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Seep community at White Oak Creek Woods Natural Area, Mason County, Illinois.

ABSTRACT

A series of small seeps were studied at White Oak Creek Woods Natural Area, Mason County, Illinois. The seep community in full sunlight had 23 species present in the plots (n = 30). Apios americana (groundnut) was dominant with a mean cover of 33.88% and an IV of 31.5 (possible 200), followed by Impatiens capensis (spotted touch-me-not), Leersia oryzoides (rice cut grass), Decodon verticillatus (swamp loosestrife), and Saururus cernuus (lizard's-tail) all with IV's >15.0. In the shaded seep, of the 19 species in the plots (n = 30), Symplocarpus foetidus (skunk cabbage) was dominant with a mean cover of 39.75% and an IV of 70.2. Impatiens capensis and Saururus cernuus were second and third with IV's of 41.7 and 40.7 respectively, the remaining species being of minor importance. Within the seep complex, a total of 140 vascular plant species representing 110 genera and 57 families were documented. During the present study many more species were found than were reported in an earlier study of the site conducted in 1912.

INTRODUCTION

Seep communities occur in areas with saturated soil caused by groundwater reaching the surface in a diffuse flow (White and Madany 1978). These seeps, which rarely exceed 0.5 ha, are common along the lower slopes of glacial moraines, in ravines, and on stream terraces. Ground water of seeps usually flows through sand and gravel deposits above an impervious soil layer to the outlet area where it forms a distinct seep line. The resulting seeps are permanently wet, marshy areas. Seeps may have some localized areas of concentrated flow (springs), but these areas are not obvious. It is common, however, that some of the seep waters flow from the site in shallow runs.

Seeps are relatively common throughout Illinois, but most are small, and these communities and the associated plant species seldom have been studied in detail. Most seeps discussed in the literature, were studied because they contained an unusual assemblage of plant species or because a few endangered or threatened species were present. The first study describing seep community vegetation in Illinois was by Gates (1912) in his analysis of a "bog" community in Mason County, Illinois. He was interested in this site because "it is so far south of the usual southern limits of peat-bog plants as outlined by Transeau" (1903). More recently Phipps and Speer (1958) and later Parker and Ebinger (1971) studied the vascular plant species of a hillside marsh in Coles County, east-central Illinois. Later Ebinger (1978) examined the flora of seven seeps in east-central Illinois, while Henry and Scott (1984) studied the vegetation of four seeps in McDonough County in northwestern Illinois. In southern Illinois Voigt and Mohlenbrock (1964) described a few seeps from the Shawnee Hills Natural Division, while Schwegman (1969) examined a series of seep springs in the Cretaceous Hills Section of the Coastal Plain Natural Division (Schwegman 1973).

The present study was undertaken to determine the composition and structure of the vegetation of this series of seeps at White Oak Creek Natural Area in Mason County, and compare the result to those observed by Gates (1912).

DESCRIPTION OF THE STUDY AREA

The seep complex studied is located along White Oak Creek about 6 km south of Havana, Mason County, Illinois (NW1/4 S23 T21N R9W). At this site numerous small seeps are located along both sides of White Oak Creek on a sandy terrace about 400 m east of the Illinois River. The largest seep is nearly 125 m long and varies in width from 1 to 8 m. This seep is in an open area with nearly continual sunlight throughout the day (sunny seep). The remaining seeps are smaller with the largest approximately 100 m long by about 10 m wide, and shaded nearly the entire day (shaded seep). The seeps are in a narrow valley surrounded by steep slopes 3 to 5 m high. The upland forest surrounding the seeps is dominated by Quercus velutina (black oak) and Q. alba (white oak). The soils are excessively drained Plainfield sand (Calsyn 1995), part of the dune and swale topography known as the Parkland Formation (Willman and Frye 1970), which occurs in the Illinois River Section of the Illinois River and Mississippi River Sand Areas Natural Division (Schwegman 1973).

The timbered land north of White Oak Creek, was designated "grade B" dry to dry-mesic sand forest by the Illinois Natural Areas Inventory (White 1978). The larger white oak trees were selectively logged north of White Oak Creek in 1980, resulting in that area being designated "grade C" forest (Lerczak 2000). Registered as an Illinois Natural Heritage Landmark since 1983, it is now designated as the Speckman-Stelter Woods Land and Water Reserve by the Illinois Nature Preserves Commission (Lerczak 2000).

METHODS

The seep complex was visited seven times each year throughout the growing seasons of 2004 and 2005. Voucher specimens of plant species listed in Appendix I were collected, identified, and deposited in the Stover-Ebinger Herbarium of Eastern Illinois University, Charleston, Illinois (EIU), or the herbarium of the Illinois Natural History Survey, Champaign, Illinois (ILLS). Nomenclature follows Mohlenbrock (2002).

In late July of 2005, a transect 90m long was located along the centerline of the sunny seep. At 3 m intervals along this transect [m.sup.2] plots were located randomly 0 to 2 m from the transect line, even number plots to the right, odd to the left (n=30). In the shaded seep two transects 45 m long were randomly located and the same procedure followed (n=30).

Species cover was determined using the Daubenmire (1959) cover class system as modified by Bailey and Poulton (1968). The modified Daubenmire cover scale is as follows: class 1 = 0 to 1%; class 2 = 1 to 5%; class 3 = 5 to 25%; class 4 = 25 to 50%; class 5 = 50 to 75%; class 6 = 75 to 95%; class 7 = 95 to 100%. Mean cover was then calculated by using the mid-point of the cover classes. Importance value (IV) for each species was determined by summing relative cover and relative frequency.

The Sorensen Index of Similarity (ISs) was used to determine the degree of vegetation similarity between the two seep communities studied (Mueller-Dombois and Ellenberg 1974). In this index [ISs = 2C/A+B x 100], A equals the number of species in the first community, B equals the number of species in the second community, and C equals the number of species common between the two communities.

RESULTS

Within the seep complex, a total of 140 vascular plant species representing 110 genera and 57 families were documented (Appendix I). Of the species listed in Appendix I, four were ferns and fern-allies, 27 were monocots in 22 genera and nine families, while 109 were dicots in 85 genera and 45 families. Although Gates (1912) reported the occurrence of the state endangered Mimulus glabratus (yellow monkey flower), and the presently state threatened Veronica scutellata (marsh speedwell), neither of these two species nor any other state threatened or endangered species were encountered during the present study (Herkert and Ebinger 2002).

The sunny seep community had 23 species recorded in the plots. The perennial vine, Apios americana (groundnut) and the annual, Impatiens capensis (spotted touch-me-not) were the dominant species with mean cover values of 33.88% and 26.78% and IV's of 31.5 and 30.3, respectively (Table 1). In many parts of the seep, Apios americana overtopped and covered the other vegetation. Leersia oryzoides (rice cut grass), Decodon verticillatus (swamp loosestrife), and Saururus cernuus (lizard's-tail) all had IV's greater than 15, and mean covers that ranged from 8.35% to 15.57%. The only non-native species encountered was Lysimachia nummularia (moneywort), with an IV of 4.4. This species formed scattered, cushion-like patches beneath the much taller vegetation.

Of the 19 species in the plots of the shaded seep, Symplocarpus foetidus (skunk cabbage) was dominant with a mean cover of 39.75% and an IV of 70.2. Impatiens capensis and Saururus cernuus were second and third, with IV's of 41.7 and 40.7 respectively (Table 1). The remaining 16 species encountered had much lower mean covers and IV's. The non-native Alliaria petiolata (garlic mustard), a pervasive adventive species of many Illinois sand deposits, was occasionally found in the shaded seep plots (IV of 2.5) and was commonly observed in the forests surrounding the seeps.

The mean cover of the sunny seep was 153.12%, nearly twice the mean cover of the shaded seep (80.61%). Consistent with this, the average mean cover for bare ground and litter was nearly eight times greater in the shaded seep (39.43%) than in the sunny seep (4.50%). Twelve species encountered in the plots were present in both of the seep communities resulting in a Sorensen Index of Similarity (ISs) of 57.1. Impatiens capensis was second in IV in both seep communities, and Saururus cernuus was fifth in IV in the sunny seep (IV of 15.4) and third in the shaded seep (IV of 40.7). Three of the top five species encountered in the sunny seep (Apios americana, Leersia oryzoides, Decodon verticillatus) were not observed in the shaded seep. Symplocarpus foetidus, the dominant species of the shaded seep (IV of 70.2), was occasionally found beneath the taller vegetation in the sunny seep (IV of 4.9). Of the species restricted to the shaded seep, Fraxinus nigra (black ash) seedlings were the most common (IV of 7.2), followed by the nonnative Alliaria petiolata, and five other species with low IVs that were rarely encountered (Table 1).

Fraxinus nigra was common in many parts of the terrace, being particularly abundant in and at the edges of the shaded seeps. Seedlings and a few saplings of F. nigra were common in the shaded seep along with a few trees that rarely exceeded 25 cm dbh. In the sunny seep, Salix discolor (pussy willow) and S. interior (sandbar willow) was observed throughout, mostly being less than 3m tall. The slightly woody Decodon verticillatus was the dominant species in parts of the seep, while the shrubs Cephalanthus occidentalis buttonbush), Ribes americanum (Missouri gooseberry), and Rosa palustris (swamp rose) were occasionally observed. Most of the remaining tree and shrub species listed in Appendix I were found at the edges of the seeps and along White Oak Creek.

DISCUSSION

In addition to the endangered and threatened plant species listed by Gates (1912), 17 other species, cited by Gates (1912), were not found during the present study. A few are very common species, while some are relatively uncommon [Amsonia tabernaemontana (blue star), Cicuta bulbifera (bulblet water hemlock), Oxypolis rigidior (cowbane), Ranunculus pensylvanicus (bristly crowfoot), Sparganium eurycarpum bur-reed)]. The reason for these losses is not known, but is probably related to changes in water flow and water levels in the seeps, down cutting and erosion by White Oak Creek, and the loss of habitat resulting from human disturbances since 1912. Additionally, Gates (1912) mentioned springs along the bluff and listed many species, which could not be located during the current study, as occurring in these spring communities. Sand bluffs resulting from undercutting by White Oak Creek were found near the seep communities, but no springs were present at the base. Also, Cornus amomum (swamp dogwood) and Ulmus thomasii (rock elm) probably represent misidentifications by Gates (1912) as they are generally not found in this part of Illinois.

The soils of this seep and most seeps in Illinois have a deep, peaty, soil with water that is usually neutral to slightly alkaline. Ebinger and Bacone (1980) found the waters to be slightly alkaline for two seeps at Turkey Run State Park, Parke County, Indiana. In their examination of four seeps in McDonough County, Illinois, Henry and Scott (1984) found the pH of seep water in three seeps to be neutral to slightly alkaline (pH of 7.0-7.6), while water of the fourth seep was very acidic (pH of 3.5). This acid seep, referred to as a coal seep, was dominated by Sphagnum and was heavily shaded by dense mesic woods.

Many species found in the present study were listed for other Illinois seeps. Only a few species, however, have consistently been found in nearly all seeps examined, including Agrostis gigantea (red top), Aster lateriflorus (side-flowering aster), Carex lurida (lurid sedge), Equisetum arvense (common horsetail), Eupatorium perfoliatum (perfoliate bone set), Glyceria striata (fowl manna grass), Impatiens capensis, Leersia virginica (white grass), Mimulus ringens (sessile monkey flower), Lobelia siphilitica (great blue lobelia), and Pilea pumila (clearweed). Two of these species consistently found with relatively high IV's are Impatiens capensis and Pilea pumila. The remainder, however, usually have low IV's, and are widely scattered or grow in localized clumps.

APPENDIX I.

Vascular plant species encountered in a seep community at White Oak Creek Woods Natural Area, Mason County, Illinois, are listed alphabetically by family under major plant groups. Collecting numbers preceded by an E represent specimens collected by John E. Ebinger and are deposited in the Stover-Ebinger Herbarium, Eastern Illinois University, Charleston, Illinois (EIU). Collecting numbers preceded by a P represent specimens collected by Loy R. Phillippe and are deposited in the Illinois Natural History Survey Herbarium, Champaign, Illinois (ILLS). The species reported by Gates (1912) are preceded by asterisks. A few Gates specimens were located at the University of Illinois Herbarium (ILL), but vouchers for most are missing.

FERNS AND FERN-ALLIES

Equisetaceae

Equisetum arvense L.: E31982

Equisetum laevigatum A. Br.: E31983

Onocleaceae

Onoclea sensibilis L.: P37204

Thelypteridaceae

*Thelypteris palustris Schott: P37227

MONOCOTS

Alismataceae

Alisma subcordatum Raf.: P37294

*Sagittaria latifolia Willd.: P37285

Araceae

*Peltandra virginica (L.) Schott: P37223

*Symplocarpus foetidus (L.) Nutt.: E31818

Cyperaceae

Bolboschoenus fluviatilis (Torr.) Sojak: P37295

Carex alopecoidea Tuckerm.: E31985

Carex blanda Dewey: E31984

Carex cristatella Britt.: P37221

*Carex lurida Wahl: P37212

Cyperus strigosus L.: P37213

Scirpus atrovirens Willd.: P37205

Dioscoreaceae

Dioscorea villosa L.: E31986

Iridaceae

*Iris shrevei Small: E31987

Lemnaceae

Lemna minor L.: E31994

Poaceae

*Agrostis gigantea Roth

Agrostis perennans (Walt.) Tuckerm.: P37274

*Cinna arundinacea L.: P37185

Dichanthelium clandestinum (L.) Gould: P37273

Elymus virginicus L.: P37296

Festuca subverticillata (Pers.) E.B. Alexeev.: E31988

Glyceria striata (Lam.) Hitchc.: E31989

Leersia oryzoides (L.) Swartz: E32059

*Leersia virginica Willd.: P37194

Muhlenbergia frondosa (Poir.) Fern.: P37220

*Poa sylvestris Gray: E31990

Smilacaceae

Smilax tamnoides L.: E32110

Sparganiaceae

*Sparganium eurycarpum Engelm.

DICOTS

Aceraceae

Acer negundo L.: E31972

*Acer saccharinum L.: E32060

Anacardiaceae

Toxicodendron radicans (L.) Kuntze: P37198

Annonaceae

Asimina triloba (L.) Dunal: E31973

Apiaceae

*Berula erecta (Huds.) Coville: E32122

*Cicuta bulbifera L.

*Cicuta maculata L.: P37217

Osmorhiza longistylis (Torr.) DC.: E31974

*Oxypolis rigidior (L.) Raf.

Sanicula canadensis L.: P37206

Sium suave Walt.: E32061

Apocynaceae

*Amsonia tabernaemontana Walt.

Asclepiadaceae

*Asclepias incarnata L.

Asteraceae

Ageratina altissima (L.) R.M. King & H. Rob.: E32062

Arnoglossum atriplicifolium (L.) H. Rob.: E31992

Aster lanceolatus Willd.: E32111

Aster lateriflorus (L.) Britt.: E32112

Aster ontarionis Wieg.: P37292

Bidens cernua L.: P37281

*Bidens comosa (Gray) Wieg.: E32063

Bidens frondosa L.: E32064

Erigeron philadelphicus L.: E31975

Eupatoriadelphus maculatus (L.) R.M. King & H. Rob.: P37193

Eupatoriadelphus purpureus (L.) R.M. King & H. Rob.: E32065

*Eupatorium perfoliatum L.: P37210

Eupatorium serotinum Michx.: P37209

Lactuca floridana (L.) Gaertn.: P37203

Solidago gigantea Ait.: E32066

Solidago ulmifolia Muhl.: E32113

Vernonia gigantea (Walt.) Trel.: P37287

Xanthium strumarium L.: P37283

Balsaminaceae

*Impatiens capensis Meerb.: P37189

Betulaceae

*Betula nigra L.: E32067

Bignoniaceae

*Campsis radicans (L.) Seem.

Boraginaceae

Hackelia virginiana (L.) I.M. Johnston: P37201

Brassicaceae

Cardamine bulbosa (Muhl.) BSP.: E32124

Nasturtium officinale R. Br.: E31976

Caesalpiniaceae

*Gleditsia triacanthos L.: E32068

Gymnocladus dioicus (L.) K. Koch: P37224

Campanulaceae

Campanulastrum americanum (L.) Small: P37228

Lobelia cardinalis L.: P37219

Lobelia siphilitica L.: E32069

Caprifoliaceae

*Sambucus canadensis L.: P39163

Viburnum lentago L.: P37197

Cornaceae

*Cornus amomum Mill.

Cornus drummondii C.A. Mey.: P37207

Cornus sericea L.: P37230

Corylaceae

Carpinus caroliniana Walt.: P39164

Cucurbitaceae

Sicyos angulatus L.: P37284

Cuscutaceae

Cuscuta gronovii Willd.: P37226

Ebenaceae

*Diospyros virginiana L.: E32070

Euphorbiaceae

Acalypha rhomboidea Raf.: P37275

Fabaceae

Amorpha fruticosa L.: P37225

*Apios americana Medic.: P37216

Desmodium paniculatum (L.) DC.: E32071

Gentianaceae

Gentiana andrewsii Griseb.: E32114

Grossulariaceae

Ribes americanum Mill.: E31820

Juglandaceae

*Juglans nigra L.: E32072

Lamiaceae

Lycopus americanus Muhl.: P37214

Lycopus virginicus L.:P37290

*Mentha arvensis L.: P37202

Physostegia speciosa (Sweet) Sweet: P37229

*Scutellaria lateriflora L.: P37199

Stachys tenuifolia Willd.: P37188

Lythraceae

*Decodon verticillatus (L.) Ell.: P37215

Menispermaceae

Menispermum canadense L.: P39165

Moraceae

Morus rubra L.: E32073

Oleaceae

*Fraxinus lanceolata Borkh.: E32074

*Fraxinus nigra Marsh.: E31821

Ligustrum vulgare L.: E32115

Oxalidaceae

Oxalis stricta L.: E31977

Platanaceae

*Platanus occidentalis L.

Polygonaceae

Antenoron virginianum (L.) Roberty & Vautier: P37190

Persicaria pensylvanica (L.) Small: P37282

Persicaria punctata (Ell.) Small: P37222

*Rumex orbiciulatus Gray

Primulaceae

*Lysimachia ciliata L.

Lysimachia nummularia L.: E32075

Ranunculaceae

Anemone canadensis L.: E31978

*Ranunculus abortivus L.: E32123

*Ranunculus pensylvanicus L.f.

Ranunculus recurvatus Poir.: E31979

Rosaceae

Geum canadense Jacq.: E32116

*Rosa palustris Marsh.: P37187

Rubiaceae

*Cephalanthus occidentalis L.: P37218

Galium aparine L.: E31981

*Galium trifidum L.

Galium triflorum Michx.: E32117

Salicaceae

*Salix discolor Muhl.: E32077

*Salix interior Rowlee: E32076

Saururaceae

*Saururus cernuus L.: P37196

Scrophulariaceae

*Chelone glabra L.: P37291

*Mimulus glabratus HBK.

Mimulus ringens L.: P37208

*Veronica scutellata L.

Tiliaceae

*Tilia americana L.

Ulmaceae

*Celtis occidentalis L.: E32118

*Ulmus americana L.: E32078

Ulmus rubra Muhl.: E31980

*Ulmus thomasii Sarg.

Urticaceae

*Boehmeria cylindrica (L.) Sw.: P37191

*Laportea canadensis (L.) Wedd.: E32079

*Pilea pumila (L.) Gray: P37195

Urtica gracilis Ait.: P37211

Verbenaceae

*Phyla lanceolata (Michx.) Greene

*Verbena hastata L.

Vitaceae

Ampelopsis cordata Michx.: P37280

Parthenocissus quinquefolia (L.) Planch.: E32121

Vitis vulpina L.: E31993

ACKNOWLEDGMENTS

The authors thank the Illinois Department of Natural Resources for access to the natural area inventory data for the seep; Tom Lerczak, Illinois Nature Preserves Commission, for his help and advice, the land owner for permission to enter the property, and the two anonymous reviewers for their helpful comments.

received 10/1/06

accepted 5/6/07

LITERATURE CITED

Bailey, A.W., and C.E. Poulton. 1968. Plant communities and environmental relationships in a portion of the Tillamook burn, northwestern Oregon. Ecology 49:1-13.

Calsyn, D.E. 1995. Soil survey of Mason County, Illinois. Soil Report 146, University of Illinois Agricultural Experiment Station, Urbana. ix+211 pp.

Daubenmire, R. 1959. A canopy coverage method of vegetation analysis. Northwest Science 33:4364.

Ebinger, J.E. 1978. Vascular flora of hillside seeps in east-central Illinois. Transactions of the Illinois State Academy of Science 71:109-114.

Ebinger, J.E. and J.A. Bacone. 1980. Vegetation survey of hillside seeps at Turkey Run State Park. Proceedings of the Indiana Academy of Science 90:390-394.

Gates, F.C. 1912. A bog in central Illinois. Torreya 11:205-211.

Henry, R.D. and A.R. Scott. 1984. The wetland vascular flora of four seeps in McDonough County, Illinois. Phytologia 56:1-15.

Herkert, J.R. and J.E. Ebinger. 2002. editors. Endangered and threatened species of Illinois: Status and distribution. Volume 1. Plants. Illinois Endangered Species Protection Board, Springfield, Illinois. 161 pp.

Lerczak, T.V. 2000. Proposal to register Speckman-Stelter Woods in Mason County as an Illinois Land and Water Reserve. Illinois Nature Preserves Commission, Springfield, Illinois. ii+14 pp.

Mohlenbrock, R.H. 2002. Vascular flora of Illinois. Southern Illinois University Press, Carbondale, Illinois. x + 490 pp.

Mueller-Dombois, D. and H. Ellenberg. 1974. Aims and methods of vegetation ecology. John Wiley and Sons, New York.

Parker, H.M. and J.E. Ebinger. 1971. Ecological study of a hillside marsh in east-central Illinois. Transactions of the Illinois State Academy of Science 64:362-369.

Phipps, R. and J. Speer. 1958. A hillside marsh in east-central Illinois. Transactions of the Illinois State Academy of Science 51:37-42.

Schwegman, J.E. 1969. Vegetation of some seep springs in the Cretaceous Hills of southern Illinois. Masters Thesis, Southern Illinois University, Carbondale, Illinois.

Schwegman, J.E. 1973. Comprehensive plan for the Illinois nature preserves system. Part 2. The natural divisions of Illinois. Illinois Nature Preserves Commission, Rockford, Illinois. 32 pp.+map.

Transeau, E.N. 1903. On the geographic distribution and ecological relationships of the bog plant societies of North America. Botanical Gazette 36:401-420.

Voigt, J.W. and R.H. Mohlenbrock. 1964. Plant Communities of Southern Illinois. Southern Illinois University Press, Carbondale, Illinois. xviii+202 pp.

White, J. 1978. (editor) Illinois Natural Areas Inventory, Technical Report. Illinois Natural Areas Inventory, Urbana, Illinois. xix + 426 pp.

White, J., and M.H. Madany. 1978. Classification of natural communities in Illinois. Pp. 310-405 in Illinois Natural Areas Inventory. Technical report. (J. White, Editor). Illinois Natural Areas Inventory, Urbana, Illinois.

Willman, H.B. and J.C. Frye. 1970. Pleistocene stratigraphy of Illinois. Illinois State Geological Survey Bulletin 94:1-204.

(1) William E. McClain, (2) Loy R. Phillippe, and (2) John E. Ebinger

(1) Adjunct Research Associate in Botany Illinois State Museum, Springfield, Illinois 62701

(2) Illinois Natural History Survey, Champaign, Illinois 61820
Table 1. Average cover and importance value of ground layer
species encountered during the summer of 2006 in a sunny seep
and a shaded seep at White Oak Creek Woods Natural Area, Mason
County, Illinois.

 Sunny seep
 Mean
Species Freq. % Cover I.V.

Apios americana 63 33.88 31.5
Impatiens capensis 87 26.78 30.3
Leersia oryzoides 73 15.57 21.0
Decodon verticillatus 47 17.12 18.1
Saururus cernuus 67 8.35 15.4
Pilea pumila 63 6.12 13.4
Rosa palustris 23 9.27 9.5
Carex spp. 43 4.05 9.0
Peltandra virginica 23 8.08 8.7
Aster lanceolatus 37 3.42 7.6
Boehmeria cylindrica 30 2.02 5.7
Cicuta maculata 27 2.67 5.6
Symplocarpus foetidus 17 3.60 4.9
Lysimachia nummularia 10 4.67 4.4
Ribes americanum 13 3.50 4.3
Parthenocissus quinquefolia 13 1.38 2.9
Leersia virginica 13 1.12 2.7
Iris shrevei 7 0.52 1.3
Stachys tenuifolia 7 0.20 1.1
Vitis vulpina 3 0.50 0.8
Acer saccharinum 3 0.10 0.6
Cinna arundinacea 3 0.10 0.6
Eupatoriadelphus maculatus 3 0.10 0.6
Fraxinus nigra -- -- --
Alliaria petiolata -- -- --
Smilax tamnoides -- -- --
Antenoron virginianum -- -- --
Acer negundo -- -- --
Thelypteris palustris -- -- --
Celastrus scandens -- -- --

Totals 153.12 200.0
Average bare ground/litter 4.50

 Shaded seep
 Mean
Species Freq. % Cover I.V.

Apios americana -- -- --
Impatiens capensis 73 19.80 41.7
Leersia oryzoides -- -- --
Decodon verticillatus -- -- --
Saururus cernuus 97 14.65 40.7
Pilea pumila 20 0.75 5.6
Rosa palustris -- -- --
Carex spp. 3 0.10 0.9
Peltandra virginica -- -- --
Aster lanceolatus 6 0.20 1.8
Boehmeria cylindrica 17 0.82 4.9
Cicuta maculata -- -- --
Symplocarpus foetidus 90 39.75 70.2
Lysimachia nummularia -- -- --
Ribes americanum 3 0.02 0.8
Parthenocissus quinquefolia 43 2.65 13.4
Leersia virginica 10 0.22 2.5
Iris shrevei -- -- --
Stachys tenuifolia -- -- --
Vitis vulpina 3 0.50 1.4
Acer saccharinum -- -- --
Cinna arundinacea 3 0.10 0.9
Eupatoriadelphus maculatus -- -- --
Fraxinus nigra 30 0.15 7.2
Alliaria petiolata 10 0.22 2.5
Smilax tamnoides 6 0.12 1.7
Antenoron virginianum 3 0.50 1.4
Acer negundo 3 0.02 0.8
Thelypteris palustris 3 0.02 0.8
Celastrus scandens 3 0.02 0.8

Totals 80.61 200.0
Average bare ground/litter 39.43
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Author:McClain, William E.; Phillippe, Loy R.; Ebinger, John E.
Publication:Transactions of the Illinois State Academy of Science
Article Type:Report
Geographic Code:1U3IL
Date:Jul 1, 2008
Words:3846
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