Printer Friendly

Remarks on the genus Necroscia (Phasmatodea, Diapheromeridae, Necrosciinae) from China, with two new records, one new synonym, and one new combination and key to the species.


There are nine Necroscia species recognized in China, among which N. inflata (Redtenbacher) and N. marginata (Gray) are reported for the first time from China. One new synonym is discovered: N. maculata (Chen & He) is regarded as a junior synonym of N. multicolor (Redtenbacher) n. syn. First description is given for the female and egg of N. hainanensis (Chen & He) n. comb., which is transferred from Sosibia Stal. A list and key to all Chinese species are given.

Key words

Necroscia, Necroscia inflata, Necroscia marginata, Necroscia hainanensis, new records, new synonym, new combination, China


Necroscia Audinet-Serville, 1838 is the second biggest genus of Necrosciinae Brenner von Wattenwyl, 1893 and contains 61 species in the tropical and warm subtropical regions of the Oriental region (Phasmida Species File Online by Brock). The number of Necroscia species is probably underestimated because of their canopy behavior, which causes difficulty collecting specimens.

In China, three Necroscia species are recognized in Hennemann et al. (2008: 17). Chen and He (2008) described several new taxa of Necroscia and Aruanoidea Brenner von Wattenwyl, 1893 in their monograph of Chinese phasmid. However, the latter one was a junior synonym to Necroscia (Hennemann 1998). Subsequently, eight species were recognized in Chen and He's monograph. There are also two other publications which involve the Chinese Necroscia, including a new species in Yunnan Province (Chen & Zhang 2008) and a newly discovered synonym (Ho 2010).

This study aims to provide taxonomic notes on the genus Necroscia in China, of which nine species are currently recognized in China. Two new species from China, N. inflata (Redtenbacher, 1908) and N. marginata (Gray, 1835), are reported for the first time. A new synonym and a new combination are proposed: N. maculata (Chen & He, 2000) is regarded as synonym of N. multicolor (Redtenbacher, 1908) n. syn.; and N. hainanensis (Chen & He, 2002) n. comb. is transferred from Sosibia Stal, 1875. The morphology of the female and structure of the egg of the latter species are firstly described and illustrated. A list and a key to Necroscia of China are provided in this paper to advance our understanding of this winged species in China.

Material and methods

The sequence of species account is alphabetical. Morphological terms largely follow Bragg (1997, 2001). Ootaxonomic description refers to Clark (1976a, 1976b, 1988, 1998) and Clark-Sellick (1997). Drawings of abdominal structure of Necroscia inflata (Redtenbacher) and N. marginata (Gray) are based on the Chinese specimens. Geographical distribution of each species provided in the checklist is based on Otte & Brock (2005), Chen & He (2008), Hennemann et al. (2008), and Phasmida Species File Online.

The materials mentioned in this paper are deposited in the following collections: Natural History Museum, London, England (BMNH); China Agricultural University, Beijing, China (CAU); Institut Royal des Sciences Naturelles de Belgique, Brussels, Belgium (ISNB); Institute of Zoology, Chinese Academy of Sciences, Beijing, China (IZCAS); Naturhistorisches Museum, Vienna, Austria (NHMW); Shanghai Entomological Museum, China Academy of Sciences, Shanghai, China (SEM); Shanghai Normal University, Shanghai, China (SNU); Zoologisches Museum der Universitat, Hamburg, Germany (ZMUH); Museum of Biology, Zhongshan (Sun Yat-Sen) University, Guangzhou, Guangdong, China (ZSU); and the private collection of Ho, G.W.G., Hong Kong, China (GH).


Necroscia Audinet-Serville, 1838

Type species.--Necroscia roseipennis Audinet-Serville, 1838: 250, by subsequent designation of Kirby, 1904: 436.

= Aruanoidea Brunner von Wattenwyl, 1893 [Synonymised by Hennemann, 1998: 121].

Notes.--Nine species are currently recognized in China. Two newly described taxa, Aruanoidea pseudocerca Chen & He, 2008 [= Necroscia pseudocerca] and Aruanoidea bidentata Chen & He, 2008 [= Necroscia bidentata] are misplaced and not discussed in this study. A publication is in progress to deal with these species (He in prep.). The specimens of Necroscia inflexipes (Olivier, 1792) are believed lost (Hennemann et al. 2008) and, therefore, are not included in the key.

Species recorded in China.--

1. Necroscia flavescens (Chen & Wang, 1998) [China (Guangxi) & Vietnam].

2. Necroscia hainanensis (Chen & He, 2002) n. comb. [China (Hainan)].

3. Necroscia inflata (Redtenbacher, 1908) [China, Singapore, Peninsular Malaysia and Sumatra].

4. Necroscia inflexipes (Olivier, 1792) [China]. = Phasma curvipes Stoll, 1813. = Phasma valgum Lichtenstein, 1796.

5. Necroscia marginata (Gray; 1835) [China (Yunnan), Singapore, Peninsular Malaysia and India].

6. Necroscia mista (Chen & He, 2008) [China (Guangdong)].

7. Necroscia multicolor (Redtenbacher, 1908) [China (Guangxi & Guangdong) & Vietnam]. = Necroscia maculata (Chen & He, 2000) n. syn.

8. Necroscia notata (Chen N Zheng, 2008) [China (Yunnan)].

9. Necroscia shukayi (Bi, Zhang & Lau, 2001) [China (Guangdong & Hong Kong)]. = Necroscia ovata Chen & He, 2008.

Key to females of the Chinese Necroscia

1. Pronotum with spines N. notata --Pronotum lacking spine, only granulated or smooth 2

2. Anal region of alae with mottled spots 3 --Anal region of alae lacking mottled spots, uniformly colored 4

3. Metafemora and metatibiae armed with fin-like lamellae N. hainanensis --Metafemora and metatibiae lacking armature N. shukayi

4. Body yellowish 5 --Body greenish 6

5. Head with ocelli N. flavescens --Head lacking ocellus N. multicolor

6. Tegmina with narrow margin in green, length of cerci distinctly long than the anal segment, body length 62-80 mm.... N. inflata --Tegmina with narrow margin in yellow, length of cerci as long as or shorter than the anal segment, body length 54-64 mm N. marginata

Key to males of the Chinese Necroscia

1. Mesonotum with spine-like tubercles N. mista --Mesonotum lacking spine-like tubercle 2

2. Anal region of alae with mottled spots 3-- Anal region of alae lacking mottled spot, uniformly colored 4

3. Tegmina with narrow outer margin in creamy white N. shukayi --Tegmina with narrow outer margin in green N. hainanensis

4. Body yellowish 5 --Body greenish 6

5. Body slender, head with ocelli N. flavescens--Body stout, head lacking ocellus N. multicolor

6. Tegmina with narrow outer margin in yellow, body length 40-44 mm N. marginata --Tegmina with narrow outer margin in green, body length 50-59 mm N. inflata

Necroscia hainanensis (Chen & He, 2002) n. comb. (Figs 1-4, 12)

Sosibia hainanensis, Chen & He (in Chen et al.), 2002: 107, figs. 9 a-b. Holotype [male], Jianfengling, Hainan Province, China, 29.VI. 1981, Rao Shuqing (ZSU).

Hennemann et al., 2008: 40.

Chen & He, 2008: 196, figs 161 a-b.

Further material.--[male], Jianfengling, Hainan Province, China, 24.VII.1984, Hua Lizhong (ZSU); [male][female] and 5 eggs, Jianfengling, Hainan Province, China, 7.VI.2008, Ho, G.W.C. (GH).

Diagnostic features.--The female is well distinguished from other species by antero-ventral carinae of metafemora and metatibia armed with fin-like lamellae. The male is generally similar to the female, but is lacking lamella on metafemora and mesofemora. Grayish to purplish green body with darker markings, which is distinctive from other Necroscia.

Description.--Female: (Figs 1, 2, 12) Grayish to purplish green with darker markings. Head and thorax covered with granules and tubercles. Larger and more robust than male.

Head: Oval, longer thanwide, slightly shorter than pronotum. Vertex with two small oval elevations between eyes, anterior margin of the elevations black. Occiput gently convex, with yellowish tubercle-like granules. Median furrow distinct. Eyes big, oval and protruding, dark brown. Ocelli distinct, placed above eyes, green and oblong. Antennae very long, longer than forelegs; the first segment 1.5x longer than second segment.

Thorax: Pronotum with few granules, parallel-sided, slightly longer than head, hind margin rounded. Mesonotum 2.5x longer than pronotum, shorter than mesofemora, median carina distinct; surface sparsely covered with small granules and few tubercle-like granules with a short carina before half being laterad of lateral margins. Mesopleurum set with four to five short spine-like tubercles. Mesosternum with yellowish granules. Metapleurum with oblong granules.

Abdomen: Cylindrical and tapering posteriorly. Sixth to 8th tergites with two lateral carinae. Anal segment as long as 9th tergum, notched at hind margin. Supra-anal plate small, triangular. Subgenital plate boat-shaped, apex slightly notched, reaching end of the anal segment. Cerci long and straight, apices rounded, slightly shorter than anal segment.

Legs: Brownish to grayish green, with irregular sizes of black mottling. Hindlegs as long as forelegs. All carinae of profemora, mesofemora, protibiae and mesotibiae smooth and unarmed. Antero-ventral carina of metafemora and metatibiae with irregular sizes of fin-like lamellae. Postero-ventral carina of metafemora smooth. Posteroventral carina of metatibiae slightly waved.

Wings: Tegmina dull green, oval, with blunt elevation, margin with a blackish stripe. Alae very long, reaching 7th tergum, costal region dull green to grayish green, with pale or green mottling, anal region buff with blackish spots.

Measurements (mm): Bodylength 79, head 5, antennae 65, pronoturn 5, mesonotum 10, metanotum 5, median segment 7, profemora 21, mesofemora 14, metafemora 23, protibiae 22.5, mesotibiae 15, metatibiae 24, tegmina 10.5, alae 46.

Eggs (Figs 3, 4): Capsule bluish green to dull green, bullet-shape, with wrinkles. Narrowing moderately towards posterior end, apex obtuse in dorsal view. Operculum flat, blackish green surrounded by dull green with wrinkles as capsule, central area elevated with a small granule-like capitulum, same coloration as capsule. Opercular collar thick and smooth. Micropylar plate oblong, anterior apex pointed while posterior apex slightly rounded. Measurements (mm): Length 5.4, width 1.7, height 1.5.

Notes.--Chen and He (in Chen et al. 2002) originally placed this species in Sosibia Stal, 1875. However, the curved inward hind margin of the anal segment in the female and the bullet-shaped egg capsule of this species match with the features of Necroscia rather than Sosibia (rounded hind margin of the anal segment in female and oval egg capsule, see Figs 5-7) and should, therefore, be transferred. The female and eggs are described for the first time.

Distribution.--Hainan (Jianfengling), China. Probably an endemic species to Hainan.

Necroscia inflata (Redtenbacher, 1908) (Figs 8-9, 13)

Aruanoidea inflata, Redtenbacher, 1908: 526. Synwpes 2[male][male], Palembang, Sumatra, G. Wober leg. ded., 11.IX.1891, 163-164 (ZMUH); Syntypes 2[female[female], Palembang, Sumatra, G. Wolber leg. ded., 11.IX.1891, 166-167 (ZMUH); Syntype [female], Deli, Palembang, Sumatra (ZMUH); Syntype [female], Nias Island, Sumatra, 1897, U.Raap, 'Mus. Genova' (NHMW); Syntype [female], Palambang, Sumatra (NHMW); Syntypes 2[male][male], Perak, Peninsular Malaysia, Staudinger (NHMW); Syntypes l [male], 5 [female][female], Kuala Kangsar, Perak, Peninsular Malaysia, 1902, Grubauer (NHMW); Syntypes [male][female], Perak, Peninsular Malaysia, lachau (NHMW); Syntype [male], Kelantan, Peninsular Malaysia, Rolle (NHMW). [Assessed by Phasmid Species File Online's images]

Necroscia inflata, Seow-Choen et al., 1994a: 11.

Seow-Choen et al., 1994b: 8.

Seow-Choen et al., 1994c: 394. [Note on food-plants].

Brock, 1995: 91.

Seow-Choen, 1997: 92. [Illustration].

Brock, 1998: 35.

Brock, 1999: 99. [Redescription and egg illustrations].

Seow-Choen, 1999: 299.

Seow-Choen, 2000: 23. [Illustration].

Zompro, 2002: 190.

Otte & Brock, 2005: 212.

Seow-Choen, 2005: 60. [Illustration].

Further material.--[female], 12000391 (SEM).

Diagnostic features.--The dark green thorax and costal region of alae lacking marking is separated from other similar species.

Description of both sexes.--Dark green species. Both sexes similar. Male smaller and slimmer than the female. Head square, almost as long as wide. Eyes dark brown. Mesonotum with small granulations. Female anal segment slightly longer than 9th tergum, slightly incised in center at hind margin. Female cerci stout and long (lacking right cercus in SEM specimen), cylindrical, apex rounded. Male cerci small, hidden beneath anal segment. Legs green as body. Tegmina green as body, elevated angle blunt and brown. Margin of costal region of alae light green, anal region pinkish with dusky areas, usually reaching 6th tergum or 7th tergum.

Distribution.--China, Singapore, Peninsular Malaysia (Perak and Kelantan) and Sumatra (Palembang and Nias Islands).

Notes.--The specimen in SEM was not labeled and was presumably collected from Yunnan or Guangxi among the neighboring Chinese specimens in the collection. With the similar distribution and occurrence in China of N. marginata [described in this study] and Trachythorax atrosignatus (Brunner von Wattenwyl, 1893) [recently reported from China in Chen & He 2008: 179], segregated distributions of this species between China and Peninsular Malaysia reflect the potential populations in Vietnam, Myanmar and Thailand. Redescription and egg illustrations were provided by Brock (1999: 99). The description is based on Brock (1999: 99) and the Chinese specimen.

Necroscia marginata (Gray, 1835) (Figs 10-11, 14)

Phasma marginatum, Gray, 1835: 23. Holotype [female], Malabar, India, catalog no 844793 (BMNH). [Assessed by Phasmid Species File Online's images].

Necroscia marginata, Westwood, 1859: 148.

Kirby, 1904: 375.

Brock, 1995: 90.

Brock, 1999: 93. [Egg illustration].

Seow-Choen, 2000: 22. [Illustration].

Otte & Brock, 2005: 213.

Seow-Choen, 2005: 54. [Illustration].

Aruanoidea marginata, Redtenbacher, 1908: 525.

Further material.--2 [female][female], Xishuangbanna, Yunnan Province, China, 8.VII.2003, Hu Jiayao & Tang Liang (SNU).

Diagnostic features.--This species is separated from other Necroscia by tegmina, and costal region of alae green with narrow margin in yellowish or reddish brown and anal region of alae rosy pink.

Description of both sexes.--Female body robust, male smaller than female. Head, thorax and wings green, abdomen brown. Head as wide as long. Genae with yellowish postocular band. Pronotum rectangular with yellowish or reddish brown margins. Mesonotum with small granules. Female anal segment truncate at hind margin. Female subgenital plate boat-shaped, apex notched and reaching end of the anal segment. Male anal segment incised in center. Cerci broadened at apex in both sexes. Legs reddish brown with dark stripes on all femora. Tegmina and costal region of alae green with narrow margin in yellowish brown (Chinese specimens) or reddish brown, followed by narrow white margin (indistinct in Chinese specimens). Mae usually reaching 7th tergum, anal region rosy pink.

Distribution.--Yunnan (Xishuangbanna), China. India [Type locality] and Peninsular Malaysia (Kedah, Pahang and Perak).

Notes.--This species is new to China and only found in Xishuang-banna of Yunnan. With the similar distribution and occurrence in China of N. inflata (described in this study) and Trachythorax atrosignatus (Brunner von Wattenwyl, 1893) (recently reported from China in Chen & He 2008: 179), segregated distributions of this species between the southwestern China and Peninsular Malaysia probably reflect the potential populations in Vietnam, Myanmar and Thailand. The description is based on Brock (1999: 93) and the two Chinese specimens. Coloration is assessed by the illustrations in Seow-Choen (2000: 22; 2005: 54). The coloration of the Chinese specimen is slightly different from the type, and the Malaysian specimen differs by its yellowish-brown margins of pronotum, mesonotum, tegmina and alae. Generally, the Chinese specimen is morphologically matched with this species.

Necroscia multicolor (Redtenbacher, 1908)

Aruanoidea multicolor, Redtenbacher, 1908: 523. Syntypes 5 [male][male], 2 [female][female], Than-Moi, Tonkin (Vietnam), Juni-Juli, Fruhstorfer, H. (NHMW); Syntypes 3 [male][male], Than-Moi, Tonkin (Vietnam), Fruhstorfer, H. (ISNB). [Assessed by Phasmid Species File Online's images] Brock, 1998: 44. Necroscia multicolor, Otte & Brock, 2005: 213.

= Aruanoidea incerta, Carl, 1913:54 [Synonymised by Gunther, 1935: 140].

= Aruanoidea maculata, Chen & He, 2000: 396, fig. 2 [Holotype [male], 240 m., Longgang, Longzhou Country, Guangxi Autonomous Region, China, 5.V.1982, Yang Chikun (CAU).] n. syn. Necroscia maculata, Otte & Brock, 2005: 213. Hennemann et al., 2008: 17. Chen & He, 2008: 114, figs. 79 a-b.

Further material.--[female], Xiangshuichangcun, Longzhou, Guangxi Autonomous Region, China, 11.VI.1980, Xi Fusheng (ZSU); [male][female], Nankunshan, Guangdong Province, China, 7.VI. 1983, Weng Yong (ZSU); 2[male][male], Heishiding, Fengkai, Guangdong Province, China, 5.VII. 1986, Tat Hong (ZSU); [male], Nankunshan, Guangdong Province, China, 9-14.VI.1987, Lu Yongjun (ZSU); 4 [male][male], 330-350 m., Longzhou, Guangxi Autonomous Region, China, 13-15.VI.2000, Li Wenzhu (IZCAS); 3 [male][male], 2 [female][female], Wuming, Guangxi Autonomous Region, China, 30-31.VII.2012, Ho, G.W.C. (GH).

Diagnostic features.--Yellowish green with bright yellow maculations and purple stripe on costal region of alae is distinctive from other species.

Description of both sexes.--Medium species. Both sexes similar, with charming coloration, eyes dark brown. Antennae bluish green, sparsely covered with long hairs, some segments with dark blue coloration ventrally. Mouthparts pale. Thorax with yellowish green granules and spine-like tubercles. Pronotum with two small spine-like tubercles. Mesonotum 3x longer than pronotum, lateral margins with seven to nine small spine-like tubercles. Abdomen dorsal surface yellowish green with bright yellow longitudinal stripe, and a purple spot medio-posteriorly. Ventral surface white. Female anal segment broadly emarginated at hind margin, lateral angles pointed. Female supra-anal plate semi-circular, median carina distinct. Female subgenital plate boat-shaped, incurved at end, slightly incised in center, projecting over the half of anal segment. Male poculum pointed at apex. Cerci very long, longer than anal segment, cylindrical and straight, apices rounded. Legs unarmed, with short setae, yellowish green. Metafemora with a purple band near apex. Metatibiae with two purple bands, one at base, another near apex. Tegmina oval, yellowish green with bright yellow maculations, elevated near middle, with u-shape purple marking and bright yellow spot laterad of the elevation. Alae reaching 6th tergum, yellowish green with bright yellow maculations, anal region pale white.

Notes.--Necroscia maculata (Chen & He, 2000), originally from Guangxi, is perfectly matched with Vietnamese Necroscia multicolor (Redtenbacher, 1908) and is no doubt a junior synonym of the latter species. Guangdong is a new distribution record for this species in China.

Distribution.--Guangxi (Longzhou and Wuming) and Guangdong (Nankunshan and Heishiding), China. Also Vietnam [Type locality].


I wish to deeply thank the Institute of Zoology, China Academy of Sciences, China Agricultural University, Shanghai Entomological Museum, Shanghai Normal University and Museum of Biology, and Zhongshan University for their kind assistance and access to the collections. I would also like to give thanks to Mrs Li Sze Wai for proofreading, and anonymous referees for commenting on the manuscript.


Audinet-Serville I.G.A. 1838 (1839). Histoire naturelle des Insectes. Orthopteres. Librairie Encyclopedique de Roret. 776 pp.

Bi D.Y., Zhang W.N., Lau C.S.K. 2001. Study on the walking stick insects (Phasmatodea) and the genus Sipyloidea from Hong Kong district of China. Entomotaxonomia 23: 253-258.

Bragg P.E. 1997. A glossary of terms used to describe phasmids. Phasmid Studies 6: 24-33.

Bragg P.E. 2001. Phasmids of Borneo. Natural History Publications, Kota Kinabalu, Borneo. 772 pp.

Brock P.D. Phasmida Species File Online. Version 2.1/3.4. Available from (accessed 5 March 2012).

Brock P.D. 1995. Catalogue of stick and leaf-insects associated with Peninsular Malaysia and Singapore. Malayan Nature Journal 49: 83-102.

Brock P.D. 1998. Catalogue of type specimens of stick & leaf-insects in Naturhistorisches Museum Wien (Insecta: Phasmida). Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museums in Wien 13: 1-75.

Brock P.D. 1999. Stick and Leaf Insects of Peninsular Malaysia and Singapore. Malaysian Nature Society, Kuala Lumpur. 223 pp.

Brenner von Wattenwyl K. 1893 Revision du Systeme des Orthopteres et description des especes rapportees par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale Giacomo Doria, Genova 13: 1-230.

Carl J. 1913. Phasmides nouveaux ou peu connus du Museum de Geneve. Revue Suisse de Zoologie 21: 1-56.

Chen S.C., He Y.H. 2000. Three new species of Phasmatodea from Shiwan Dashan, Guangxi (Phasmatodea: Heteronemiidae, Phasmatidae). Acta Zootaxonomica Sinica 25: 395-397.

Chen S.C., He Y.H. 2008. Phasmatodea of China. China Forestry Publishing House, Beijing. 476 pp.

Chen S.C., Wang J.J. 1998. A new species of Aruanoidea Brenner attacking forest from Guangxi (Phasmatodea: Heteronemiidae). Journal of Guangxi Academy of Sciences 14: 1-2.

Chen S.C., Zhang P.Y. 2008. Five new species of stick insects from Yunnan Province and description of male Cnipsomorpha colorantis (Chen & He) (Phasmatodea: Heteronemiidae, Phasmatidea). Entomotaxonomia 30: 245-254.

Chen S.C., He Y.H., Li Y. 2002. Phasmatodea, pp. 100-116. In: Huang F.S. (Ed) Forest insects of Hainan. Science Press, China.

Clark J.T. 1976a. The capitulum of phasmid eggs (Insecta: Phasmida). Zoological Journal of the Linnean Society, London 59:365-375.

Clark J.T. 1976b. The eggs of stick insects (Phasmida): a review with descriptions of the eggs of eleven species. Systematic Entomology 1 : 95-105.

Clark J.T. 1988. The capitula of phasmid eggs: an update with a review of the current state of phasmid ootaxonomy. Zoological Journal of the Linnean Society, London 93: 273-282.

Clark J.T. 1998. The micropylar plate of the eggs of Phasmida, with a survey of the range of plate form within the order. Systematic Entomology 23: 203-228.

Clark-Sellick J.T.C. 1997. The range of egg capsule morphology within the Phasmatodea and its relevance to the taxonomy of the order. Italian Journal of Zoology 64:97-104.

Gray G.R. 1835. Synopsis of the species of insects belonging to the family of Phasmidae. Longman, London. 48 pp.

Gunther K. 1935. Uber einige Phasmoiden aus der Sammlung des Herrn Dr. C. Willemse, Eijgelshoven. Natuurhistorisch Maandblad, Maastricht 24:123-126 & 138-140.

Hennemann F.H. 1998. Ein Beitrag zur Kenntnis der Phasmidenfauna von Sulawesi. Mit einem Katalog der bisher bekanntgewordenen Arten. Mitteilungen aus dem Museum fur Naturkunde Berlin, Zoologische Reihe 74: 95-128.

Hennemann F.H., Conle O.V., Zhang W.W. 2008. Catalogue of the Stick and Leaf-insects (Phasmatodea) of China, with a faunistic analysis, review of recent ecological and biological studies and bibliography (Insecta: Orthoptera: Phasmatodea). Zootaxa 1735: 1-76.

Ho G.W.C. 2010. Taxonomic note on Necroscia shukayi (Bi, Zhang & Lau 2001) comb. nov. (Phasmida: Necrosciinae: Necrosciini) and first description of its egg. Journal of Orthoptera Research 19: 1-5.

Ho G.W.C. 2012. Description of male and egg of Sosibia truncata Chen & Chen, 2000 (Phasmida: Diapheromeridae: Necrosdinae). Bulletin of Hong Kong Entomological Society 4: 38-40.

Kirby W.F. 1904. A synonymic catalogue of Orthoptera. I. Orthoptera Euplexoptera, Cursoria et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae). British Museum, London. 501 pp + ext. 25 pp.

Lichtenstein A.A.H. 1796. Catalogus Musei zoologici ditissimi Hamburgi, d III Februar 1796. Auctionis lege distrahendi 3: 1-222.

Olivier A.G. 1792. Mante, pp. 619-643. In: Encyclopedie Methodique, Histoire Naturelle, Insectes Vol. 7. Panckoucke, Paris.

Otte D., Brock P.D. 2005. Phasmida Species File. Catalog of stick and leaf insects of the world. The Insect Diversity Association and the Academy of Natural Sciences, Philadelphia. 414 pp.

Redtenbacher J. 1908. Die Insektenfamilie der Phasmiden. III. Phasmidae Anareolatae (Phibalosomini, Acrophyllini, Necrosciini). Wilhelm Engelmann, Leipzig. 248 pp.

Seow-Choen F. 1997. A guide to the stick and leaf insects of Singapore. Singapore Science Centre, Singapore. 160 pp.

Seow-Choen F. 1999. Stick and leaf insect (Phasmida: Insects) biodiversity in the nature reserves of Singapore. Garden's Bulletin Singapore 49:297-312.

Seow-Choen F. 2000. An illustrated guide to the stick and leaf insects of Peninsular Malaysia and Singapore. Natural History Publications, Borneo. 173 pp.

Seow-Choen F. 2005. Phasmids of Peninsular Malaysia and Singapore. Natural History Publications, Borneo. 120 pp.

Seow-Choen F., Brock P.D., Seow-En I. 1994a. The stick insects of Singapore. Singapore Scientist 70: 10-14.

Seow-Choen F., Brock, P.D., Seow-En I. 1994b. An introduction to the stick and leaf-insects of Singapore. Malaysian Naturalist 48: 7-11.

Seow-Choen F., Pin T.E., Brock RD., Seow-En I. 1994c. Foodplants of some stick-insects (Phasmida = Phasmatodea) from Singapore. Malayan Nature Journal 47: 393-396.

Stal C. 1875. Recensio orthopterorum. 3. Revue critique des Orthopteres decrits par Linne, DeGeer et Thunberg. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar 32: 1-105.

Stoll C. 1813. Representation des Spectres ou Phasmes, des Mantes, des Sauturelles, des Grillons, des Criquets et des Blattes des quatre Parties du Monde. L'Europe, L'Asia, L'Afrique et L'Amerique; ressemblees et descrites. Amsterdam.

Westwood J.O. 1859. Catalogue of the orthopterous in sects in the collection of the British Museum. Part 1. Phasmidae. British Museum, London. 196 pp.

Zompro O. 2002. Catalogue of type material of the insect order Phasmatodea at the Zoologisches Museum der Universitat Hamburg (Insecta: Orthoptera: Phasmatodea). Mitteilungen des Hamburger Zoologischen Museums und Instituts 99: 179-201.


Hong Kong Entomological Society. P.O. Box No. 73749, Kowloon Central Post Office, Hong Kong. Email:
COPYRIGHT 2013 The Orthopterists' Society
No portion of this article can be reproduced without the express written permission from the copyright holder.
Copyright 2013 Gale, Cengage Learning. All rights reserved.

Article Details
Printer friendly Cite/link Email Feedback
Author:Ho, George Wai-Chun
Publication:Journal of Orthoptera Research
Article Type:Report
Geographic Code:9CHIN
Date:Jan 1, 2013
Previous Article:A note on the molecular phylogeny of a small sample of Catantopine grasshoppers.
Next Article:A new species of Entoria Stal, 1875 from Hong Kong (Phasmatodea: Phasmatidae: Clitumninae).

Terms of use | Privacy policy | Copyright © 2022 Farlex, Inc. | Feedback | For webmasters |