 Rd., E of Greenleaf Rd.

42 Freedom Woods on S side of 41 12 35.9 81 06 59.6
 intersection of Nonhline
 & Slagle Rd.

43 Charlestown Halfway down E. side of 41 10 17.5 81 07 06.4
 Greenleaf Rd. at Hinkley
 Crk trib. crossing.

44 Windham Near 90[infinity] bend in 41 12 39.6 81 02 36.6
 Winklepeck Rd & Sand
 Creek, NE of Wilcox-
 Wayland Rd.

45 Paris Powerline cut B&O Y area 41 11.728 81 05 36
 N of S Patrol, E of RR
 tracks

46 Charlestown S Patrol Rd E of Rt 80 41 10 01 81 09 16
 near exterior fenceline

47 Windham Boy Scout Camp W of 41 13 37 81 04 40
 Wadsworth Rd, S of N
 Patrol, walnut stand

48 Windham N Patrol Rd woods near 41 13 32 81 05 47
 V Patrol Rd pond

49 Charlestown W of Fuze/Booster, 122 m 41 11 28 81 5.80
 behind 1st bldg S of
 junction with Demolition
 Spur

50 Freedom W end of wood lot, 41 11.80 81 6.81
 Block D

51 Paris Guard property E side, 45 62 426 50 11 55
 woods, near bridge over
 trib. to Mahoning River

52 Charlestown N of S Patrol near gate 41 10.44 81 8.58
 22 W side Arsenal off
 of Rt. 80

53 Windham Trib to S. fork Eagle 41 12 33 81 06 59
 Creek at Slagle Rd.
 bridge S. of
 Northline Rd.

54 Charlestown SW corner S Patrol Rd on 41 10.25 81 11.11
 dirt rd leading to oil
 pumps

55 Charlestown 457.2 m W of Rt 80 on S 41 10.03 81 9.24
 Patrol Rd stream

56 Windham E of stone arch bridge 41 13.37 81 5.16
 crossing S Fork Eagle
 Creek on Wadsworth Rd

57 Windham Woods adjacent to ammo 41 12 37 81 02 09
 sectioning area off
 Rematia W of Randall Rd

58 Paris Newton Falls Rd. 0.8km W 41 11 27 81 04 57
 of Paris Windham Rd

59 Paris Woods on E side George 41 11 27 81 04 57
 Rd just S of Newton
 Falls Rd.

60 Charlestown Midway btwn Block B and 41 11.61 81 9.10
 Rt 80 on both sides of
 Newton Falls Rd

61 Charlestown W and behind building 41 19.34 81 15.78
 SM-7 on Magazine Rd

62 Charlestown 125 km N of S Patrol Rd 45 57.03 48 59.21
 on Knapp Rd

TABLE 2
List of the species of Coleoptera in the Ravenna Training
and Logistics Site 1999.

Taxon Total Range of Dates

Anthicidae
 Anthicus cervinus (La Ferte) 4 05-19 to 08-18
 Anthicus ephippium LaFerte 1 08-18
 Omonadus floralis (L.) 1 04-28

Biphyllidae
 Diplococelus brunneus LeConte 1 10-19

Buprestidae
 Chrysobothris rugosiceps Melsheimer 1 06-09
 Taphrocerus gracilis (Say) 1 06-09

Byturidae
 Byturus unicolor Say 1 05-12

Cantharidae
 Silis sp. 13 05-12 to 06-03

Cerambycidae
 Analeptura lineola (Say) 7 06-23 to 07-26
 Bellamira scalaris (Say) 1 06-23
 Brachysomida bivittata (Say) 5 05-19 to 06-09
 Dorcaschema alternatum (Say) 1 07-14
 Gaurotes cyanipennis (Say) 2 07-14
 Orthosoma brunneum (Forster) 17 04-21 to 07-15
 Parandra (Neandra) brunnea brunnea 2 08-04 to 08-25
 (F.)
 Stictoleptura canadensis Canadensis 1 07-27
 (Olivier)
 Strangalia luteicornis (F.) 6 6-29 to 07-27
 Tetraopes tetrophthalmus (Forster) 11 06-29 to 07-19
 Tylonotus bimaculatus (Haldeman) 1 07-27
 Typocerus velutinus velutinus 2 07-14 to 07-21
 (Olivier)

Cerylonidae
 Cerylon castaneum Say 7 05-05 to 06-09
 Philothermus glabriculus LeConte 1 04-21

Chrysomelidae
 Altica spp. 3 05-05 to 07-30
 Brachycoryna melsheimeri (Crotch) 1 05-19
 Brachypmoea margaretae (Schultz) 9 06-16 to 07-30
 Calligrapha multipunctata bigsbyana 1 06-16
 (Kirby)
 Chaetocnema fuscata White 6 04-21 to 05-19
 Chaetocnema pulicaria Melsheimer 1 04-28
 Chrysochus auratus (F.) 15 07-14 to 08-18
 Chrysolina auripennis (Say) 2 06-23 to 06-27
 Crepidodera browni Parry 1 04-28
 Diabrotica undecimpunctata howardi 11 07-19 to 11-10
 Barber
 Disonycha pensylvanica (Illiger) 8 08-24 to 10-04
 Donacia sp. 1 08-10
 Exema canadensis Pierce 1 not recorded
 Exema sp. 1 06-16
 Kuschelina miniata (F.) 1 04-28
 Longitarsus sp. 4 04-28 to 05-12
 Luperaltica senilis (Say) 3 08-17
 Microrhopala sp. 1 09-15
 Odontota sp. 20 06-09 to 08-10
 Ophraella conferta (LeConte) 3 06-16 to 06-29
 Ophraella cribrata (LeConte) 1 06-03
 Ophraella notata (F.) 1 05-05
 Pachybrachys sp. 1 08-03
 Paria thoracica (Melsheimer) 8 06-29 to 08-17
 Phyllotreta sp. 2 04-28 to 05-19
 Plateumaris shoemakeri (Schaeffer) 1 05-19
 Rhabdopterus praetextus (Say) 3 07-07 to 07-21
 Scelodyperus sp.nr liriophilus Wilcox 1 06-16
 Strabala rufa (Illiger) 1 05-19
 Systena blanda Melsheimer 1 10-19
 Tricholochmaea sp. 5 05-19 to 08-18
 Tymnes tricolor (F.) 1 07-07

Ciidae
 Cis levetti (Casey) 13 04-07 to 08-04
 Orthocis punctatus (Mellie) 3 05-12 to 06-09

Cleridae
 Enoclerus nigripes (Say) 14 09-15 to 11-04
 Necrobia rufipes DeGeer 9 06-29
 Necrobia violacea L. 20 04-14 to 06-29
 Placopterus thoracicus (Olivier) 1 06-03
 Thanasimus dubius (F.) 3 05-05 to 05-19
 Zenodosus sanguineus (Say) 3 05-05 to 05-19

Coccinellidae
 Brachiacantha rotunda Gordon 3 06-16 to 07-14
 Coccinella septempunctata L. 1 05-19
 Coleomegilla maculata lengi Timberlake 4 05-19 to 08-03
 Cycloneda munda (Say) 6 04-28 to 10-01
 Harmonia axyridis (Pallas) 7 04-28 to 10-19
 Hippodamia parenthesis (Say) 3 04-14 to 08-17
 Psyllobora vigintimaculata (Say) 6 04-21 to 07-27

Colydiidae
 Colydium lineola Say 1 06-03-99


Corylophidae
 Molamba sp. 1 05-12-99

Cryptophagidae
 Telmatophilus americanus LeConte 1 07-27-99

Cucujidae
 Cathartosilvanus imbellis (LeConte) 1 09-29
 Cucujus clavipes F. 2 05-05-99
 Pediacus fuscus Erichson 1 06-09-99
 Silvanus muticus Sharp 2 06-16-99
 Uleiota dubius (F.) 1 09-29

Curculionidae
 Acoptus suturalis LeConte 1 06-29
 Anthonomus consimilis Dietz? 1 04-28
 Barypeithes pellucidus (Boheman) 98 05-19 to 07-21

 Caulophilus dubius (Horn) 5 07-31 to 08-16
 Ceutorhynchus septentrionalis Gyll. 1 04-28
 Conotrachelus anaglypticus (Say) 1 09-09
 Conotrachelus aratus (Germar) 2 09-08 to 09-09
 Conotrachelus nenuphar (Herbst) 1 07-14
 Conotrachelus posticatus Boheman 2 09-09 to 09-15
 Cossonus impressifrons Boheman 1 07-21
 Cyrtepistomus castaneus (Roelofs) 1 10-27
 Dorytomus imbecillus Faust 1 06-22
 Dorytomus laticollis LeConte 1 06-22
 Dryophthornus americanus Bedel 3 06-16 to 07-07
 Euparius marmoreus Olivier 1 09-29
 Eurhoptus pyriformis LeConte 1 07-21
 Gymnetron pascuorum (Gyllenhall) 1 06-29
 Lechriops oculata (Say) 1 05-12
 Listronotus sparsus Say 1 04-21
 Mecinus pyraster (Herbst) 5 05-12 to 08-31
 Odontocorynus salebrosus (Casey) 19 06-09 to 07-19
 Otiorhynchus rugosostriatus (Goeze) 5 05-26 to 08-25
 Otiorhynchus sp. 2 08-05
 Otiorhynchus sulcatus (F.) 1 08-18
 Perigaster liturata (Dietz) 1 06-16
 Phyllobius oblongus (L.) 9 05-16 to 06-03
 Phyxelis rigidus (Say) 1 09-29
 Pseudobaris nigrina (Say) 1 04-28
 Rhynchaenus pallicornis (Say) 1 04-28
 Sciaphilus asperatus (Bonsdorff) 8 04-28 to 08-05
 Stenoscelis brevis (Boheman) 17 06-03 to 07-14
 Tachyerges niger (Horn) 2 07-19
 Tychius picirostris (F.) 2 04-28 to 05-19
 Tychius stephensi Schoenherr 1 07-07

Dermestidae
 Dermestes rattus rattus (LeConte) 3 07-30 to 08-04

Derodontidae
 Derodontus maculatus (Melsheimer) 4 10-02 to 11-04

Elateridae
 Aeolus mellilus (Say) 1 08-27
 Agriotes quebecensis Brown 13 04-28 to 06-29
 Ampedus areolatus (Say) 1 06-09
 Ampedus linteus (Say) 1 08-24
 Ampedus nigricollis (Herbst) 1 05-12
 Ampedus sanguinipennis (Say) 1 04-28
 Ampedus semicinctus (Randall) 1 06-09
 Athous brightwelli (Kirby) 1 06-23
 Athous cucullatus (Say) 1 08-12
 Ctenicera caricinus (Germar) 2 04-21
 Ctenicera cylindriformis (Herbst) 6 04-28 to 05-05
 Ctenicera hamata (Say) 1 06-23
 Ctenicera lobatus (VanDyke) 1 04-28
 Ctenicera pyrrhos (Herbst) 16 07-14 to 07-27
 Ctenicera vernalis (Hentz) 1 04-28
 Dalopius spp. 4 05-05 to 06-23
 Elater abruptus Say 2 07-21 to 07-30
 Hemicrepidius bilobatus (Say) 7 08-06 to 08-18
 Hemicrepidius memnonius (Herbst) 8 07-15 to 08-12
 Lacon discoidea (Weber) 2 06-16
 Melanotus castanipes (Paykull) 2 06-03 to 06-09
 Melanotus morosus Candeze 1 07-07
 Melanotus sagittarius (LeConte) 1 05-26
 Melanotus similis (Kirby) 13 06-03 to 09-22
 Melanotus sp. 1 06-03

Endomychidae
 Aphorista vittata (F.) 1 08-10
 Lycoperdina ferruginea LeConte 2 08-12 to 08-16

Erotylidae
 Megalodacne fasciata (F.) 4 07-15 to 08-18
 Triplax festiva Lacordaire 1 08-12
 Triplax thoracica (Say) 2 05-12 to 05-19
 Tritoma biguttata (Say) 4 07-21 to 07-27
 Tritoma mimetica (Crotch) 1 08-18
 Tritoma sanguinipennis (Say) 4 05-26 to 06-29

Eucnemidae
 Dirhagus triangularis (Say) 9 06-03 to 06-23
 Dirhagus pectinatus (LeConte) 1 06-09
 Isorhipis ruficornis (Say) 2 06-03 to 07-07

Geotrupidae
 Bolboceras filicornis (Say) 2 07-15 to 08-25
 Geotrupes splendidus splendidus (F.) 8 05-19 to 08-24

Heteroceridae
 Heterocerus sp. 2 07-27
 Heterocerus tristis Mannerheim 2 07-27
 Tropicus pusillus (Say) 1 07-27

Histeridae
 Aeletes floridae (Marseul) 4 05-12 to 06-16
 Euspilotus assimilis (Paykull) 1 07-21
 Hister abbreviatus F. 2 05-05 to 07-21
 Hister margaranotus LeConte 5 05-05 to 08-04
 Hololepta aequalis (Say) 2 05-19 to 06-03
 Hololepta lucida LeConte 1 06-29
 Platysoma coarctatum LeConte 1 05-05
 Platysoma lecontei Marseul 16 05-05 to 09-15

Hydraenidae
 Hydraena pennsylvanica Kiesenwetter 1 06-11

Laemophloeidae
 Laemophloeus fasciatus Melsheimer 4 05-05 to 11-04
 Lampyridae
 Ellychnia corrusca (L.) 1 06-16
 Lucidota sp. 1 06-29
 Photinus carolinus Green 2 06-11 to 06-29
 Photinus consanguineus LeConte 1 06-29
 Photinus sp. 1 08-04
 Photuris sp. 2 06-16
 Pyropyga decipiens (Harris) 2 06-16 to 06-23
 Pyropyga sp. 7 06-16 to 06-29

Languriidae
 Toramus sp. 1 05-19

Leptodiridae
 Catops sp. 6 07-07 to 07-21
 Prionochaeta opaca (Say) 2 08-04

Lucanidae
 Ceruchus piceus (Weber) 4 06-23 to 10-01
 Platycerus virescens (F.) 1 05-19

Melandryidae
 Dircaea liturata (LeConte) 1 7-27
 Eustrophus tomentosus Say 1 08-18
 Hallomenus sp. 1 09-01
 Melandrya striata Say 3 05-19 to 06-09
 Synchroa punctata Newman 6 06-09 to 07-27

Meloidae
 Epicauta pensylvanica (DeGeer) 1 08-18
 Meloe americanus (Leach) 1 05-05

Monotomidae
 Pycnotomina cavicolle (Horn) 4 05-05 to 06-03

Mordellidae
 Mordella marginata Melsheimer 3 06-16 to 08-17
 Mordellistena aspersa (Melsheimer) 1 06-09
 Mordellistena bihamata (Melsheimer) 1 07-14

Mycetophagidae
 Litargus balteatus LeConte 4 06-09 to 11-04
 Litargus tetraspilotus LeConte 1 06-09
 Mycetophagus melsheimeri LeConte 1 05-05

Oedemeridae
 Asclera ruficollis (Say) 3 04-28 to 05-19

Passandridae
 Catogenus rufus (F.) 2 06-09 to 09-29

Phengodidae
 Phengodes fusciceps LeConte 1 not recorded

Pyrochroidae
 Dendroides canadensis Latreille 4 06-29 to 08-18
 Neopyrochroa flabellata (F.) 7 06-29 to 07-21

Rhizophagidae
 Picnotomina cavicolle (Horn) 1 05-12

Salpingidae
 Rhinosimus viridiaenus Randall 2 05-26 to 10-19

Scarabaeidae
 Aphodius fimetarius (L.) 1 07-26
 Aphodius granarius (L.) 6 08-12 to 09-09
 Aphodius leopardus Horn 1 08-27
 Aphodius lutulentus Haldeman 2 07-27
 Aphodius rujipes (L.) 19 07-15 to 09-09

 Aphodius ruricola Melsheimer 8 04-14 to 09-01
 Ataenius strigatus (Say) 10 04-28 to 10-05
 Copris fricator fricator (F.) 2 09-09 to 09-29
 Cyclocephala borealis Arrow 3 06-29 to 07-15
 Dialytes truncatus Melsheimer 6 08-27 to 09-01
 Dialytes ulkei Horn 7 08-12 to 09-01
 Dichelonyx diluta (Fall) 1 06-29
 Gnorimella maculosa (Knoch) 1 06-03
 Hoplia trifasciata Say 1 04-28
 Onthophagus hecate hecate (Panzer) 21 04-28 to 09-29
 Onthophagus striatulus striatulus 2 06-22 to 06-23
 (Beauvois)
 Osmoderma scabra (Beauvois) 1 07-21
 Phyllophaga balia (Say) 3 07-14 to 07-15
 Phyllophaga knochii (Schoenherr & 1 07-15
 Gyllenhal)
 Popillia japonica Newman 3623 07-07 to 10-12
 Serica intermixta Blatchley 3 07-15 to 07-27
 Serica serica (Illiger) 1 07-27
 Serica vespertina (Gyllenhal) 8 06-09 to 09-09

Scirtidae: (=Helodidae)
 Cyphon nebulosus (LeConte) 2 08-03 to 11-04
 Cyphon neovariabilis Klausnitzer 1 06-03
 Prionocyphon limbatus LeConte 2 08-06 to 09-09

Silphidae
 Necrodes surinamensis (F.) 2 04-14
 Nicrophila americana (L.) 91 06-09 to 08-04
 Nicrophorus orbicollis Say 123 05-26 to 09-08

 Nicrophorus sayi Laporte 6 04-14 to 07-27
 Nicrophorus tomentosus Weber 53 07-07 to 09-15
 Oiceoptoma inaequale (F.) 2 04-14 to 07-21
 Oiceoptoma noveboracense (Forster) 44 04-14 to 08-04

Staphylinidae
 Aleochara sp. 1 07-21
 Aleocharinae sp. 1 10 04-14 to 06-03
 Aleocharinae sp. 2 1 06-09
 Aleocharinae sp. 3 1 05-12
 Aleocharinae sp. 4 1 05-12
 Belonuchus formosus (Gravenhorst) 1 06-03
 Bisnius blandus (Gravenhorst) 1 05-19
 Carphacis intrusus (Horn) 2 05-05 to 06-23
 Creophilus maxillosus (L.) 2 04-14
 Erichsonius brachycephalus Frank 1 not recorded
 Erichsonius cinerascens (Gravenhorst) 1 09-29
 Gabrius sp. 1 07-21
 Hesperus apicialis (Say) 2 05-05 to 05-12
 Homaeotarsus sp. 1 05-12
 Laetulonthus laetulus (Say) 3 04-14 to 05-19
 Lathrobium sp. 1 09-01
 Neohypnus sp. 1 04-28
 Ontholoestes cingulatus (Gravenhorst) 9 05-19 to 08-04
 Oxyporus quinquemaculatus LeConte 1 06-29
 Philonthus politus (L.) 1 04-14
 Philonthus? sp. 3 1 05-05
 Platydracus violaceus (Gravenhorst) 1 09-15
 Quedius fulgidus (F.) 2 09-22
 Quedius plagiatus (=laevigatus) 1 08-18
 Sepedophilus versicolor (Casey) 2 09-29
 Siagonium punctatum LeConte 2 05-19
 Tachinus fimbriatus Gravenhorst 38 06-16 to 09-08
 Tachinus fumipennis (Say) 1 05-19
 Tachinus luridus Erichson 1 04-28
 Xestolinus abdominalus Casey 14 04-28 to 05-05

Tenebrionidae
 Alobates pennsylvanica (DeGeer) 2 08-24 to 08-25
 Anaedus aeneus (Ziegler) 4 07-07 to 08-16
 Bolitotherus cornutus (Panzer) 4 07-27 to 08-10
 Corticeus praetermissus (Fall) 1 05-05
 Diaperis maculata (Olivier) 4 06-15 to 08-04
 Meracantha contracta (Beauvois) 3 06-16 to 06-29
 Neatus tenebrioides (Beauvois) 2 06-11 to 08-24
 Neomida bicornis (F.) 1 06-09
 Paratenetus punctatus Spinola 1 06-16-99
 Uloma impressa Melsheimer 2 07-21 to 09-29

Throscidae
 Aulonothroscus sp. 1 10-27
 Trixagus sp. 8 05-05 to 11-04

Trogidae
 Trox aequalis Say 3 08-12 to 09-29
 Trox capillaris Say 1 04-14
 Trox hamatus Robinson 2 07-27
 Trox unistriatus Beauvois 2 08-27 to 09-09
 Trox variolatus Melsheimer 5 05-29 to 08-18

Trogositidae
 Grynocharis quadrilineata Melsheimer 1 04-28
 Ostomida sp. 1 07-14
 Tenebroides corticalis Melsheimer 2 05-05 to 06-19
 Thymalus marginicollis Chevrolat 1 07-14

Taxon Abun Location

Anthicidae
 Anthicus cervinus (La Ferte) A 1,45,54
 Anthicus ephippium LaFerte A 54
 Omonadus floralis (L.) C 4

Biphyllidae
 Diplococelus brunneus LeConte C 2

Buprestidae
 Chrysobothris rugosiceps Melsheimer C 1
 Taphrocerus gracilis (Say) U 2

Byturidae
 Byturus unicolor Say C [DELTA] *

Cantharidae
 Silis sp. N/A ** 1,2

Cerambycidae
 Analeptura lineola (Say) A 1,2,3
 Bellamira scalaris (Say) U 3
 Brachysomida bivittata (Say) C 1,2,4,11
 Dorcaschema alternatum (Say) C 9
 Gaurotes cyanipennis (Say) C 2
 Orthosoma brunneum (Forster) A 1,3,14
 Parandra (Neandra) brunnea brunnea A 2
 (F.)
 Stictoleptura canadensis Canadensis U 3
 (Olivier)
 Strangalia luteicornis (F.) A 2
 Tetraopes tetrophthalmus (Forster) A 3,33,34
 Tylonotus bimaculatus (Haldeman) U [DELTA]
 Typocerus velutinus velutinus A 1,33
 (Olivier)

Cerylonidae
 Cerylon castaneum Say C 1,2
 Philothermus glabriculus LeConte C 1

Chrysomelidae
 Altica spp. N/A 1,2,3
 Brachycoryna melsheimeri (Crotch) R, SI 4
 Brachypmoea margaretae (Schultz) C 1,19,30,41
 Calligrapha multipunctata bigsbyana C 1
 (Kirby)
 Chaetocnema fuscata White C 4
 Chaetocnema pulicaria Melsheimer A 19
 Chrysochus auratus (F.) C 1,19,30,41
 Chrysolina auripennis (Say) C 2,[DELTA]
 Crepidodera browni Parry C 19
 Diabrotica undecimpunctata howardi A 1,3,19,22,25
 Barber
 Disonycha pensylvanica (Illiger) A 22,[DELTA]
 Donacia sp. N/A 19
 Exema canadensis Pierce A [DELTA]
 Exema sp. N/A 2
 Kuschelina miniata (F.) C 2
 Longitarsus sp. N/A 4
 Luperaltica senilis (Say) A 30
 Microrhopala sp. N/A 4
 Odontota sp. N/A 1,2,3,22,30,34,41
 Ophraella conferta (LeConte) A 9
 Ophraella cribrata (LeConte) A 2
 Ophraella notata (F.) A 1
 Pachybrachys sp. N/A 34
 Paria thoracica (Melsheimer) A 1,6,9,19,34
 Phyllotreta sp. N/A 3
 Plateumaris shoemakeri (Schaeffer) C 11
 Rhabdopterus praetextus (Say) C 2
 Scelodyperus sp.nr liriophilus Wilcox A 3
 Strabala rufa (Illiger) C 2
 Systena blanda Melsheimer A 2
 Tricholochmaea sp. N/A 4,19,34
 Tymnes tricolor (F.) R 2

Ciidae
 Cis levetti (Casey) C 1,2
 Orthocis punctatus (Mellie) U 2

Cleridae
 Enoclerus nigripes (Say) A 1,3
 Necrobia rufipes DeGeer C 4
 Necrobia violacea L. C 39
 Placopterus thoracicus (Olivier) C 2
 Thanasimus dubius (F.) C 2
 Zenodosus sanguineus (Say) C 2,4

Coccinellidae
 Brachiacantha rotunda Gordon C 1,4
 Coccinella septempunctata L. C 4
 Coleomegilla maculata lengi Timberlake C 1,3,30
 Cycloneda munda (Say) C 4,19,[DELTA]
 Harmonia axyridis (Pallas) A 1,2,19,30,33
 Hippodamia parenthesis (Say) U 3,19,[DELTA]
 Psyllobora vigintimaculata (Say) C 1,2,3,30

Colydiidae
 Colydium lineola Say U 1

Corylophidae
 Molamba sp. N/A 4

Cryptophagidae
 Telmatophilus americanus LeConte U 30

Cucujidae
 Cathartosilvanus imbellis (LeConte) C 1
 Cucujus clavipes F. U 2,[DELTA]
 Pediacus fuscus Erichson U [DELTA]
 Silvanus muticus Sharp C 1
 Uleiota dubius (F.) C 1

Curculionidae
 Acoptus suturalis LeConte C 2
 Anthonomus consimilis Dietz? C 19
 Barypeithes pellucidus (Boheman) C 1,3,4,20,25,27,28,
 29,33,35,47,48
 Caulophilus dubius (Horn) C 1,37
 Ceutorhynchus septentrionalis Gyll. C 4
 Conotrachelus anaglypticus (Say) C 16
 Conotrachelus aratus (Germar) C 2,16
 Conotrachelus nenuphar (Herbst) C 32
 Conotrachelus posticatus Boheman C 16,28
 Cossonus impressifrons Boheman C 1
 Cyrtepistomus castaneus (Roelofs) C 2
 Dorytomus imbecillus Faust R 48
 Dorytomus laticollis LeConte C 48
 Dryophthornus americanus Bedel C 2
 Euparius marmoreus Olivier C 1
 Eurhoptus pyriformis LeConte A 1
 Gymnetron pascuorum (Gyllenhall) C 41
 Lechriops oculata (Say) C 2
 Listronotus sparsus Say C 4
 Mecinus pyraster (Herbst) C,SI 1
 Odontocorynus salebrosus (Casey) C 1,19,33,39,41
 Otiorhynchus rugosostriatus (Goeze) C 15,28,37
 Otiorhynchus sp. N/A 46
 Otiorhynchus sulcatus (F.) C 2
 Perigaster liturata (Dietz) R 2
 Phyllobius oblongus (L.) C 1,2,3,4
 Phyxelis rigidus (Say) C 1
 Pseudobaris nigrina (Say) C 4
 Rhynchaenus pallicornis (Say) C 4
 Sciaphilus asperatus (Bonsdorff) C 2,3,4,21,39,41
 Stenoscelis brevis (Boheman) C 2,28,[DELTA]
 Tachyerges niger (Horn) C 19
 Tychius picirostris (F.) C 4
 Tychius stephensi Schoenherr C 1

Dermestidae
 Dermestes rattus rattus (LeConte) SI 2,[DELTA]

Derodontidae
 Derodontus maculatus (Melsheimer) C 2

Elateridae
 Aeolus mellilus (Say) A 16
 Agriotes quebecensis Brown C 2,3,4
 Ampedus areolatus (Say) C [DELTA]
 Ampedus linteus (Say) A 22
 Ampedus nigricollis (Herbst) A 2
 Ampedus sanguinipennis (Say) A 2
 Ampedus semicinctus (Randall) A 2
 Athous brightwelli (Kirby) A 2
 Athous cucullatus (Say) C 28
 Ctenicera caricinus (Germar) A 2
 Ctenicera cylindriformis (Herbst) A 2
 Ctenicera hamata (Say) C 2
 Ctenicera lobatus (VanDyke) C 2
 Ctenicera pyrrhos (Herbst) A 2,15,18,21
 Ctenicera vernalis (Hentz) C 2
 Dalopius spp. N/A 1,2,3
 Elater abruptus Say A 2
 Hemicrepidius bilobatus (Say) A 3,24,28
 Hemicrepidius memnonius (Herbst) C 1,2,16,18,21,24
 Lacon discoidea (Weber) A 2
 Melanotus castanipes (Paykull) C 2
 Melanotus morosus Candeze A 4
 Melanotus sagittarius (LeConte) A 2
 Melanotus similis (Kirby) A 1,2,4,16
 Melanotus sp. N/A 3

Endomychidae
 Aphorista vittata (F.) C 50
 Lycoperdina ferruginea LeConte C 24,59

Erotylidae
 Megalodacne fasciata (F.) A 14,21,22
 Triplax festiva Lacordaire C 14
 Triplax thoracica (Say) A 2
 Tritoma biguttata (Say) A 1,18
 Tritoma mimetica (Crotch) R 55
 Tritoma sanguinipennis (Say) A 2,3

Eucnemidae
 Dirhagus triangularis (Say) C 2
 Dirhagus pectinatus (LeConte) C 2
 Isorhipis ruficornis (Say) C 2,3

Geotrupidae
 Bolboceras filicornis (Say) C 3,[DELTA]
 Geotrupes splendidus splendidus (F.) C 1,3,22

Heteroceridae
 Heterocerus sp. N/A 30,60
 Heterocerus tristis Mannerheim A 30,60
 Tropicus pusillus (Say) A 60

Histeridae
 Aeletes floridae (Marseul) U 1
 Euspilotus assimilis (Paykull) A 39
 Hister abbreviatus F. A 39
 Hister margaranotus LeConte A 1,2,22
 Hololepta aequalis (Say) C 1
 Hololepta lucida LeConte U 1
 Platysoma coarctatum LeConte C 2
 Platysoma lecontei Marseul A 1,2,3

Hydraenidae
 Hydraena pennsylvanica Kiesenwetter U [DELTA]

Laemophloeidae
 Laemophloeus fasciatus Melsheimer C 2
 Lampyridae
 Ellychnia corrusca (L.) A 2
 Lucidota sp. N/A 2
 Photinus carolinus Green U 2,29
 Photinus consanguineus LeConte C 1
 Photinus sp. N/A 1
 Photuris sp. N/A 4
 Pyropyga decipiens (Harris) C 1,4
 Pyropyga sp. N/A 1

Languriidae
 Toramus sp. N/A 4

Leptodiridae
 Catops sp. N/A 2
 Prionochaeta opaca (Say) U 2

Lucanidae
 Ceruchus piceus (Weber) C 2,22
 Platycerus virescens (F.) C 3

Melandryidae
 Dircaea liturata (LeConte) A 20
 Eustrophus tomentosus Say U 55
 Hallomenus sp. N/A 41
 Melandrya striata Say C 2
 Synchroa punctata Newman A 2,[DELTA]

Meloidae
 Epicauta pensylvanica (DeGeer) A 19
 Meloe americanus (Leach) C 2

Monotomidae
 Pycnotomina cavicolle (Horn) 12 2,3

Mordellidae
 Mordella marginata Melsheimer A 4,19,25
 Mordellistena aspersa (Melsheimer) A 1
 Mordellistena bihamata (Melsheimer) C 3

Mycetophagidae
 Litargus balteatus LeConte A 2
 Litargus tetraspilotus LeConte U 2
 Mycetophagus melsheimeri LeConte R 2

Oedemeridae
 Asclera ruficollis (Say) C 2,3

Passandridae
 Catogenus rufus (F.) U 1,2

Phengodidae
 Phengodes fusciceps LeConte R [DELTA]

Pyrochroidae
 Dendroides canadensis Latreille C 3,21,30
 Neopyrochroa flabellata (F.) A 23,15

Rhizophagidae
 Picnotomina cavicolle (Horn) R 2

Salpingidae
 Rhinosimus viridiaenus Randall 3

Scarabaeidae
 Aphodius fimetarius (L.) A [DELTA]
 Aphodius granarius (L.) A 5,9,24
 Aphodius leopardus Horn C 28
 Aphodius lutulentus Haldeman A 18,21
 Aphodius rujipes (L.) A 2,3,5,15,16
 18,24,28,41
 Aphodius ruricola Melsheimer C 2,21,24,28,30,39
 Ataenius strigatus (Say) A 2,4,9,29,60
 Copris fricator fricator (F.) A 9,42
 Cyclocephala borealis Arrow C 4,15
 Dialytes truncatus Melsheimer C 60,61,62
 Dialytes ulkei Horn C 13,47,60,61,62
 Dichelonyx diluta (Fall) C 2
 Gnorimella maculosa (Knoch) C 2
 Hoplia trifasciata Say C 2
 Onthophagus hecate hecate (Panzer) A 1,3,4
 Onthophagus striatulus striatulus C 2,3
 (Beauvois)
 Osmoderma scabra (Beauvois) C 2
 Phyllophaga balia (Say) A 3,16
 Phyllophaga knochii (Schoenherr & C 16
 Gyllenhal)
 Popillia japonica Newman A 2,4
 Serica intermixta Blatchley C 16,18,30
 Serica serica (Illiger) C 18
 Serica vespertina (Gyllenhal) A 2,5,14,18

Scirtidae: (=Helodidae)
 Cyphon nebulosus (LeConte) C 1
 Cyphon neovariabilis Klausnitzer C 4
 Prionocyphon limbatus LeConte C 5,56

Silphidae
 Necrodes surinamensis (F.) U 39
 Nicrophila americana (L.) A 1,2,3,4
 Nicrophorus orbicollis Say A 1,2,3,4,5,6,8,14
 18,20,21,24,25,28
 Nicrophorus sayi Laporte C 2,3
 Nicrophorus tomentosus Weber C 1,2,3,4
 Oiceoptoma inaequale (F.) C 2,39
 Oiceoptoma noveboracense (Forster) C 1,2,3,4,39

Staphylinidae
 Aleochara sp. N/A 39
 Aleocharinae sp. 1 N/A 1,2,4
 Aleocharinae sp. 2 N/A 3
 Aleocharinae sp. 3 N/A 1
 Aleocharinae sp. 4 N/A 4
 Belonuchus formosus (Gravenhorst) A 1
 Bisnius blandus (Gravenhorst) A 3
 Carphacis intrusus (Horn) C 1,3
 Creophilus maxillosus (L.) A 39
 Erichsonius brachycephalus Frank NSR *** 47
 Erichsonius cinerascens (Gravenhorst) U 2
 Gabrius sp. N/A 57
 Hesperus apicialis (Say) U 1,2
 Homaeotarsus sp. N/A 2
 Laetulonthus laetulus (Say) U 1
 Lathrobium sp. N/A 22
 Neohypnus sp. N/A 2
 Ontholoestes cingulatus (Gravenhorst) A 1,3,4,22
 Oxyporus quinquemaculatus LeConte U 1
 Philonthus politus (L.) A 39
 Philonthus? sp. 3 N/A 2
 Platydracus violaceus (Gravenhorst) C 1
 Quedius fulgidus (F.) C 1
 Quedius plagiatus (=laevigatus) C 2
 Sepedophilus versicolor (Casey) C 2
 Siagonium punctatum LeConte NSR 1
 Tachinus fimbriatus Gravenhorst A 1,2,3
 Tachinus fumipennis (Say) A 3
 Tachinus luridus Erichson A 1
 Xestolinus abdominalus Casey NSR 1,4

Tenebrionidae
 Alobates pennsylvanica (DeGeer) A 15,22
 Anaedus aeneus (Ziegler) U 20,37
 Bolitotherus cornutus (Panzer) C 3,14,20,21
 Corticeus praetermissus (Fall) R 2
 Diaperis maculata (Olivier) C 20,22,29
 Meracantha contracta (Beauvois) U 1,28,36
 Neatus tenebrioides (Beauvois) A 1,28,36
 Neomida bicornis (F.) C 2
 Paratenetus punctatus Spinola C 27
 Uloma impressa Melsheimer R 1,41

Throscidae
 Aulonothroscus sp. N/A 2
 Trixagus sp. N/A 2,3,6,30,45

Trogidae
 Trox aequalis Say C 42,55
 Trox capillaris Say U 39
 Trox hamatus Robinson C 21,30
 Trox unistriatus Beauvois C 62
 Trox variolatus Melsheimer A 2,3

Trogositidae
 Grynocharis quadrilineata Melsheimer U 2
 Ostomida sp. N/A 2
 Tenebroides corticalis Melsheimer U 2
 Thymalus marginicollis Chevrolat U 2

* [DELTA] Denotes a specimen within the compound,
exact location not specified.

** N/A Denotes specimens identified to genus only.

*** NSR Denotes new state record.


ACKNOWLEDGMENTS. We are especially pleased with the cadre of specialists who have made the determinations of the many specimens which were captured.

We wish to thank all of the other collectors that were active in the Training Grounds in 1999 for their help and cooperation for collecting beetles for this project. Many of these beetles were unique. Specifically we want to thank Michael Gilligan, Michael Hoggarth, Megan Michael, Daniel Rice, and Daniel Riggs for the number of beetles they caught and had included in the survey. We wish to thank Larry Roche for his enthusiasm and moral support throughout the season.

At the OARDC in the Communications and Technology Department we wish to thank Kenneth Chamberlain, Photographer, for his fine work both in the field and photo lab, for his expertise and technical work for photos in this report and on the Coleoptera poster; Jesse Ewing, Graphic Designer, for his technical work on slides and photos for the report and poster. Also, many thanks to Bonnie Bing for her patience, graphics advice, and technical skills in the creation of the poster and report cover. Jody Lanham, Systems Specialist in the Computing and Statistical Services Department, used her software expertise to help us put together this report, for which we are very grateful.

We also wish to thank the fine team of Coleoptera specialists who have helped with the difficult task of getting the specimens identified. M. Scan Ellis was our Master Separator of material of the entire order of Coleoptera. He separated the field collected material into families. Scan also identified all the sap beetles (Nitidulidae) to species. Other specialists were: Robert Androw (Cerambycidae, Buprestidae & Cleridae); Richard Beal (Dermestidae, esp. Amhrenus); Marc Branham (Lampyridae); Donald Chandler (Anthrihidae); Shawn Clark (Chrysomelidae); Wills Flowers (Chrysomelidae); Michael Goodrich (Byturidae, Biphyllidae, Corylophidae, Erotylidae, Oxyporinae); Michael Ivie (Trogossitidae; Stenotrachelidae, Colydiidae; Lycidae); Sophien Kamoun (Carabidae); George Keeney (Silphidae); Peter Kovarik (Histeridae); John Lawrence (Ciidae); Wenhua Lu (Mordellidae); Alfred F. Newton (Staphylinidae); Charles O'Brien (Curculionidae); Philip Perkins (Hydraenidae); Robert Rabaglia (Scolytidae); Steve Schott (Heteroceridae); Paul Skelley (Anthribidae, Biphyllidae, Endomychidae, Cucujidae & Cryptophagidae); Andrew Smith (Scarabaeidae); Margaret Thayer (Staphylinidae); Michael Thomas (Cucujidae); C. A. Triplehom (Tenebrionidae and several other families); Samuel Wells (Elateridae & Eucnemidae); and Daniel Young (Scirtidae & Pyrochroidae). Most of these workers identified additional families which greatly facilitated the process of identification.

At The Ravenna Training and Logistics Site we wish to thank Timothy Morgan, Natural Resources Manager, and Lt. Col. Thomas A. Tadsen. Deputy Training Site Commander, for their valued cooperation. Also thanks to Captain Thomas Daugherty, Environmental Protection Specialist for the Ohio National Guard, for his keen interest in preserving the environment, and his interest in beetles, which made this project possible. Guards Frank Hertig and Emma Lamp at the gate who kept us informed of daily activities at the Arsenal, were extremely helpful with their assistance on directions, and passing along pertinent information to other researchers.

LITERATURE CITED

Braun EL. 1989. The woody plants of Ohio. Columbus (OH): The Ohio State University Pr. p 19-32.

Peck SB, Davies AE. 1980. Collecting small beetles with large area window traps. Coleopterists Bull 34(2):237-9.

Tertuliani JS. 1999. Aquatic macroinvertebrates collected at Ravenna Army Ammunition Plant, Portage and Trumbull Counties, Ohio. 1998 Water-Resources Investigations Report 99-4202. US Dept of the Interior, US Geological Survey. In cooperation with the Ohio Army National Guard. 38 p.

Williams RN, Ellis MS, Fickle DS. 1995. Insects in the Killbuck Marsh Wildlife Area: 1993 Survey. Ohio J Sci 95(3):226-32.

(1) Manuscript received 17 March 2002 and in revised form 4 November 2002 (#02-05).

ROGER WILLIAMS AND DIANE HARTZLER, Department of Entomology, OARDC, The Ohio State University, Wooster, OH 44691-4096
COPYRIGHT 2004 Ohio Academy of Science
No portion of this article can be reproduced without the express written permission from the copyright holder.
Copyright 2004 Gale, Cengage Learning. All rights reserved.

Article Details
Printer friendly Cite/link Email Feedback
Author:Williams, Roger; Hartzler, Diane
Publication:The Ohio Journal of Science
Geographic Code:1U3OH
Date:Apr 1, 2004
Words:7094
Previous Article:What is at stake with high stakes testing? A discussion of issues and research (1).
Next Article:Comparison of standing vegetation and seed bank composition one year following hardwood reforestation in Southwestern Ohio (1).
Topics:

Terms of use | Copyright © 2017 Farlex, Inc. | Feedback | For webmasters