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Marine biodiversity of Bahia Culebra, Guanacaste, Costa Rica: published records.

Bahia Culebra (10[degrees]36'N-85[degrees]39'W) is located on the north Pacific coast of Costa Rica. It is a relatively deep (30 m) and enclosed bay covering about 24 [km.sup.2]. This region of Costa Rica is undergoing the greatest coastal alteration due to tourism and other anthropongenic activities (Jimenez 2001, Fernandez-Garcia 2007). The bay is protected and deep enough to serve as an anchorage for US battleships between 1933 and 1937 (Sanchez -Noguera 2012). Based on his review of the history of marine research in Bahia Culebra by Cortes (2012a) the first period of active research in the area occurred between 1933 and 1938. During 1933 and 1934 the motor yacht Velero III on an expedition sponsored by the Allan Hancock Foundation, visited the bay (Fraser 1943). These expeditions have resulted in a significant number of publications and reports. In 1933, core samples were taken with the USS Hannibal, deposited at Scripps Oceanographic Institution, and the foraminifera analyzed (Bandy & Arnal 1957). The other important expedition was conducted by the New York Zoological Society as part of their eastern Pacific Expeditions. The steam yacht Taca, visited Bahia Culebra in 1938 (Beebe 1938) and resulted in a significant number of publications. Few studies subsequently occurred until the 1950's when individual researchers studied the bay's fauna and flora, e.g., Elmer Y. Dawson who collected algae (Dawson 1960).

Since the mid 1980's scientists from the Centro de Investigacion en Ciencias del Mar y Limnologia (CIMAR: Marine Science and Limnology Research Center), Universidad de Costa Rica, have been carrying out research on the marine environments of Bahia Culebra, mainly on the coral reefs (Cortes & Murillo 1985). This work has been later expanded by Carlos E. Jimenez in the 1990s (Jimenez 1998) and also included studies of the diversity of marine organisms. The contributions by Arturo Dominici-Arosemena and co-workers (2001) added an important number of fishes to the list of Bahia Culebra. Karina Rodriguez (Rodriguez-Saenz 2005, Rodriguez-Saenz & Gasca 2009, Rodriguez-Saenz & Segura-Puertas 2009) significantly increased the knowledge on the diversity of the gelatinous zooplankton of Bahia Culebra. Jorge Cabrera-Pena and his team from the Universidad Nacional, Heredia, Costa Rica, published three ecological papers in which marine species were mentioned (Madrigal-Castro et al. 1984, Cabrera-Pena et al. 1994, Cabrera-Pena & Solano-Lopez 1996). Angel Valdes and Yolanda E. Camacho-Garcia (2004) add seven species of gastropods to the list of species of Bahia Culebra.

As emphasized by Knowlton and Jackson (2008) baseline information is fundamental to evaluate changes. This is important for regions rich in coralline environments (Cortes & Jimenez 2003) with high marine biodiversity (this paper), and applies especially for areas that are being altered at a rapid rate such as Bahia Culebra. For this reason we have compiled this list of published records as a historical document and as baseline for other papers published both in this issue and in the future.

MATERIALS AND METHODS

We have compiled all reports of marine species and related topics dealing with Bahia Culebra (Cortes 2012b) and a list of all the species reported, organized by taxonomic group, has been organized. The numbers of species per taxa and the year they were reported were recorded. For the algae we used the classification and updated names proposed by Guiry and Guiry (2010). For marine animals we referred mostly to WoRMS (World Register of Marine Species, http://www.marinespecies.org) and ITIS (Integrated Taxonomic Information System, http://www.itis.gov).

RESULTS AND DISCUSSION

The published records of marine species of Bahia Culebra are listed in Table 1 together with their references. A total of 577 species have been documented, in 22 phyla (Table 1 and 2). The Crustacea had the most species reported (147 spp., mostly decapods: 94 spp.); followed by the cnidarians (100 spp., mostly hydroids: 75 spp.); mollusks (93 spp., mostly bivalves: 55 spp.); and chordates (92 spp., mostly fishes: 82 spp.) (Table 2). These numbers is a reflection of marine biodiversity in general (IUCN 2010) but also the interest and dedication of certain researchers such as is the case of the hydroids by Charles McLean Fraser and Karina Rodriguez-Saenz, or bivalves where most reports are by Leo G. Hertlein and Archibald M. Strong.

Most reports were published in the 1940's and 2000's (Fig. 1). The first pulse of publications resulted from the support of the Allan Hancock Foundation based on collections made aboard the Velero III (Fraser 1943), and the American Museum based on the collections of the New York Zoological Society Pacific expeditions aboard the Taca (Beebe 1938). The second pulse of publications has occurred in the last ten years based on investigations sponsored by the Universidad de Costa Rica through the Centro de Investigacion en Ciencias del Mar y Limnologia (CIMAR: Marine Science and Limnology Research Center) and the Museo de Zoologia (Zoology Museum).

Though known to be present in Bahia Culebra there are no published accounts for several groups of marine organisms, including the black corals, flatworms and nemerteans. Furthermore, conspicuos species such as marine turtles have been seem in the bay but not reported anywhere. Even important ecosystems, such as the soft bottom benthos as well as the rocky intertidal communities have not been studied and much more work needs to be done in the mangrove forests.

At present corals reefs, coral communities, sea urchins and some environmental parameters are being monitored by CIMAR, but other environments and organisms should be assessed periodically. The only way to determine changes due to anthropogenic and natural effects is through monitoring.

ACKNOWLEDGMENTS

We thank Cindy Fernandez for help with the algae, Juan Jose Alvarado with the echinoderms, Harlan Dean with the polychaetes. We appreciate the review of the manuscript by Richard Heard, Harlan Dean and Juan Jose Alvarado. Projects in Bahia Culebra have been funded over the years by the Vicerrectoria de Investigacion (Universidad de Costa Rica), Ecodesarrollo Papagayo, USAID-CDR program, and the Centro de Investigacion en Ciencias del Mar y Limnologia (CIMAR).

REFERENCES

Ajtai, P., Y. Camacho-Garcia & I.S. Wehrtmann. 2003. Range extension of Crosslandia daedali (Nudibranchia: Scyllaeidae). Rev. Biol. Trop. 51: 274.

Alpermann, T.J. 2001. The fisheries of ornamental fishes in Guanacaste, Costa Rica with special emphasis on the population dynamics of the Cortez Rainbow wrasse, Thalassoma lucasanum (Gill 1863). Tesis de Maestria, ISATEC, Univ. Bremen. Bremen, Alemania. 84 p.

Alvarado. J.J. 2008. Seasonal occurrence and aggregation behavior of the sea urchin Astropyga pulvinata (Echinodermata: Echinoidea) in Bahia Culebra, Costa Rica. Pac. Sci. 62: 579-592.

Augener, H. 1922. Ueber litorale Polychaeten von Westindien. Sitz. Ges. Naturf. Freunde Berlin 1922 (3-5): 38-53.

Bandy, O.L. & R.E. Arnal. 1957. Distribution of Recent Foraminifera off west coast of Central America. Bull. Amer. Assoc. Petrol. Geol. 41: 2037-2053.

Barnard, J.L. 1954. Amphipoda of the family Ampelisceidae collected in the eastern Pacific by the Velero III and Velero IV. Allan Hancock Pac. Exped. 18: 1-137.

Barnard, J.L. & C.M. Barnard. 1982. Revision of Foxiphalus and Eobrolgus (Crustacea: Amphipoda: Phoxocephalidae) from American oceans. Smithson. Contr. Zool. 372: 1-35.

Bednarski, M.K. 2001. Macrozooplankton of Culebra Bay, Costa Rica, with emphasis on copepods. M.Sc. thesis, ISATEC, Univ. Bremen. Bremen, Germany. 155 p.

Bednarski, M. & A. Morales-Ramirez. 2004. Composition, abundance and distribution of macrozooplankton in Culebra Bay, Gulf of Papagayo, Pacific coast of Costa Rica and its value as bioindicator of pollution. Rev. Biol. Trop. 52 (Suppl. 2): 105-118.

Beebe, W. 1938. Eastern Pacific expeditions of the New York Zoological Society, XIV. Introduction, itinerary, list of stations, nets and dredges of the eastern Pacific Zaca expedition, 1937-1938. Zoologica 23: 287-298.

Beebe, W. 1942. Eastern Pacific expeditions of the New York Zoological Society, XXX. Atlantic and Pacific fishes of the genus Dixonina. Zoologica 27: 43-48.

Berry, S.S. 1958. Notices of new eastern Pacific Mollusca --II. Leafl. Malacol. 1: 83-90.

Breedy, O. 2009. Octocorals. Text: Pp. 161-167, Species list: CD Pp. 108-111. In: I.S. Wehrtmann and J. Cortes (Eds.). Marine Biodiversity of Costa Rica, Central America. Springe, Berlin.

Breedy, O. & H.M. Guzman. 2003. Octocorals from Costa Rica: the genus Pacifigorgia (Coelenterata: Octocorallia: Gorgoniidae). Zootaxa 281: 1-60.

Breedy, O. & H.M. Guzman. 2005. A new species of Leptogorgia (Coelenterata: Ocotocorallia: Gorgoniidae) from the shallow waters of the eastern Pacific. Zootaxa 899: 1-11.

Breedy, O. & H.M. Guzman. 2007. A revision of the genus Leptogorgia Milne Edwards & Haime, 1857 (Coelenterata: Octocorallia: Gorgoniidae) in the eastern Pacific. Zootaxa 1407: 1-90.

Breedy, O. & H.M. Guzman. 2011. A revision of the genus Heterogorgia Verrill, 1868 (Coelenterata: Octocora llia: Plexauridae. Zootaxa 2995: 27-44.

Brusca, R.C. & E.W. Iverson. 1985. A guide to the marine isopod Crustacea of Pacific Costa Rica. Rev. Biol. Trop. 33 (Suppl. 1): 1-77.

Bussing, W.A. & M. Lopez. 2009. Marine fish. Text: Pp. 453-458, Species list: CD Pp. 412-473. In: I.S. Wehrtmann and J. Cortes (Eds.). Marine Biodiversity of Costa Rica, Central America. Springer, Berlin.

Cabrera-Pena, J. & Y. Solano-Lopez. 1996. Tamanos y frecuencia de Pontonia margarita (Crustacea: Palaemonidae) asociada a Pinctada mazatlanica (Bivalvia: Pteriidae), Costa Rica. Rev. Biol. Trop. 44: 915-917.

Cabrera-Pena, J., F. Vives-Jimenez & Y. Solano-Lopez. 1994. Tamanos y proporcion sexual de Ucides occidentalis (Crustacea: Gecarcinidae) en el manglar un manglar de Costa Rica. Uniciencia 11: 97-99.

Castellanos, I.A., A. Morales-Ramirez & E. Suarez-Morales. 2009. Appendicularians (Urochordata). Texto: Pp. 445-452, Lista de especies: Disco Compacto Pp. 411. In: I.S. Wehrtmann and J. Cortes (Eds.). Marine Biodiversity of Costa Rica, Central America. Springer, Berlin.

Castro, P. 1996. Eastern Pacific species of Trapezia (Crustacea, Brachyura: Trapeziidae), sibling species symbiotic with reef corals. Bull. Mar. Sci. 58: 531-554.

Cernohorsky, W.O. 1976. The Mitridae of the world, Part I: The Subfamily Mitrinae. Indo-Pac. Mollusca 3(17): 273-528.

Child, C.A. 1979. Shallow-water Pycnogonida of the Isthmus of Panama and the coast of Middle America. Smithsonian Contr. Zool. 293: 1-86.

Clark, H.L. 1940. Eastern Pacific expeditions of the New York Zoological Society. XXI. Notes on Echinoderms from the West Coast of Central America. Zoologica 25: 331-352.

Clark, H.L. 1948. A report on the Echini of the warmer eastern Pacific based on the collections of the Velero III. Allan Hancock Pac Exped. 8: 225-352.

Cortes, J. 2001. Requiem for an eastern Pacific seagrass bed, Bahia Culebra, Costa Rica. Rev. Biol. Trop. 49 (Suppl. 2): 273-278.

Cortes, J. 2012a. Historia de la investigacion marina en Bahia Culebra, Guanacaste, Costa Rica. Rev. Biol. Trop. 60 (Supl. 2): 19-37.

Cortes, J. 2012b. Bibliografia anotada sobre organismos, ambientes y procesos marinos en Bahia Culebra, Guanacaste, Costa Rica. Rev. Biol. Trop. 60 (Supl. 2): 231-242.

Cortes, J. & H.M. Guzman. 1998. Organismos de los arrecifes coralinos de Costa Rica: Descripcion, distribucion geografica e historia natural de los corales zooxantelados (Anthozoa: Scleractinia) del Pacifico. Rev. Biol. Trop. 46: 55-91.

Cortes, J. & C.E. Jimenez. 2003. Corals and coral reefs of the Pacific of Costa Rica: history, research and status: 361-385. In: J. Cortes (Ed.). Latin American Coral Reefs. Elsevier, Amsterdam.

Cortes, J. & M.M. Murillo. 1985. Comunidades coralinas y arrecifes del Pacifico de Costa Rica. Rev. Biol. Trop. 33: 197-202.

Crane, J. 1941a. Eastern Pacific Expeditions of the New York Zoological Society. XXVI. Crabs of the Genus Uca from the west coast of Central America. Zoologica 26: 145-208.

Crane, J. 1941b. Eastern Pacific Expeditions of the New York Zoological Society. XXIX. On the growth and ecology of Brachyuran crabs of the Genus Ocypode. Zoologica 26: 297-310.

Crane, J. 1947. Eastern Pacific Expeditions of the New York Zoological Society. XXXVIII. Intertidal brachygnathous crabs from the west coast of tropical America with special reference to ecology. Zoologica 32: 69-95.

Crane, J. 1975. Fiddler Crabs of the World. Ocypodidae: Genus Uca. Princeton Univ. Press, Princeton, New Jersey. 738 p.

Crouch, R.W. & C.W. Poag. 1987. Benthic Foraminifera of the Panamanian Province: distribution and origins. J. Foram. Res. 17: 153-176.

Cushman, J.A. & I. McCulloch. 1939. A report on some arenaceous Foraminifera. Allan Hancock Pac. Exped. 6: 1-113.

Cushman, J.A. & I. McCulloch. 1940. Some Nonionidae in the Collections of the Allan Hancock Foundation. Allan Hancock Pac. Exped. 6: 145-178.

Cushman, J.A. & I. McCulloch. 1942. Some Virgulininae in the Collections of the Allan Hancock Foundation. Allan Hancock Pac. Exped. 6: 179-230.

Cushman, J.A. & I. McCulloch. 1948. The species of Bulimina and related genera in the collections of the Allan Hancock Foundation. Allan Hancock Pac. Exped. 6: 231-294.

Dawson, E.Y. 1957. Marine algae from the Pacific Costa Rican gulfs. Los Angeles County Mus. Contr. Sci. 15: 1-28.

Dawson, E.Y. 1960. New records of marine algae from Pacific Mexico and Central America. Pac. Nat. 1(20): 31-52.

Dawson, E.Y. 1961. A guide to the literature and distributions of Pacific benthic algae from Alaska to the Galapagos Islands. Pac. Sci. 15: 370-461.

Dawson, E.Y. & P.T. Beaudette. 1959. Field notes from the 1959 eastern Pacific cruise of the Stella Polaris. Pac. Nat. 1(13): 1-24.

Dean, H.K. 2004. Marine biodiversity of Costa Rica: Class Polychaeta (Annelida). Rev. Biol. Trop. 52 (Suppl. 2): 131-181.

Deichmann, E. 1958. The Holothurioidea collected by the Velero III and IV during the years 1932 to 1954. Part II. Aspidochirota. Allan Hancock Pac. Exped. 11: 253-349.

Dominici-Arosemena, A., E. Brugnoli-Olivera, J. CortesNunez, H. Molina-Urena & M. Quesada-Alpizar. 2005. Community structure of eastern Pacific reef fishes (Gulf of Papagayo, Costa Rica). Tecnociencia 7(2): 19-41.

Drouet, F. 1936. Myxophyceae of the G. Allan Hancock Expedition of 1934, collected by Wm. R. Taylor. Allan Hancock Pac. Exp. 3: 15-32.

Fernandez-Garcia, C. 2007. Propagacion del alga Caulerpa sertularioides (Chlorophyta) en Bahia Culebra, Golfo de Papagayo, Pacifico norte de Costa Rica. M.Sc. thesis, Universidad de Costa Rica, San Jose, Costa Rica. 92 p.

Fernandez, C. & J.J. Alvarado. 2008. Chlorophyta de la costa Pacifica de Costa Rica. Rev. Biol. Trop. 56 (Suppl. 4): 149-162.

Fernandez, C. & J. Cortes. 2005. Reef Site: Caulerpa sertularioides, a green alga spreading aggressively over coral reef communities in Culebra Bay, North Pacific of Costa Rica. Coral Reefs 24: 10.

Fraser, C.M. 1938. Hydroids of the 1934 Hancock Pacific Expeditions. Allan Hancock Pac. Exped. 4: 1-105.

Fraser, C.M. 1943. General account of the scientific work of the Velero III in the eastern Pacific, 1931-1941, Part II: Geographic and biological associations. Allan Hancock Pac. Exped. 1: 49-258.

Fraser, C.M. 1948. Hydroids of the Allan Hancock Pacific Expeditions since March, 1938. Allan Hancock Pac. Exped. 4: 179-335.

Fritzsche, R.A. 1980. Revision of the Eastern Pacific Syngnathidae (Piscesisyngnathiformes), including both Recent and fossil forms. Proc. California Acad. Sci. 42: 181-227.

Garth, J.S. 1940. Some new species of Brachyuran crabs from Mexico and the Central and South American mainland. Allan Hancock Pac. Exped. 5: 53-127.

Garth, J.S. 1958. Brachyura of the Pacific coast of America. Oxyrhyncha. Tables and Plates. Allan Hancock Pac. Exped. 21: 501-854.

Garth, J.S. 1959. Eastern Pacific Expeditions of the New York Zoological Society. XLIV. Non-intertidial brachygnathous crabs from the west coast of Tropical America. Part 1: Brachygnatha, Oxyrhyncha. Zoologica 44: 105-126.

Garth, J.S. 1961. Eastern Pacific Expeditions of the New York Zoological Society. XLV. Non-intertidial brachygnathous crabs from the west coast of Tropical America. Part 2: Brachygnatha Brachyrhyncha. Zoologica 46: 133-160.

Garth, J.S. 1966. Eastern Pacific Expeditions of the New York Zoological Society. XLVI. Oxystomatous and allied crabs from the west coast of Tropical America. Zoologica 51: 1-16.

Glynn, P.W., E.M. Druffel & R.B. Dunbar. 1983. A dead Central American coral reef tract: possible link with the Little Ice Age. J. Mar. Res. 41: 605-637.

Gore, R. 1982. Porcellanid crabs from the coasts of Mexico and Central America (Crustacea: Decapoda: Anomura). Smithsonian Contr. Zool. 363: 1-34.

Gravel, P., K. Johanning, J. McLachlan, J.A. Vargas & E. Oberdorster. 2006. Imposex in the intertidal snail Thais brevidentata (Gastropoda: Muricidae) from the Pacific coast of Costa Rica. Rev. Biol. Trop. 54 (Suppl. 1): 21-26.

Guiry, M.D. & Guiry, G.M. 2010. AlgaeBase. World-wide electronic publication, National University of Ireland, Galway. http://www.algaebase.org; searched on 23 August 2010.

Haig, J. 1960. The Porcellanidae (Crustacea: Anomura) of the eastern Pacific. Allan Hancock Pac. Exped. 24: 1-440.

Haig, J. 1968. Eastern Pacific expeditions of the New York Zoological Society. Porcellanid crabs (Crustacea: Anomura) from the west coast of Tropical America. Zoologica 53: 57-74.

Hanna, G.D. & A.M. Strong. 1949. West American mollusks of the genus Conus. Proc. Calif. Acad. Sci., 4th ser., 26: 247-322.

Hartman, O. 1940. Polychaetous Annelids. Part II. Chrysopetalidae to Goniadidae. Allan Hancock Pac. Exped. 7: 173-287.

Hartman, O. 1944a. Polychaetous annelids. Pt. VI. Paraonidae, Magelonidae, Longosomidae, Ctenodrilidae and Sabellariidae. Allan Hancock Pac. Exped. 10: 311-389.

Hartman, O. 1944b. Polychaetous annelids. Pt. V. Eunicea. Allan Hancock Pac. Exped. 10: 1-238.

Hendrickx, M.E. & M.K. Wicksten. 1989. Los Pandalidae (Crustacea: Caridea) del Pacifico mexicano, con una clave para su identificacion. Caldasia 16: 71-86.

Henry, D.P. & P.A. McLaughlin. 1967. A revision of the subgenus Solidabalanus Hoek (Cirripedia, Thoracica) including a description of a new species with complemental males. Crustaceana 12: 43-58.

Henry, D.P. & P.A. McLaughlin. 1975. The barnacles of the Balanus amphitrite complex (Cirripedia, Thoracica). Zool. Verh. 141: 1-254.

Hertlein, L.G. & A.M. Strong. 1943. Eastern Pacific Expeditions of the New York Zoological Society. XXXII. Mollusks from the West Coast of Mexico and Central America. Part II. Zoologica 28: 149-168.

Hertlein, L.G. & A.M. Strong. 1946a. Eastern Pacific Expeditions of the New York Zoological Society. XXXIV. Mollusks from the West Coast of Mexico and Central America. Part III. Zoologica 31: 53-76.

Hertlein, L.G. & A.M. Strong. 1946b. Eastern Pacific Expeditions of the New York Zoological Society. XXXV. Mollusks from the West Coast of Mexico and Central America. Part IV. Zoologica 31: 93-120.

Hertlein, L.G. & A.M. Strong. 1947. Eastern Pacific Expeditions of the New York Zoological Society. XXXVI. Mollusks from the West Coast of Mexico and Central America. Part V. Zoologica 31: 129-150.

Hertlein, L.G. & A.M. Strong. 1948. Eastern Pacific Expeditions of the New York Zoological Society. XXXIX. Mollusks from the West Coast of Mexico and Central America. Part VI. Zoologica 33: 163-198.

Hertlein, L.G. & A.M. Strong. 1949a. Eastern Pacific Expeditions of the New York Zoological Society. XL. Mollusks from the West Coast of Mexico and Central America. Part VII. Zoologica 34: 63-97.

Hertlein, L.G. & A.M. Strong. 1949b. Eastern Pacific Expeditions of the New York Zoological Society. XLI. Mollusks from the West Coast of Mexico and Central America. Part VIII. Zoologica 34: 239-258.

Hertlein, L.G. & A.M. Strong. 1950. Eastern Pacific Expeditions of the New York Zoological Society. XLII. Mollusks from the West Coast of Mexico and Central America. Part IX. Zoologica 35: 217-252.

Hertlein, L.G. & A.M. Strong. 1951. Eastern Pacific Expeditions of the New York Zoological Society. XLIII. Mollusks from the West Coast of Mexico and Central America. Part X. Zoologica 36: 66-120.

Hoberg, E.P., D.R. Brooks, H. Molina-Urena & E. Erbe. 1998. Echinocephalus janzeni n.sp. (Nematoda: Gnathostomatidae) in Himantura pacifica (Chondrichthyes: Myliobatiformes) from the Pacific coast of Costa Rica and Mexico, with historial biogeographic analysis of the genus. J. Parasitol. 84: 571-581.

IUCN. 2010. IUCN Red List of Threatened Species. Version 2010.3. www.iucnredlist.org. Downloaded on 09 September 2010.

Jimenez, C.E. 1997. Corals and coral reefs of Culebra Bay, Pacific coast of Costa Rica: Anarchy in the reef. Proc. 8th Int. Coral Reef Symp., Panama 1: 329-334.

Jimenez, C.E. 1998. Arrecifes y comunidades coralinas de Bahia Culebra, Pacifico Norte de Costa Rica (Golfo de Papagayo). M.Sc. thesis, Universidad de Costa Rica, San Jose, Costa Rica. 218 p.

Jimenez, C. 2001. Arrecifes y ambientes coralinos de Bahia Culebra, Pacifico de Costa Rica: aspectos biologicos, economico-recreativos y de manejo. Rev. Biol. Trop. 49 (Suppl. 2): 215-231.

Jung, P. 1989. Revision of the Strombina-Group (Gastropoda: Columbellidae), fossil and living: distribution, biostratigraphy and systematics. Schweiz. Palaont. Abhandl. 111: 1-298.

Kelmo, F. & R. Vargas. 2002. Anthoathecatae and Leptothecatae hydroids from Costa Rica (Cnidaria: Hydrozoa). Rev. Biol. Trop. 50: 599-627.

Kim, W. & L.G. Abele. 1988. The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contr. Zool. 454: 1-119.

Knowlton, N. & J.B.C. Jackson. 2008. Shifting baselines, local impacts, and global change on coral reefs. PLoS Biology 6: 215-220.

Lalicker, C.G. & I. McCulloch. 1940. Some Textulariidae of the Pacific Ocean. Allan Hancock Pac. Exped. 6: 115-143.

Madrigal-Castro, E., J. Cabrera-Pena, J. Monge-Esquivel & F. Perez-Acuna. 1984. Comparacion entre dos poblaciones de Acanthina brevidentata (Gastropoda: Mollusca) en dos zonas rocosas de Playa Panama, Guanacaste, Costa Rica. Rev. Biol. Trop. 32: 11-15.

Manning, R.B. 1971. Eastern Pacific Expeditions of the New York Zoological Society. Stomatopod Crustacea. Zoologica 56: 95-113.

Manning, R.B. 1974. Stomatopods collected by Th. Mortensen in the eastern Pacific Region (Crustacea, Stomatopoda). Steenstrupia 3: 101-109.

Manter, H.W. 1940. Digenetic trematodes of fishes from the Galapagos Islands and the neighboring Pacific. Allan Hancock Pac. Exped. 2: 329-497.

Marques, F., D.R. Brooks & H. Molina-Urena. 1996. Two new species of Tetraphyllidean Cestodes in Himantura pacifica (Chondrichthyes: Myliobatiformes: Dasyatididae) from the northwest Coast of Costa Rica. J. Parasitol. 82: 302-306.

May-Collado, L. 2009. Marine mammals. Text: Pp. 479-495, Species list: CD Pp. 485-490. In: I.S. Wehrt mann and J. Cortes (Eds.). Marine Biodiversity of Costa Rica, Central America. Springer, Berlin.

May-Collado, L. & A. Morales-Ramirez. 2005. Presencia y patrones de comportamiento del delfin manchado costero, Stenella attenuata (Cetacea: Delhinidae) en el Golfo de Papagayo, Costa Rica. Rev. Biol. Trop. 53: 265-276.

May-Collado, L., T. Gerrodette, J. Calambokidis, K. Rasmussen & I. Sereg. 2005. Patterns of cetacean sighting distribution in the Pacific Exclusive Economic Zone of Costa Rica based on data collected from 1979-2001. Rev. Biol. Trop 53: 249-263.

McLaughlin, P.A. 1981. Revision of Pylopagurus and Tomopagurus (Crustacea: Decapoda: Paguridae), with the descriptions of new genera and species: Part II. Rhodochirus McLaughlin and Phimochirus McLaughlin. Bull. Mar. Sci. 31: 329-365.

McLean, J.H. & R. Poorman. 1971. New species of tropical eastern Pacific Turridae. Veliger 14: 89-113.

Meserve, F.G. 1938. Some monogenetic trematodes from the Galapagos Islands and the neighboring Pacific. Allan Hancock Pac. Exped. 2: 31-89.

Monks, S., F. Marques, V. Leon-Regagnon & G. PerezPonce de Leon. 1997. Koronacantha pectinaria n. comb. (Acanthocephala: Illiosentidae) from Microlepidotus brevipinnis (Haemulidae) and redescription of Tegorhynchus brevis. J. Parasitol. 83: 485-494.

Morales-Ramirez, A., R. Viquez, K. Rodriguez & M. Vargas. 2001. Marea roja producida por Lingulodinium polyedrum (Peridiniales, Dinophyceae) en Bahia Culebra, Golfo de Papagayo, Costa Rica. Rev. Biol. Trop. 49 (Suppl. 2): 19-23.

Osburn, R.C. 1950. Bryozoa of the Pacific Coast of America. Part 1, Cheilostomata-Anasca. Allan Hancock Pac. Exped. 14: 1-269.

Osburn, R.C. 1952. Bryozoa of the Pacific Coast of America. Part 2, Cheilostomata-Ascophora. Allan Hancock Pac. Exped. 14: 271-611.

Osburn, R.C. & J.D. Soule. 1953. Order Ectoprocta, Suborder Ctenostomata: Pp. 726-758 In: R.C. Osburn, Bryozoa of the Pacific Coast of America. Part 3, Cyclostomata, Ctenostomata, Entroprocta, and Addenda. Allan Hancock Pac. Exped. 14: 613-841.

Perez-Ponce de Leon, G., V. Leon-Regagnon & S. Monks. 1998. Theletrum lamothei sp. nov. (Digenea), parasite of Echidna nocturna from Cuajiniquil, Guanacaste, and other digenes of marine fishes from Costa Rica. Rev. Biol. Trop. 46: 345-354.

Pfeiler, E. 2008. Resurrection of the name Albula pacifica (Beebe, 1942) for the shafted bonefis (Albuniformes: Albulidae) from the eastern Pacific. Rev. Biol. Trop. 56: 839-844.

Rathbun, M.J. 1937. The Oxystomatous and allied crabs of America. U.S. Natl. Mus. Bull. 166: 1-272.

Reid, D.G. 2002. The genus Nodilittorina von Martens, 1897 (Gastropoda: Littorinidae) in the eastern Pacific Ocean, with a discussion of Biogeographic Provinces of the rocky-shore fauna. Veliger 45: 85-170.

Rodriguez-Ortiz, B., L. Garcia-Prieto & G. Perez-Ponce de Leon. 2004. Checklist of the helminth parasites of vertebrates in Costa Rica. Rev. Biol. Trop. 52: 313-354.

Rodriguez-Saenz, K. 2005. Distribucion espacial y temporal de la biomasa, composicion y abundancia del zooplancton, con enfasis en hidromedusas de Bahia Culebra durante La Nina 1999 y el 2000. M.Sc. thesis, Universidad de Costa Rica, San Jose, Costa Rica. 156 p.

Rodriguez-Saenz, K. & R. Gasca. 2009. Siphonophores. Text: Pp. 151-156, Species list: CD Pp. 101-104. In: I.S. Wehrtmann and J. Cortes (Eds.). Marine Biodiversity of Costa Rica, Central America. Springer, Berlin.

Rodriguez-Saenz, K. & J. Rodriguez-Fonseca. 2004. Avistamientos del delfin manchado, Stenella attenuata (Cetacea: Delphinidae) en Bahia Culebra, Costa Rica, 1999-2000. Rev. Biol. Trop. 52 (Suppl. 2): 189-193.

Rodriguez-Saenz, K. & L. Segura-Puertas. 2009. Hydrozoa, Scyphozoa, and Cubozoa (Medusozoa). Text: Pp. 143-149, Species list: CD Pp. 94-100. In: I.S. Wehrtmann and J. Cortes (Eds.). Marine Biodiversity of Costa Rica, Central America. Springer, Berlin.

Rost, H. 1955. A report on the Family Arcidae (Pelecypo da). Allan Hancock Pac. Exped. 20: 177-249.

Sanchez-Noguera, C. 2012. Entre historias y culebras: mas que una bahia (Bahia Culebra, Guanacaste, Costa Rica). Rev. Biol. Trop. 60 (Supl. 2): 000-000. Schmitt, W.L. 1940. The Stomatopods of the West Coast

of America, based on collections made by the Allan Hancock Expeditions, 1933-38. Allan Hancock Pac. Exped. 5: 129-225.

Skoglund, C. 2001. Panamic Province Molluscan Literature. Additions and changes from 1971 through 2000. I. Bivalvia. Festivus 32 (Supplement): 119 p.

Skoglund, C. 2002. Panamic Province Molluscan Literature. Additions and changes from 1971 through 2001. III Gastropoda. Festivus 33 (Supplement): 286 p.

Soot-Ryen, T. 1955. A report on the Family Mytilidae (Pelecypoda). Allan Hancock Pac. Exped. 20: 1-175.

Spongberg, A.L. 2006. PCB concentrations in intertidal sipunculan (Phylum Sipuncula) marine worms from the Pacific coast of Costa Rica. Rev. Biol. Trop. 54 (Suppl. 1): 27-33.

Suarez-Morales, E. & A. Morales-Ramirez. 2003. A new species of Cymbasoma (Crustacea: Copepoda: Monstrilloida) from the Pacific coast of Costa Rica, Central America. Proc. Biol. Soc. Wash. 116: 206-214.

Taylor, E.H. 1953. Early records of the seasnake Pelamis platurus in Latin America. Copeia 1953: 124.

Taylor, W.R. 1945. Pacific marine algae of the Allan Hancock Pacific Expeditions to the Galapagos Islands. Allan Hancock Pacific Exped. 12: 1-528.

Tsuda, R.T. 1968. Additional records of the marine algae from Costa Rica. Carib. J. Sci. 8: 103-104.

Tu, A.T. 1976. Investigation of the sea snake, Pelamis platurus (Reptilia, Serpentes, Hydrophiidae), on the Pacific coast of Costa Rica, Central America. J. Herpetol. 10: 13-18.

Turkay, M. 1970. Die Gecarcinidae Amerikas. Mit einem Anhang uber Ucides Rathbun. Sencken. Biol. 51: 333-354.

Valdes, A. & Y.E. Camacho-Garcia. 2004. "Cephalaspidean" Heterobranchs (Gastropoda) from the Pacific Coast of Costa Rica. Proc. California Acad. Sci. 55: 459-497.

Van Cleave, H.J. 1940. The Acanthocephala collected by the Allan Hancock Pacific Expedition, 1934. Allan Hancock Pac. Exped. 2: 501-527.

Vargas, R. & J. Cortes. 2006. Biodiversidad marina de Costa Rica: Crustacea: Infraorden Anomura. Rev. Biol. Trop. 54: 461-488.

Vargas-Montero, M. 2004. Floraciones algales en Costa Rica, su relacion con algunos factores meteorologicos y consideraciones sobre sus efectos sociecomicos. M.Sc. thesis, GIACT, Univ. Costa Rica. San Pedro, Costa Rica. 90 p.

Wehrtmann, I.S. & R. Vargas. 2003. New records and range extensions of shrimps (Decapoda: Penaeoidea, Caridea) from the Pacific and Caribbean coasts of Costa Rica, Central America. Rev. Biol. Trop. 51: 268-274.

Wicksten, M.K. 1983. Shallow water caridean shrimps of the Gulf of California, Mexico. Allan Hancock Monogr. Mar. Biol. 13: 1-59.

Wicksten, M.K. & M.E. Hendrickx. 1992. An updated checklist of benthic marine and brackish water shrimps (Decapoda: Penaeoidea, Stenopodidea, Caridea) from the Eastern Tropical Pacific: 49-76. In: M.E. Hendrickx (ed.), Contributions to the Study of East Pacific Crustaceans, 2 [Contribuciones al estudio de los Crustaceos del Pacifico Este, 2]. Instituto de Ciencias del Mar y Limnologia, UNAM.

Wilson, C.B. 1937. Parasitic copepods taken during the third Hancock expedition to the Galapagos Islands. Allan Hancock Pac. Exped. 2: 23-31.

Ziesenhenne, F.C. 1940. New ophiurans of the Allan Hancock Pacific Expeditions. Allan Hancock Pac. Exped. 8: 9-59.

Jorge Cortes (1, 2), Rita Vargas-Castillo (3) & Jaime Nivia-Ruiz1

(1.) Centro de Investigacion en Ciencias del Mar y Limnologia (CIMAR), Universidad de Costa Rica, San Pedro, 115012060 San Jose, Costa Rica; jorge.cortes@ucr.ac.cr

(2.) Escuela de Biologia, Universidad de Costa Rica, San Pedro, 11501-2060 San Jose, Costa Rica

(3.) Museo de Zoologia, Universidad de Costa Rica, San Pedro, 11501-2060 San Jose, Costa Rica; rita.vargas@ucr.ac.cr

Received 23-II-2011. Corrected 28-XI-2011. Accepted 15-II-2012.
TABLE 1

List of marine species reported in the literature for Bahia Culebra,
Guanacaste, Costa Rica

Clasification: Phylum              Species
(Subphylum): Class: Order
(Infraorder): Family

Cyanobacteria: Cyanophyceae:       Merismopoedia glauca
Chroococcales: Microcystaceae      (Ehrenberg) Kutzing, 1845

Cyanobacteria: Cyanophyceae:       Isactis plana (Harvey) Thuret
Nostocales: Rivulariaceae          ex Bornet & Flahault, 1886

Cyanobacteria: Cyanophyceae:       Blennothrix cantharidosma
Oscillatoriales:                   (Gomont)
Oscillatoriaceae

                                   Anagnostidis & Komarek, 1988
                                   as Hydrocoleum cantharidosmum

Cyanobacteria: Cyanophyceae:       Phormidium monile Setchell &
Oscillatoriales: Phormidiaceae     Gardner, 1930 as Lyngbya
                                   gracilis

                                   Trichodesmium erythraeum
                                   Ehrenberg ex Gomont 1892 Tas
                                   Trichodesmium erytraeum

Bacillariophyta:                   Pseudo-nitzschia pungens
Bacillariophyceae:                 (Grunow ex P.T. Cleve, 1897)
Bacillariales: Bacillariaceae      Hasle, 1993

                                   Pseudo-nitzschia spp.

Bacillariophyta:                   Guinardia sp.
Coscinodiscophyceae:
Rhizosoleniales:
Rhizosoleniaceae

Dinoflagellata: Dinophyceae:       Dinophysis caudata Saville-
Dinophysiales: Dinophyceae         Kent, 1881

Dinoflagellata: Dinophyceae:       Ceratium dens Ostenfeld &
Gonyaulacales: Ceratiaceae         Schmidt, 1859

                                   Ceratium furca (Ehrenberg,
                                   1833)

Dinoflagellata: Dinophyceae:       Alexandrium cf. ostenfeldii
Gonyaulacales: Goniodomataceae
                                   Alexandrium monilatum (Howell,
                                   1953)

Dinoflagellata: Dinophyceae:       Gonyaulax spinifera (Claparede
Gonyaulacales: Gonyaulacaceae      & Lachmann, 1859)

                                   Lingulodinium polyedrum
                                   (Stein, 1883)

Dinoflagellata: Dinophyceae:       Cochlodinium polykrikoides
Gymnodiniales: Gymnodiniaceae      Margalef, 1961

                                   Gymnodinium catenatum Grahm,
                                   1943

Dinoflagellata: Dinophyceae:       Noctiluca scintillans
Noctilucales: Noctilucaceae        (Macartney) Kofoid & Swezy,
                                   1921

Dinoflagellata: Dinophyceae:       Peridinium quinquecorne Abe,
Peridiniales: Peridinaceae         1927 in lit Protoperidinium
                                   quinquecrones

Dinoflagellata: Dinophyceae:       Prorocentrum lima (Ehrenberg)
Protocentrales: Protocentraceae    Dodge, 1975

                                   Prorocentrum micans Ehrenberg,
                                   1834

                                   Prorocentrum spp.

Chlorophyta: Bryopsidophyceae:     Caulerpa sertularioides (S.
Bryopsidales: Caulerpaceae         Gmelin) Howe, 1905

Chlorophyta: Bryopsidophyceae:     Halimeda discoidea Descaine,
Bryopsidales: Halimedaceae         1842

Chlorophyta: Ulvophyceae:          Rhizoclonium lubricum Setchell
Cladophorales: Cladophoraceae      & Gardner, 1819 as Lola
                                   lubrica

                                   Cladophora crystallina (Roth)
                                   Kutzing, 1843

Chlorophyta: Ulvophyceae:          Parvocaulis parvulus (Solms-
Dasycladales: Polyphysaceae        Laubach) Berger,

                                   Fettweiss, Gleissberg, Liddle,
                                   Richter, Sawitsky &
                                   Zuccarello, 2003

Chlorophyta: Ulvophyceae:          Cladophoropsis gracillima
Siphonocladales: Boodleacea        Dawson, 1950

                                   Phyllodictyon sp. as Struvea
                                   sp.

Chlorophyta: Ulvophyceae:          Ulva fasciata Delile, 1813
Ulvales: Ulvaceae
                                   Ulva flexuosa Wulfen, 1803 as
                                   Enteromorpha lingulata

                                   Ulva lactuca Linnaeus, 1753

Rhodophyta: Florideophyceae:       Acrochaetium infestans Howe &
Acrochaetiales:                    Hoyt, 1916
Acrochaetiaceae

Rhodophyta: Florideophyceae:       Ceramium avalonae Dawson, 1949
Ceramiales: Ceramiaceae
                                   Antithamnionella breviramosa
                                   (Dawson) Wollaston, 1970 as
                                   Antithamnion breviramosus

Rhodophyta: Florideophyceae:       Hypoglossum attenuatum var.
Ceramiales: Delesseriaceae         abyssicola (Taylor) Dawson,
                                   1962 as Hypoglossum
                                   abyssicolum

Rhodophyta: Florideophyceae:       Acanthophora spicifera
Ceramiales: Rhodomelaceae          (M.Vahl) B0rgesen, 1910 as
                                   Acanthophora specifera

                                   Chondria acrorhizophora
                                   Setchell & Gardner, 1924 as
                                   Chondria californica

                                   Chondria platyclada Taylor,
                                   1945

Rhodophyta: Florideophyceae:       Spyridia filamentosa (Wulfen)
Ceramiales: Spyridiaceae           Harvey, 1833

                                   Spyridia sp.

Rhodophyta: Florideophyceae:       Amphiroa sp.
Corallinales: Corallinaceae
                                   Jania tenella var. zacae
                                   Dawson, 1953

                                   Jania ungulata (Yendo) Yendo,
                                   1905

Rhodophyta: Florideophyceae:       Lithothamnion validum Foslie,
Corallinales: Hapalidiaceae        1906 as Lithothamnium validum

Rhodophyta: Florideophyceae:       Sarcodiotheca ecuadoreana
Gigartinales: Areschougiaceae      Taylor, 1945

                                   Sarcodiotheca furcata
                                   (Setchell & Gardner) Kylin,
                                   1925

Rhodophyta: Florideophyceae:       Gracilaria symmetrica Dawson,
Gracilariales: Gracilariaceae      1949

                                   Gracilaria tepocensis (Dawson)
                                   Dawson, 1940 as Gracilaria
                                   crockery

                                   Gracilariopsis costaricensis
                                   Dawson, 1949

                                   Gracilariopsis panamensis
                                   (Taylor) Dawson, 1949

Rhodophyta: Florideophyceae:       Cryptonemia angustata
Halymeniales: Halymeniaceae        (Setchell & Gardner) Dawson,
                                   1954

                                   Cryptonemia limensis (Kutzing)
                                   Lewis, 1990 in lit Cryptonemia
                                   guaymasensis

Rhodophyta: Florideophyceae:       Galaxaura filamentosa Taylor,
Nemaliales: Galaxauraceae          1945

Rhodophyta: Florideophyceae:       Liagora sp.
Nemaliales: Liagoraceae

Rhodophyta: Florideophyceae:       Scinaia complanata (Collins)
Nemaliales: Scinaiaceae            Cotton, 1907

                                   Scinaia confusa (Setchell)
                                   Huisman, 1985 as Gloiphloea
                                   confuse

                                   Scinaia johnstoniae Setchell,
                                   1914

Rhodophyta: Florideophyceae:       Lomentaria baileyana (Harvey)
Rhodymeniales: Lomentariaceae      Farlow, 1876

                                   Lomentaria hakodatensis Yendo,
                                   1920

Ochrophyta: Phaeophyceae:          Dictyota crenulata J. Agardh,
Dictyotales: Dictyotaceae          1847

                                   Dictyota flabellata (Collins)
                                   Setchell & Gardner, 1924

                                   Padina durvillei Bory
                                   Saint-Vincent, 1827

                                   Padina sp.

Ochrophyta: Phaeophyceae:          Feldmannia indica (Sonder)
Ectocarpales: Actinetosporaceae    Womersley & Bailey, 1970 as
                                   Ectocarpus indicus

Ochrophyta: Phaeophyceae:          Asteronema breviarticulatum
Scytothamnales: Scytothamnaceae    (J. Agardh) Ouriques & Bouzon,
                                   2000 as Ectocarpus
                                   breviarticulatus

Magnoliophyta: Liliopsida:         Ruppia maritima Linnaeus, 1753
Najadales: Ruppiaceae

Magnoliophyta: Liliopsida:         Halophila baillonis Ascherson,
Alismatales: Hydrocharitaceae      1874

Magnoliophyta: Magnoliopsida:      Avicennia sp.
Lamiales: Verbenaceae:

Magnoliophyta: Magnoliopsida:      Laguncularia sp.
Myrtales: Combretaceae

Magnoliophyta: Magnoliopsida:      Rhizophora sp.
Rhizophorales: Rhizophoraceae

Protozoa: Sarcodina:               Bolivina vaughani (Natland,
Foraminiferida: Bolivinitidae      1938)

                                   Brizalina pygmaea (Brady,
                                   1881) as Bolivina pygmaea

Protozoa: Sarcodina:               Buliminella parallela Cushman
Foraminiferida: Buliminidae        & Parker, 1931

                                   Bulimina pagoda var. denudata
                                   Cushman, 1927 as Bulimina
                                   denudata

Protozoa: Sarcodina:               Cibicides fletcheri Galloway &
Foraminiferida: Cibicididae        Wissler, 1927

Protozoa: Sarcodina:               Elphidium crispum (Linnaeus,
Foraminiferida: Elphidiidae        1758)

Protozoa: Sarcodina:               Eponides antillarum
Foraminiferida: Eponididae         (d'Orbigny, 1839)

Protozoa: Sarcodina:               Virgulina schreibersiana
Foraminiferida: Fusenkoinidae      Czjzek, 1848

Protozoa: Sarcodina:               Globigerina bulloides
Foraminiferida: Globigerinidae     D'Orbigny, 1826

                                   Globigerina subcretacea
                                   Chapman, 1902

                                   Globigerinella siphonifera
                                   (d'Orbigny, 1839) as
                                   Globigerinella aequilateralis

                                   Globigerinoides ruber
                                   (d'Orbigny, 1839) as
                                   Globigerinaoides rubra

Protozoa: Sarcodina:               Globorotalia menardii
Foraminiferida:                    (d'Orbigny, 1826)
Globorotaliidae

Protozoa: Sarcodina:               Haplophragmoides sp.
Foraminiferida:
Haplophragmoididae

Protozoa: Sarcodina:               Loxostomum limbatum (Brady,
Foraminiferida: Loxostomatidae     1881) as Loxostoma limbatum

Protozoa: Sarcodina:               Quinqueloculina dutemplei
Foraminiferida: Miliolidae         (d'Orbigny, 1846)

                                   Quinqueloculina lamarckiana
                                   (d'Orbigny, 1839)

                                   Massilina inaffecta (a)

                                   Massilina patula (a)

                                   Massilina robustior Cushman &
                                   Valentine, 1930

Protozoa: Sarcodina:               Nonionella atlantica (Cushman,
Foraminiferida: Nonionidae         1947)

Protozoa: Sarcodina:               Nouria polymorphinoides
Foraminiferida: Nouriidae          Heron-Allen & Earland, 1914

Protozoa: Sarcodina:               Planorbulina mediterranensis
Foraminiferida:                    (d'Orbigny, 1826)
Planorbulinidae

Protozoa: Sarcodina:               Reussella aequa Cushman &
Foraminiferida: Reussellidae       McCulloch, 1948

                                   Reussella pacifica Cushman &
                                   McCulloch, 1948

Protozoa: Sarcodina:               Rectobolivina hancocki
Foraminiferida:                    (Cushman & McCulloch, 1942)
Sphaeroidinidae

Protozoa: Sarcodina:               Spiroloculina dentata Cushman
Foraminiferida:                    & Tood, 1944
Spiroloculinidae

Protozoa: Sarcodina:               Poritextularia panamensis (a)
Foraminiferida: Textulariidae
                                   Textularia candeiana
                                   d'Orbigny, 1839

                                   Textularia foliacea Cushman,
                                   1922

                                   Textularia secasensis Lalicker
                                   & McCulloch, 1940

                                   Textularia vola Lalicker &
                                   McCulloch, 1940

Protozoa: Sarcodina:               Trochammina pacifica Cushman,
Foraminiferida: Trochamminidae     1925

Cnidaria: Anthozoa:                Carijoa sp. as Telesto sp.
Alcyonacea: Clavulariidae

Cnidaria: Anthozoa:                Leptogorgia alba Duchassaing &
Alcyonacea: Gorgoniidae            Michelotti, 1864

                                   Leptogorgia cofrini Breedy &
                                   Guzman, 2005

                                   Leptogorgia cuspidata Verrill,
                                   1865

                                   Pacifigorgia firma Breedy &
                                   Guzman, 2003

Cnidaria: Anthozoa:                Heterogorgia verrucosa
Alcyonacea: Plexauridae            Verrill, 1868

Cnidaria: Anthozoa:                Leptoseris papyracea (Dana,
Scleractinia: Agariciidae          1846)

                                   Pavona clavus (Dana, 1846)

                                   Pavona gigantea Verrill, 1869

                                   Pavona maldivensis (Gardiner,
                                   1905)

                                   Pavona varians Verrill, 1864

Cnidaria: Anthozoa:                Fungia (Cycloseris) curvata
Scleractinia: Fungiidae            (Verrill, 1870)

                                   Fungia (Cycloseris) distorta
                                   Michelin, 1842

Cnidaria: Anthozoa:                Pocillopora capitata Verrill,
Scleractinia: Pocilloporidae       1864

                                   Pocillopora damicornis
                                   (Linnaeus, 1758)

                                   Pocillopora elegans Dana, 1846

                                   Pocillopora eydouxi Milne
                                   Edwards & Haime, 1860

                                   Pocillopora meandrina Dana,
                                   1846

Cnidaria: Anthozoa:                Porites lobata Dana, 1840
Scleractinia: Poritidae
                                   Porites panamensis Verrill,
                                   1866

Cnidaria: Anthozoa:                Oulangia bradleyi (Verrill,
Scleractinia: Rhizangiidae         1866)

Cnidaria: Anthozoa:                Psammocora stellata (Verrill,
Scleractinia: Siderastreidae       1866)

                                   Psammocora superficialis
                                   Gardiner, 1898

Cnidaria: Scyphozoa:               Nausithoe punctata Kolliker,
Coronateae: Nausithoidae           1853

Cnidaria: Scyphozoa:               Pelagia noctiluca Peron and
Semaeostomae: Pelagiidae           Lesueur, 1810

Cnidaria: Hydrozoa:                Bougainvillia crassa Fraser,
Anthoathecata:                     1938
Bougainvilliidae

                                   Bouganvillea fulva Agassiz &
                                   Mayer, 1899

                                   Bouganvillea muscus Allmar,
                                   1863

                                   Garveia gracilis (Clark, 1876)
                                   as Bimeria gracilis

                                   Koellikerina fasciculata
                                   (Peron & Lesuerur, 1810)

                                   Koellikerina multicirrata
                                   (Kramp, 1928)

                                   Leuckartiara octona (Fleming,
                                   1823) as Perigonimus repens

Cnidaria: Hydrozoa:                Heterotiara minor Vanhoffen,
Anthoathecata: Bythotiaridae       1911

                                   Kanaka pelagica Uchida, 1947
                                   incerta sedis

Cnidaria: Hydrozoa:                Vannucia forbesii (Mayer,
Anthoathecata: Corymorphidae       1894)

Cnidaria: Hydrozoa:                Dipurena ophiogaster Haeckel,
Anthoathecata: Corynidae           1879

                                   Sarsia coccometra Bigelow,
                                   1909

                                   Sarsia spp.

Cnidaria: Hydrozoa:                Euphysa aurata Forbes, 1848
Anthoathecata: Euphysidae

                                   Euphysilla pyramidata Kramp,
                                   1955

Cnidaria: Hydrozoa:                Hydractinia carolinae Fraser,
Anthoathecata: Hydractiniidae      1912

                                   Hydractinia disjuncta Fraser,
                                   1938

Cnidaria: Hydrozoa:                Turritopsis nutricula McCrady,
Anthoathecata: Oceaniidae          1857

Cnidaria: Hydrozoa:                Amphinema australis (Mayer,
Anthoathecata: Pandeidae           1900)

                                   Amphinema dilema (Peron &
                                   Lesuerur, 1810)

                                   Haliotholus intermedius
                                   (Browne, 1902)

                                   Haliotholus pauper Hartlaub,
                                   1913

                                   Leukartiara hoepplii Hsu, 1928

                                   Leukartiara octona (Fleming,
                                   1823)

                                   Pandeopsis ikarii Kramp, 1959

Cnidaria: Hydrozoa:                Pennaria disticha (Goldfuss,
Anthoathecata: Pennariidae         Pennaria tiarella 1820) as

Cnidaria: Hydrozoa:                Porpita porpita (Linnaeus,
Anthoathecata: Porpitidae          1758)

Cnidaria: Hydrozoa:                Halitiara formosa Fewkes, 1882
Anthoathecata: Protiaridae

Cnidaria: Hydrozoa:                Trichydra pudica Wright, 1858
Anthoathecata: Trichydridae

Cnidaria: Hydrozoa:                Ectopleura dumortieri (van
Anthoathecata: Tubulariidae        Beneden, 1844)

Cnidaria: Hydrozoa:                Abylopsis eschscholtzi
Calycophorae: Abylidae             (Huxley, 1859)

                                   Abylopsis tetragona (Otto,
                                   1823)

Cnidaria: Hydrozoa:                Sulculeolaria chuni (Lens &
Calycophorae: Dyphidae             van Riemsdijk, 1908)

                                   Sulculeolaria quadrivalvis
                                   Blainville, 1834

                                   Chelophyes contorta (Lens &
                                   van Riemsdijk, 1908)

                                   Diphyes bojani (Eschscholtz,
                                   1829)

                                   Diphyes dispar Chamisso &
                                   Eysenhardt, 1821

                                   Eudoxoides mitra Huxley, 1859

                                   Muggiaea atlantica Cunningham,
                                   1892

Cnidaria: Hydrozoa:                Rosacea cymbiformis (Chiaje,
Calycophorae: Prayidae             1822)

                                   Rosacea flaccida Biggs, Pugh &
                                   Carre, 1978

Cnidaria: Hydrozoa:                Aequorea coerulescens (Brandt,
Leptothecata: Aequoridae           1838)

                                   Aequorea macrodactyla (Brandt,
                                   1834)

Cnidaria: Hydrozoa:                Clytia brunescens (Bigelow,
Leptothecata: Campanulariidae      1904)

                                   Clytia hemisphaerica
                                   (Linnaeus, 1767)

                                   Clytia simplex (Browne, 1902)

                                   Clytia uchidai Kramp, 1961

                                   Obelia spp.

Cnidaria: Hydrozoa:                Dipleurosoma pacificum Agassiz
Leptothecata:                      & Mayer, 1902
Dipleurosomatidae

Cnidaria: Hydrozoa:                Eutima curva Browne, 1905
Leptothecata: Eutimidae

                                   Eutima orientales (Browne,
                                   1905)

Cnidaria: Hydrozoa:                Halecium delicatulum
Leptothecata: Haleciidae           Coughtrey, 1876 as Halecium
                                   gracile

                                   Halecium nanum Alder, 1849

Cnidaria: Hydrozoa:                Laodicea marana Agassiz &
Leptothecata: Laodiceidae          Mayer, 1899

Cnidaria: Hydrozoa:                Eucheilota comata (Bigelow,
Leptothecata: Lovenellidae         1909)

                                   Eucheilota menoni Kramp, 1959

Cnidaria: Hydrozoa:                Octophiallucium bigelowi
Leptothecata: Malagazziidae        Kramp, 1955

Cnidaria: Hydrozoa:                Melicetum sp.
Leptothecata: Melicertidae

Cnidaria: Hydrozoa:                Olindias singulares Browne,
Limnomedusae: Olindiidae           1905

Cnidaria: Hydrozoa:                Phialella sp.
Leptothecata: Phialellidae

Cnidaria: Hydrozoa:                Lytocarpus phillippinus
Leptothecata: Plumulariidae        (Kirschenpauer, 1872)

Cnidaria: Hydrozoa:                Sertularia exigua Allman, 1877
Leptothecata: Sertulariidae
                                   Thuiaria crisiodes (Lamouroux,
                                   1824)

Cnidaria: Hydrozoa:                Solmundella bitentaculata
Narcomedusae: Aeginidae            (Quoy & Gaimard, 1833)

Cnidaria: Hydrozoa:                Cunina octonaria McCrady, 1859
Narcomedusae: Cuninidae

                                   Solmaris rhodoloma (Brandt,
                                   1838)

Cnidaria: Hydrozoa:                Agalma elegans (Sars, 1846)
Physonectae: Agalmidae

                                   Agalma okeni Eschscholtz, 1825

                                   Nanomia bijuga (Chiaje, 1841)

Cnidaria: Hydrozoa:                Forskalia edwarsi Kolliker,
Physonectae: Forskaliidae          1853

Cnidaria: Hydrozoa:                Liriope tetraphylla (Chamisso
Trachymedusae: Geryoniidae         & Eysenhardt, 1821)

Cnidaria: Hydrozoa:                Aglantha sp.
Trachymedusae:
Rhopalonematidae
                                   Aglantha digitale (Muller,
                                   1766)

                                   Aglaura hemistoma Peron &
                                   Lesuerur, 1810

                                   Rhopalonema velatum Gegenbaur,
                                   1857

Ctenophora                         Unidentified species

Platyhelminthes: Cestoda:          Rhinebothrium geminum Marques,
Rhinebothriidea:                   Brooks & Molina-Urena, 1996
Rhinobothriidae

Platyhelminthes: Cestoda:          Acanthobothroides pacificus
Tetraphyllidae:                    Marques, Brooks & Molina-
Onchobothriidae                    Urena, 1996

Platyhelminthes: Monogenea:        Neomicrocotyle pacifica
Polyopisthocotylea:                (Meserve, 1938) as
Protomicrocotylidae                Protomicrocotyle pacifica

Platyhelminthes: Trematoda:        Bucephalus introversus Manter,
Plagiorchiida: Bucephalidae        1940

Platyhelminthes: Trematoda:        Larval Didymozoidae not
Plagiorchiida: Didymozoidae        identified (metacercariae)

Platyhelminthes: Trematoda:        Tergestia laticollis
Plagiorchiida: Fellodistomidae     (Rudolphi, 1819)

Platyhelminthes: Trematoda:        Bianium simonei (Travassos,
                                   Freitas &

Plagiorchiida: Lepocreadiidae      Buhenheim, 1965)

Platyhelminthes: Trematoda:        Lasiotocus costaricae (Manter,
Plagiorchiida: Monorchiidae        1940) as Proctotrema
                                   costaricae

Platyhelminthes: Trematoda:        Helicometra sinuata (Rudolphi,
Plagiorchiida: Opecoelidae         1819)

Acanthocephala:                    Koronacantha pectinaria (Van
Palaeacanthocephala:               Cleave, 1940) as
Echinorhynchida: Illiosentidae     Microlepidotus brevipinnis

                                   Tegorhynchus pectinarius Van
                                   Cleave, 1940

Nematoda: Secernentea:             Echinocephalus janzeni Hoberg,
Spirurida: Gnathostomatidae        Brooks, Molina-Urena & Erbe,
                                   1998

Annelida: Polychaeta:              Chloeia pseudeuglochis
Amphinomida: Amphinomidae          Augener, 1924

                                   Eurythoe complanata (Pallas,
                                   1766)

Annelida: Polychaeta:              Lumbrineris latreilli Audouin
Eunicida: Lumbrineridae            & Milne Edwards, 1834

Annelida: Polychaeta:              Idanthyrsus cretus Chamberlin,
Sabellida: Sabellariidae           1919 as Idanthyrsus pennatus

Sipunculida:                       Antillesoma antillarum (Grube,
Phascolosomatidea:                 1858)

Phascolosomatida:                  Phascolosoma perlucens Baird,
Phascolosomatidae                  1868

Mollusca: Bivalvia: Arcoida:       Acar bailyi Bartsch, 1931
Arcidae
                                   Anadara concinna (Sowerby,
                                   1833) as Anadara (Scapharca)
                                   cumingiana

                                   Anadara formosa (Sowerby,
                                   1833) as Arca (Anadara)
                                   Formosa

                                   Anadara multicostata (Sowerby,
                                   1833) as Arca (Larkinia)
                                   multicostata

                                   Anadara (Cunearca) nux
                                   (Sowerby, 1833)

                                   Anadara similis (Adams, 1852)
                                   as Arca (Anadara) similis

                                   Anadara tuberculosa (Sowerby,
                                   1833) as Arca (Anadara)
                                   tuberculosa

                                   Arca mutabilis Sowerby, 1833
                                   as Arca (Arca) mutabilis

Mollusca: Bivalvia: Arcoida:       Glycymeris strigilata
Glycymerididae                     (Sowerby, 1833) as Glycymeris
                                   (Tuceta) tessellata

Mollusca: Bivalvia: Myoida:        Corbula (Caryocorbula) ovulata
Corbulidae                         Sowerby, 1833 as Aloidis
                                   (Caryocorbula) ovulate

Mollusca: Bivalvia: Myoida:        Amygdalum americanum
Mytilidae                          Soot-Ryen, 1955

                                   Crenella divaricata
                                   (d'Orbigny, 1842)

                                   Lioberus salvadoricus Hertlein
                                   & Strong, 1946

                                   Lithophaga (Diberus) plumula
                                   (Hanley, 1844)

                                   Lithophaga (Myoforceps)
                                   aristata (Dillwyn, 1817)

                                   Modiolus eiseni Hertlein &
                                   Strong, 1937

                                   Septifer zeteki Hertlein &
                                   Strong, 1946

Mollusca: Bivalvia: Veneroida:     Cardita affinis (Sowerby,
Cardiidae                          1833)

                                   Lophocardium annettae (Dall,
                                   1889) as Cardium annettae

                                   Trigoniocardia guanacastense
                                   (Hertlein & Strong, 1947) as
                                   Cardium (Americardia)
                                   guanacastense

Mollusca: Bivalvia: Veneroida:     Donax asper Hanley, 1845
Donacidae
                                   Donax assimilis Hanley, 1845

                                   Iphigenia altior (Sowerby, 1833)

Mollusca: Bivalvia: Veneroida:     Cymatioa electilis Berry, 1963
Lasaeidae
                                   Rochefortia compressa Dall, 1913

                                   Solecardia eburnea Conrad, 1849

Mollusca: Bivalvia: Veneroida:     Heterodonax bimaculata
Psammobiidae                       Linnaeus, 1758

Mollusca: Bivalvia: Veneroida:     Tagelus (Tagelus) affinis
Solecurtidae                       (Adams, 1852)

                                   Tagelus (Tagelus)
                                   californianus (Conrad, 1837)

                                   Tagelus (Mesopleura)
                                   peruvianus Pilsbry & Olsson,
                                   1941

                                   Tagelus (Tagelus) violascens
                                   Carpenter, 1857

Mollusca: Bivalvia: Veneroida:     Tellina cumingii Hanley, 1844
Tellinidae                         as Tellina (Tellinela)
                                   cumingii

                                   Tellina laceridens Hanley,
                                   1844 as Tellina (Eurytellina)
                                   laceridens

                                   Tellina simulans Adams, 1852
                                   as Tellina (Eurytellina)
                                   simulans

Mollusca: Bivalvia: Veneroida:     Diplodonta sericata (Reeve,
Ungulinidae                        1850) as Taras sericatus

                                   Diplodonta subquadrata
                                   (Carpenter, 1856) as Taras
                                   subquadrata

Mollusca: Bivalvia: Veneroida:     Chione subimbricata (Sowerby,
Veneridae                          1835) as Anomalocardia
                                   subimbricata

                                   Chione subrugosa (Wood, 1828)
                                   as Anomalocardia subrugosa

                                   Cyclinella subquadrata
                                   (Hanley, 1845)

                                   Dosinia (Dosinidia) dunkeri
                                   (Philippi, 1844)

                                   Megapitaria aurantiaca
                                   (Sowerby, 1831)

                                   Megapitaria squalida (Sowerby,
                                   1835)

                                   Periglypta multicostata
                                   (Sowerby, 1835) as Antigona
                                   (Periglypta) multicostata

                                   Pitar mexicanus Hertlein &
                                   Strong, 1948

                                   Pitar unicolor (Sowerby, 1835)

                                   Protothaca grata (Say, 1830)

                                   Ventricolaria isocardia
                                   (Verrill, 1870) as Antigona
                                   (Ventricola) isocardia

Mollusca: Bivalvia: Ostreoida:     Striostrea prismatica (Gray,
Ostreidae                          1825) as Ostrea iridescens

Mollusca: Bivalvia: Pectinoida:    Nodipecten subnodosus
Pectinidae                         (Sowerby, 1833) as Pecten
                                   (Lyropecten) subnodosus

                                   Argopecten irradians
                                   concentricus (Say, 1822) as
                                   Pecten (Plagioctenium)
                                   circularis

                                   Euvola vogdesi (Arnold, 1906)
                                   as Pecten (Pecten) vogdesi

Mollusca: Bivalvia:                Spondylus princeps Broderip,
Pectinoida: Spondylidae            1833

Mollusca: Bivalvia:                Pinna rugosa (Sowerby, 1835)
Pterioidea: Pinnidae               Pinna sp.

Mollusca: Bivalvia:                Pinctada mazatlanica (Hanley,
Pterioidea: Pteriidae              1855)

Mollusca: Gastropoda:              Bullapunctulata A. Adams in
Cephalaspidea: Bullidae            Sowerby, 1850

Mollusca: Gastropoda:              Acteocina carinata (Carpenter,
Cephalaspidea: Cylichnidae         1857)

Mollusca: Gastropoda:              Cylichnella tabogaensis
Cephalaspidea: Cylichnidae         (Strong & Hertlein, 1939)

Mollusca: Gastropoda:              Atys defuncta (Baker & Hanna,
Cephalaspidea: Haminoeidae         1927)

Mollusca: Gastropoda:              Retusa paziana Dall, 1919
Cephalaspidea: Retusidae

Mollusca: Gastropoda:              Volvulella cylindrica
Cephalaspidea: Rhizoridae          (Carpenter, 1864)

Mollusca: Gastropoda:              Volvulella panamica Dall, 1919
Cephalaspidea: Rhizoridae

Mollusca: Gastropoda:              Caecum richthofeni Strong &
Littorinimorpha: Caecidae          Hertlein, 1939

                                   Elephantulum mirificum (de
                                   Folin, 1867)

Mollusca: Gastropoda:              Echinolittorina fuscolineata
Littorinimorpha: Littorinidae      (Reid, 2002) as Nodilittorina
                                   fuscolineata

Mollusca: Gastropoda:              Strombus gracilior G.B.
Littorinimorpha: Strombidae        Sowerby I, 1825

Mollusca: Gastropoda:              Cyclostremiscus
Littorinimorpha: Tornidae          salvatierrensis Herts, Myers &
                                   Gemmell, 1992

Mollusca: Gastropoda:              Cotonopsis hirundo (Gaskoin,
Neogastropoda: Columbellidae       1852)

                                   Sincola dorsata (Sowerby,
                                   1832)

                                   Sincola gibberula (Sowerby,
                                   1832)

                                   Strombina elegans (Sowerby,
                                   1832)

                                   Strombina maculosa (Sowerby,
                                   1832)

                                   Strombina recurva (Sowerby,
                                   1832)

Mollusca: Gastropoda:              Conus archon Broderip, 1833
Neogastropoda: Conidae
                                   Conus arcuatus Broderip &
                                   Sowerby, 1829

                                   Conus brunneus Wood, 1828

                                   Conus lucidus Wood, 1828

                                   Conus mahogani Reeve, 1843

                                   Conus nux Broderip, 1833

                                   Conus perplexus Sowerby, 1857

                                   Conus recurvus Broderip, 1833

                                   Conus regularis Sowerby, 1833

                                   Conus tiaratus Sowerby, 1833

                                   Strombinoturris crockeri
                                   Hertlein & Strong, 1951

Mollusca: Gastropoda:              Pustulatirus hemphilli
Neogastropoda: Fasciolariidae      (Hertlein & Strong, 1951) as
                                   Latirus hemphilli

Mollusca: Gastropoda:              Mitra (Strigatella) tristis
Neogastropoda: Mitridae            Broderip, 1836

Mollusca: Gastropoda:              Muricanthus ambiguus (Reeve,
Neogastropoda: Muricidae           1845) as Muricanthus
                                   callidinus

Mollusca: Gastropoda:              Thais brevidentata (Wood,
Neogastropoda: Muricidae           1828) as Acanthina
                                   brevidentata

Mollusca: Gastropoda:              Terebra sp.
Neogastropoda: Terebridae

Mollusca: Gastropoda:              Crassispira coracina McLean &
Neogastropoda: Turridae            Poorman, 1971

                                   Crassispira ballenaensis
                                   Hertlein & Strong, 1951 as
                                   Crassispira turricula
                                   ballanaensis

Mollusca: Gastropoda               Alaba supralirata Carpenter,
Neotaenioglossa: Litiopidae        1856

Mollusca: Gastropoda:              Crosslandia daedali Poorman &
Nudibranchia: Scyllaeidae          Mulliner, 1981

Arthropoda: Crustacea:             Ampelisca pugetica Stimpson,
Malacostraca: Amphipoda:           1864
Ampeliscidae

Arthropoda: Crustacea:             Foxiphalus golfensis Barnard &
Malacostraca: Amphipoda:           Barnard, 1982
Phoxocephalidae

Arthropoda: Crustacea:             Paguristes digueti Bouvier,
Malacostraca: Decapoda:            1893
Anomura: Diogenidae
                                   Trizopagurus magnificus
                                   (Bouvier, 1898)

Arthropoda: Crustacea:             Phimochirus californiensis
Malacostraca: Decapoda:            (Benedict, 1892)
Anomura: Paguridae

Arthropoda: Crustacea:             Pachycheles biocellatus
Malacostraca: Decapoda:            (Lockington, 1878)
Anomura: Porcellanidae
                                   Pachycheles spinidactylus
                                   Haig, 1957

                                   Pachycheles vicarius Nobili,
                                   1901

Arthropoda: Crustacea:             Petrolisthes armatus (Gibbes,
Malacostraca: Decapoda:            1850)
Brachyura: Aethridae
                                   Petrolisthes edwardsii (de
                                   Saussure, 1853)

                                   Petrolisthes glasselli Haig,
                                   1957

Arthropoda: Crustacea:             Petrolisthes haigae (Chace,
Malacostraca: Decapoda:            1962)
Brachyura: Calappidae
                                   Petrolisthes hians Nobili,
                                   1901

                                   Petrolisthes ortmanni Nobili,
                                   1901

                                   Petrolisthes tonsorius Haig,
                                   1960

                                   Petrolisthes tridentatus
                                   Stimpson, 1859

                                   Pisidia magdalenensis
                                   (Glassell, 1936)

                                   Porcellana cancrisocialis
                                   Glassell, 1936

                                   Porcellana paguriconviva
                                   Glassell, 1936

                                   Hepatus kossmanni Neumann,
                                   1878

                                   Cryptosoma bairdii (Stimpson,
                                   1870) as Cycloes bairdii

                                   Calappa convexa Saussure, 1853

Arthropoda: Crustacea:             Daira americana Stimpson, 1860
Malacostraca: Decapoda:
Brachyura: Dairidae

Arthropoda: Crustacea:             Hypoconcha panamensis Smith,
Malacostraca: Decapoda:            in Verrill, 1869
Brachyura: Dromiidae

Arthropoda: Crustacea:             Herbstia pubescens Stimpson,
Malacostraca: Decapoda:            1871
Brachyura: Epialtidae
                                   Herbstia tumida (Stimpson,
                                   1871)

                                   Microlissa aurivillium
                                   (Rathbun, 1898) as Lissa
                                   aurivillium

                                   Pitho picteti (Saussure, 1856)

Arthropoda: Crustacea:             Ethusa lata Rathbun, 1894
Malacostraca: Decapoda:
Brachyura: Ethusidae               Ethusa panamensis Finnegan,
                                   1931 as Ethusa mascarone
                                   panamensis

Arthropoda: Crustacea:             Gecarcinus lateralis
Malacostraca: Decapoda:            (Freminville, 1835) as
Brachyura: Gecarcinidae            Gecarcinus (Gecarcinus)
                                   lateralis quadratus

                                   Ucides occidentalis (Ortmann,
                                   1897)

Arthropoda: Crustacea:             Geograpsus lividus (H. Milne
Malacostraca: Decapoda:            Edwards, 1837)
Brachyura: Grapsidae
                                   Goniopsis pulchra (Lockington,
                                   1877)

                                   Grapsus grapsus Linnaeus,
                                   1758)

                                   Pachygrapsus transversus
                                   (Gibbes, 1850)

Arthropoda: Crustacea:             Eucinetops panamensis Rathbun,
Malacostraca: Decapoda:            1923
Brachyura: Inachidae
                                   Ericerodes angulatus
                                   (Finnegan, 1931)

                                   Stenorhynchus debilis (Smith,
                                   1871)

Arthropoda: Crustacea:             Collodes tenuirostris Rathbun,
Malacostraca: Decapoda:            1894
Brachyura: Inachoididae

                                   Euprognatha bifida Rathbun,
                                   1894

                                   Inachoides laevis Stimpson,
                                   1870

                                   Paradasygyius depressus (Bell,
                                   1835)

Arthropoda: Crustacea:             Ebalia magdalenensis Rathbun,
Malacostraca: Decapoda:            1933
Brachyura: Leucosiidae
                                   Iliacantha hancocki Rathbun,
                                   1935

                                   Lithadia cumingii Bell, 1855

                                   Randallia agaricias Rathbun,
                                   1898

                                   Randallia bulligera Rathbun,
                                   1898

                                   Randallia minuta Rathbun, 1935

Arthropoda: Crustacea:             Ala cornuta (Stimpson, 1860)
Malacostraca: Decapoda:
Brachyura: Majidae                 Hemus finneganae Garth, 1958

                                   Microphrys branchialis
                                   Rathbun, 1898

                                   Mithraculus denticulatus
                                   (Bell, 1835) as Mitrax
                                   (Mitrax) denticulatus

                                   Mithrax pygmaeus Bell, 1835

                                   Stenocionops ovatus (Bell,
                                   1835)

                                   Teleophrys cristulipes
                                   Stimpson, 1860

                                   Thoe panamensis Nobili, 1901
                                   as Thoe sulcata panamensis

Arthropoda: Crustacea:             Ocypode gaudichaudii H. Milne
Malacostraca: Decapoda:            Edwards & Lucas, 1843
Brachyura: Ocypodidae
                                   Ocypode occidentalis Stimpson,
                                   1860

                                   Uca latimanus (Rathbun, 1894)

                                   Uca panamensis (Stimpson, 1859)

Arthropoda: Crustacea:             Ozius tenuidactylus
Malacostraca: Decapoda:            (Lockington, 1877)
Brachyura: Oziidae

Arthropoda: Crustacea:             Celatopesia hassleri (Rathbun,
Malacostraca: Decapoda:            1925) as Cryptopodia hassleri
Brachyura: Parthenopidae
                                   Daldorfia glasselli (Garth,
                                   1958) as Thyrolambrus
                                   glasselli

                                   Daldorfia trigona (A.
                                   Milne-Edwards, 1869) as
                                   Daldorfia garthi


                                   Heterocrypta macrobrachia
                                   Stimpson, 1871

                                   Ochtholambrus stimpsoni
                                   (Garth, 1958) as Pathenope
                                   (Pseudolambrus) stimpsoni

                                   Solenolambrus arcuatus
                                   Stimpson, 1891

Arthropoda: Crustacea:             Eurypanopeus transversus
Malacostraca: Decapoda:            (Stimpson, 1860)
Brachyura: Panopeidae
                                   Eurytium tristani Rathbun,
                                   1906

                                   Hexapanopeus costaricensis
                                   Garth, 1940

                                   Malacoplax californiensis
                                   (Lockington, 1877) as
                                   Speocarcinus californiensis

                                   Panopeus chilensis H.
                                   Milne-Edwards & Lucas, 1843

                                   Panopeus purpureus Lockington,
                                   1877

Arthropoda: Crustacea:             Pilumnus pygmaeus Boone, 1927
Malacostraca: Decapoda:
Brachyura: Pilumnidae              Pilumnus xantusii Stimpson,
                                   1860

Arthropoda: Crustacea:             Callinectes arcuatus Ordway,
Malacostraca: Decapoda:            1863
Brachyura: Portunidae
                                   Cronius ruber (Lamarck, 1818)

                                   Euphylax robustus Stimpson,
                                   1860

                                   Portunus acuminatus (Stimpson,
                                   1871)

Arthropoda: Crustacea:             Raninoides benedicti Rathbun,
Malacostraca: Decapoda:            1935
Brachyura: Raninidae

Arthropoda: Crustacea:             Trapezia bidentata (Forskal,
Malacostraca: Decapoda:            1775) as Trapezia ferruginea
Brachyura: Trapeziidae
                                   Trapezia corallina
                                   Gerstaecker, 1857

                                   Trapezia formosa Stimpson,
                                   1869

Arthropoda: Crustacea:             Edwardsium lobipes (Rathbun,
Malacostraca: Decapoda:            1898) as Medaeus lobipes
Brachyura: Xanthidae
                                   Heteractaea lunata (Lucas in
                                   H. Milne Edwards & Lucas,
                                   1844)

                                   Liomera cinctimana (White,
                                   1847)

                                   Microcassiope xantusii
                                   (Stimpson, 1871)

                                   Platyactaea dovii (Stimpson,
                                   1871)

Arthropoda: Crustacea:             Alpheus cylindricus Kingsley,
Malacostraca: Decapoda:            1878
Caridea: Alpheidae

Arthropoda: Crustacea:             Macrobrachium tenellum (Smith,
Malacostraca: Decapoda:            1871)
Caridea: Palaemonidae
                                   Neopontonides henryvonprahli
                                   Ramos, 1995

                                   Periclimenes spinosus
                                   Holthuis, 1951

                                   Pontonia margarita Smith, 1869

                                   Pontonia simplex Holthuis,
                                   1951

Arthropoda: Crustacea:             Plesionika mexicana Chace,
Malacostraca: Decapoda:            1937
Caridea: Pandalidae

Arthropoda: Crustacea:             Excirolana braziliensis
Malacostraca: Isopoda:             Richardson, 1912
Cirolanidae

Arthropoda: Crustacea:             Neogonodactylus bahiahondensis
Malacostraca: Stomatopoda:         (Schmitt, 1940) in lit
Gonodactylidae                     Gonodactylus bahiahondensis

                                   Neogonodactylus
                                   lalibertadensis (Schmitt,
                                   1940) as Gonodactylus
                                   lalibertadensis

Arthropoda: Crustacea:             Meiosquilla dawsoni Manning,
Malacostraca: Stomatopoda:         1970
Squillidae
                                   Squilla parva Bigelow, 1891

Arthropoda: Crustacea:             Acartia lilljeborgii
Maxillopoda: Calanoida:            Giesbrecht, 1889
Acartiidae
                                   Acartia tonsa Dana, 1849

Arthropoda: Crustacea:             Canthocalanus pauper
Maxillopoda: Calanoida:            (Giesbrecht, 1888)
Calanidae

Arthropoda: Crustacea:             Caligus chorinemi Kroyer, 1863
Maxillopoda: Calanoida:
Caligidae                          Neocalanus gracilis (Dana,
                                   1852)

                                   Undinula vulgaris (Dana, 1849)

Arthropoda: Crustacea:             Calocalanus spp.
Maxillopoda: Calanoida:
Calocalanidae

Arthropoda: Crustacea:             Candacia catula (Giesbrecht,
Maxillopoda: Calanoida:            1888)
Candaciidae

                                   Candacia pachydactyla (Dana,
                                   1849)

Arthropoda: Crustacea:             Centropages furcatus Dana,
Maxillopoda: Calanoida:            1849
Centropagidae

                                   Centropages longicornis Mori,
                                   1932

Arthropoda: Crustacea:             Clausocalanus arcuicornis
Maxillopoda: Calanoida:            (Dana, 1849)
Clausocalanidae

Arthropoda: Crustacea:             Eucalanus elongatus (Dana,
Maxillopoda: Calanoida:            1848)
Eucalanidae
                                   Pareucalanus attenuatus (Dana,
                                   1849) as Eucalanus attenuates

Arthropoda: Crustacea:             Euchaeta indica Wolfenden,
Maxillopoda: Calanoida:            1905 as Euchaeta wolfendeni
Euchaetidae
                                   Paraeuchaeta longicornis
                                   (Giesbrecht, 1888) as Euchaeta
                                   longicornis

Arthropoda: Crustacea:             Lucicutia flavicornis (Claus,
Maxillopoda: Calanoida:            1863)
Lucicutiidae

Arthropoda: Crustacea:             Acrocalanus longicornis
Maxillopoda: Calanoida:            Giesbrecht, 1888
Paracalanidae
                                   Paracalanus parvus (Claus,
                                   1863)

Arthropoda: Crustacea:             Labidocera acuta (Dana, 1849)
Maxillopoda: Calanoida:
Pontellidae                        Pontellina plumata (Dana,
                                   1849)

Arthropoda: Crustacea:             Pseudodiaptomus wrighti
Maxillopoda: Calanoida:            Johnson, 1964
Pseudodiaptomidae

Arthropoda: Crustacea:             Scolecithricella bradyi
Maxillopoda: Calanoida:            (Giesbrecht, 1888)
Scolecitrichidae

Arthropoda: Crustacea:             Temora discaudata Giesbrecht,
Maxillopoda: Calanoida:            1889
Temoridae

Arthropoda: Crustacea:             Rhincalanus spp.
Maxillopoda: Calanoida:
Rhincalanidae

Arthropoda: Crustacea:             Subeucalanus subcrassus
Maxillopoda: Calanoida:            (Giesbrecht, 1888)
Subeucalanidae
                                   Subeucalanus subtenuis
                                   (Giesbrecht, 1888) as
                                   Eucalanus subtenuis

Arthropoda: Crustacea:             Oithona plumifera Baird, 1843
Maxillopoda: Cyclopoida:
Oithonidae
                                   Oithona setigera Dana, 1852

                                   Oithona similis Claus, 1866

Arthropoda: Crustacea:             Cymbasoma concepcionae Suarez-
Maxillopoda: Mostrilloida:         Morales & Morales-Ramirez,
Monstrillidae                      2003

Arthropoda: Crustacea:             Saphirella tropica Wolfenden,
Maxillopoda:                       1906
Poecilostomatoida:
Clausidiidae

Arthropoda: Crustacea:             Corycaeus ovalis Claus, 1863
Maxillopoda:
Poecilostomatoida: Corycaeidae     Ditrichocorycaeus brehmi
                                   (Steuer, 1910) as Corycaeus
                                   bremhi

Arthropoda: Crustacea:             Oncaea mediterranea (Claus,
Maxillopoda:                       1863)
Poecilostomatoida: Oncaeidae
                                   Oncaea venusta Philippi, 1843

Arthropoda: Crustacea:             Copilia mirabilis Dana, 1852
Maxillopoda:
Poecilostomatoida:                 Sapphirina opalina Dana, 1849
Sapphirinidae
                                   Sapphirina ovatolanceolata
                                   Dana, 1849

                                   Sapphirina scarlata
                                   Giesbrecht, 1891

Arthropoda: Crustacea:             Balanus masignotus Henry &
Maxillopoda: Sessilia:             McLaughlin, 1967
Balanidae
                                   Balanus mexicanus Henry, 1941
                                   as Balanus concavus mexicanus

                                   Balanus trigonus Darwin, 1864

Arthropoda: Crustacea:             Lernanthropus micropterygis
Maxillopoda:                       Richiardi, 1884
Siphonostomatoida:
Lernanthropidae

Arthropoda: Crustacea:             Unidentified species
Branchiopoda: Diplostraca:
Cladocera

Arthropoda: Crustacea:             Unidentified species
Ostracoda

Arthropoda: Chelicerata:           Anoplodactylus especies
Pycnogonida: Pantopoda:            indeterminada
Phoxichilidiidae

Bryozoa: Gymnolaemata:             Beania hirtissima (Heller,
Cheilostomatida: Beaniidae         1867)

Bryozoa: Gymnolaemata:             Aimulosia uvulifera (Osburn,
Cheilostomatida:                   1914)
Buffonellodidae

Bryozoa: Gymnolaemata:             Porella porifera (Hincks,
Cheilostomatida:                   1884)
Bryocryptellidae

Bryozoa: Gymnolaemata:             Mamillopora cupula Smitt, 1873
Cheilostomatida:
Mamilloporidae

Bryozoa: Gymnolaemata:             Bifustra savartii (Audouin,
Cheilostomatida:                   1826) as Membranipora savarti
Membraniporidae

Bryozoa: Gymnolaemata:             Microporella gibbosula Canu &
Cheilostomatida:                   Bassler, 1930
Microporellidae

Bryozoa: Gymnolaemata:             Petralia japonica (Busk, 1884)
Cheilostomatida: Petraliidae

Bryozoa: Gymnolaemata:             Hippodiplosia americana
Cheilostomatida:                   (Verrill, 1875)
Schizoporellidae

Bryozoa: Gymnolaemata:             Amathia convoluta (Lamarck,
Ctenostomatida: Vesiculariidae     1816)

Echinodermata: Ophiuroidea:        Ophiolepis plateia
Ophiurida: Ophiolepididae          Ziesenhenne, 1940

Echinodermata: Asteroidea:         Astropecten armatus Gray, 1840
Paxillosida: Astropectinidae

Echinodermata: Echinoidea:         Eucidaris thourasii
Cidaroida: Cidaridae               (Valenciennes, 1846)

Echinodermata: Echinoidea:         (?) Encope irregularis Clark,
Clypeasteroida: Mellitidae         1948

Echinodermata: Echinoidea:         Astropyga pulvinata (Lamarck,
Diadematoida: Diadematidae         1816)

                                   Diadema mexicaum A. Agassiz,
                                   1863

Echinodermata: Echinoidea:         Toxopneustes roseus (A.
Temnopleuroida: Toxopneustidae     Agassiz, 1863)

Echinodermata: Holothuroidea:      Holothuria (Platyperona)
Aspidochirota: Holothuriidae       difficilis Semper, 1868 as
                                   Microthele difficilis

                                   Holothuria (Selenkothuria)
                                   Deichmann, 1958 as
                                   Selenkothuria lubrica

                                   Holothuria (Thymiosycia)
                                   impatiens (Forskal, 1775) as
                                   Brandtothuria impatiens

                                   Labidodemas maccullochi
                                   (Deichmann, 1958) as
                                   Irenothuria macculloch

Chaetognatha: Sagittoidea:         Unidentified species
Sagittoidea: Sagittidae

Chordata: Appendicularia:          Oikopleura fusiformis Fol,
Copelata: Oikopleuridae            1872

                                   Oikopleura longicauda (Vogt,
                                   1854)

                                   Oikopleura rufescens Fol, 1872

Chordata: Appendicularia:          Fritillaria haplostoma Fol,
Copelata: Fritillariidae           1872

Chordata: Thaliacea: Salpida:      Unidentified species
Salpidae

Chordata: Chondrichthyes:          Ginglymostoma cirratum
Orectolobiformes:                  (Bonnaterre, 1788)
Ginglymostomatidae

Chordata: Chondrichthyes:          Himantura pacifica (Beebe &
Rajiformes: Dasyatidae             Tee-Van, 1941)

Chordata: Chondrichthyes:          Aetobatus narinari (Euphrasen,
Rajiformes: Myliobatidae           1790)

Chordata: Actinopterygii:          Albula pacifica (Beebe, 1942)
Albuliformes: Albulidae            as Dixonina pacifica

Chordata: Actinopterygii:          Gymnomuraena zebra (Shaw, 1797)
Anguilliformes: Muraenidae
Beryciformes: Holocentridae        Gymnothorax castaneus (Jordan
                                   & Gilbert, 1883)

                                   Muraena lentiginosa Jenyns,
                                   1842

                                   Myripristis sp.

                                   Sargocentron suborbitalis
                                   (Gill, 1863)

Chordata: Actinopterygii:          Fistularia commersonii
Syngnathiformes: Fistulariidae     Ruppell, 1838

Chordata: Actinopterygii:          Hippocampus ingens Girard, 1858
Syngnathiformes: Syngnathidae

Chordata: Actinopterygii:          Prionurus laticlavius
Perciformes: Acanthuridae          (Valenciennes, 1846)

Chordata: Actinopterygii:          Apogon dovii Gunther, 1862
Perciformes: Apogonidae
                                   Apogon pacificus (Herre, 1935)
                                   as Apogon pacifici

Chordata: Actinopterygii:          Ophioblennius steindachneri
Perciformes: Blenniidae            Jordan & Evermann, 1898

                                   Plagiotremus azaleus (Jordan &
                                   Bollman, 1890)

Chordata: Actinopterygii:          Caranx caballus Gunther, 1868
Perciformes: Carangidae
                                   Caranx sexfasciatus Quoy &
                                   Gaimard, 1825 as Caranx
                                   marginatus

                                   Seriola lalandi Valenciennes,
                                   1833 as Seriola dorsalis

                                   Seriola rivoliana
                                   Valenciennes, 1833

Chordata: Actinopterygii:          Chaetodon humeralis Gunther,
Perciformes: Chaetodontidae        1860

                                   Johnrandallia nigrirostris
                                   (Gill, 1862)

Chordata: Actinopterygii:          Cirrhitichthys oxycephalus
Perciformes: Cirrhitidae           (Bleeker, 1855)

                                   Cirrhitus rivulatus
                                   Valenciennes, 1846

Chordata: Actinopterygii:          Chaetodipterus zonatus
Perciformes: Ephippidae            (Girard, 1858)

Chordata: Actinopterygii:          Anisotremus taeniatus Gill,
Perciformes: Haemulidae            1861

                                   Haemulon flaviguttatum Gill,
                                   1862

                                   Haemulon maculicauda (Gill,
                                   1862)

                                   Haemulon scudderii Gill, 1862

                                   Haemulon sexfasciatum Gill,
                                   1862

                                   Haemulon steindachneri (Jordan
                                   & Gilbert, 1882)

Chordata: Actinopterygii:          Kyphosus analogus (Gill, 1862)
Perciformes: Kyphosidae
                                   Kyphosus elegans (Peters,
                                   1869)

                                   Sectator ocyurus (Jordan &
                                   Gilbert, 1882)

Chordata: Actinopterygii:          Bodianus diplotaenia (Gill,
Perciformes: Labridae              1862)

                                   Halichoeres chierchiae Di
                                   Caporiacco, 1948

                                   Halichoeres dispilus (Gunther,
                                   1864)

                                   Halichoeres melanotis
                                   (Gilbert, 1890)

                                   Halichoeres nicholsi (Jordan &
                                   Gilbert, 1882)

                                   Halichoeres notospilus
                                   (Gunther, 1864)

                                   Novaculichthys taeniourus
                                   (Lacepede, 1801) Thalassoma
                                   lucasanum (Gill, 1862)

                                   Xyrichtys victori Wellington,
                                   1992

Chordata: Actinopterygii:          Lutjanus argentiventris
Perciformes: Lutjanidae            (Peters, 1869) Lutjanus
                                   guttatus (Steindachner, 1869)

Chordata: Actinopterygii:          Malacanthus brevirostris
Perciformes: Malacanthidae         Guichenot, 1848

Chordata: Actinopterygii:          Holacanthus passer
Perciformes: Pomacanthidae         Valenciennes, 1846

                                   Pomacanthus
                                   zonipectus (Gill, 1862)

Chordata: Actinopterygii:          Abudefduf troschelii (Gill,
Perciformes: Pomacentridae         1862)

                                   Chromis atrilobata Gill, 1862

                                   Microspathodon bairdii (Gill,
                                   1862)

                                   Microspathodon dorsalis (Gill,
                                   1862)

                                   Stegastes acapulcoensis
                                   (Fowler, 1944)

                                   Stegastes flavilatus (Gill,
                                   1862)

Chordata: Actinopterygii:          Nicholsina denticulata
Perciformes: Scaridae              (Evermann & Radcliffe, 1917)

                                   Scarus compressus (Osburn &
                                   Nichols, 1916)

                                   Scarus ghobban Forsskal, 1775

                                   Scarus perrico Jordan &
                                   Gilbert, 1882

                                   Scarus rubroviolaceus Bleeker,
                                   1847

Chordata: Actinopterygii:          Odontoscion xanthops Gilbert,
Perciformes: Sciaenidae            1898

                                   Pareques viola (Gilbert, 1898)

Chordata: Actinopterygii:          Scomberomorus sierra Jordan &
Perciformes: Scombridae            Starks, 1895

Chordata: Actinopterygii:          Alphestes immaculatus Breder,
Perciformes: Serranidae            1936

                                   Alphestes multiguttatus
                                   (Gunther, 1867)

                                   Cephalopholis panamensis
                                   (Steindachner, 1877)

                                   Dermatolepis dermatolepis
                                   (Boulenger, 1895)

                                   Epinephelus labriformis
                                   (Jenyns, 1840)

                                   Hyporthodus acanthistius
                                   (Gilbert, 1892) as Epinephelus
                                   acanthistius

                                   Paranthias colonus
                                   (Valenciennes, 1846)

                                   Rypticus bicolor Valenciennes,
                                   1846

                                   Serranus psittacinus
                                   Valenciennes, 1846

Chordata: Actinopterygii:          Cosmocampus arctus coccineus
Syngnathiformes: Syngnathidae      (Herald, 1940) as Bryx
                                   (Simocampus) coccineus

Chordata: Actinopterygii:          Balistes polylepis
Tetraodontiformes: Balistidae      Steindachner, 1876

                                   Melichthys niger (Bloch, 1786)

                                   Pseudobalistes naufragium
                                   (Jordan & Starks, 1895)

                                   Sufflamen verres (Gilbert &
                                   Starks, 1904)

Chordata: Actinopterygii:          Diodon holocanthus Linnaeus,
Tetraodontiformes: Diodontidae     1758

                                   Diodon hystrix Linnaeus, 1758

Chordata: Actinopterygii:          Aluterus scriptus (Osbeck,
Tetraodontiformes:                 1765)
Monocanthidae

Chordata: Actinopterygii:          Ostracion meleagris Shaw, 1796
Tetraodontiformes: Ostraciidae

Chordata: Actinopterygii:          Arothron hispidus (Linnaeus,
Tetraodontiformes:                 1758)
Tetraodontidae
                                   Arothron meleagris (Anonymous,
                                   1798)

                                   Canthigaster punctatissima
                                   (Gunther, 1870)

Chordata: Reptilia: Squamata:      Pelamis platurus (Linnaeus,
Elapidae                           1766)

Chordata: Mammalia: Cetacea:       Megaptera novaeangliae
Balaenopteridae                    Borowski, 1781

Chordata: Mammalia: Cetacea:       Stenella attenuata Gray, 1846
Delphinidae

                                   Stenella attenuata graffmani
                                   Lonnberg, 1934

                                   Steno bredanensis Lesson, 1828

Clasification: Phylum              References
(Subphylum): Class: Order
(Infraorder): Family

Cyanobacteria: Cyanophyceae:       Drouet 1936
Chroococcales: Microcystaceae

Cyanobacteria: Cyanophyceae:       Taylor 1945
Nostocales: Rivulariaceae

Cyanobacteria: Cyanophyceae:       Drouet 1936
Oscillatoriales:
Oscillatoriaceae

Cyanobacteria: Cyanophyceae:       Taylor 1945
Oscillatoriales: Phormidiaceae

                                   Vargas-Montero 2004

Bacillariophyta:                   Vargas-Montero 2004
Bacillariophyceae:
Bacillariales: Bacillariaceae

                                   Vargas-Montero 2004

Bacillariophyta:                   Vargas-Montero 2004
Coscinodiscophyceae:
Rhizosoleniales:
Rhizosoleniaceae

Dinoflagellata: Dinophyceae:       Vargas-Montero 2004
Dinophysiales: Dinophyceae

Dinoflagellata: Dinophyceae:       Vargas-Montero 2004
Gonyaulacales: Ceratiaceae

                                   Vargas-Montero 2004

Dinoflagellata: Dinophyceae:       Vargas-Montero 2004
Gonyaulacales: Goniodomataceae
                                   Vargas-Montero 2004

Dinoflagellata: Dinophyceae:       Vargas-Montero 2004
Gonyaulacales: Gonyaulacaceae

                                   Morales-Ramirez et al. 2001

Dinoflagellata: Dinophyceae:       Vargas-Montero 2004

Gymnodiniales: Gymnodiniaceae

                                   Vargas-Montero 2004

Dinoflagellata: Dinophyceae:       Vargas-Montero 2004
Noctilucales: Noctilucaceae

Dinoflagellata: Dinophyceae:       Vargas-Montero 2004
Peridiniales: Peridinaceae

Dinoflagellata: Dinophyceae:       Vargas-Montero 2004
Protocentrales: Protocentraceae

                                   Vargas-Montero 2004

                                   Vargas-Montero 2004

Chlorophyta: Bryopsidophyceae:     Fernandez & Cortes 2005
Bryopsidales: Caulerpaceae

Chlorophyta: Bryopsidophyceae:     Fernandez & Alvarado 2008
Bryopsidales: Halimedaceae

Chlorophyta: Ulvophyceae:          Taylor 1945, Dawson 1961
Cladophorales: Cladophoraceae

                                   Dawson 1960, 1961

Chlorophyta: Ulvophyceae:          Fernandez & Alvarado 2008
Dasycladales: Polyphysaceae

Chlorophyta: Ulvophyceae:          Dawson 1960
Siphonocladales: Boodleacea

                                   Tsuda 1968

Chlorophyta: Ulvophyceae:          Tsuda 1968
Ulvales: Ulvaceae
                                   Taylor 1945, Fernandez &
                                   Alvarado 2008

                                   Fernandez & Alvarado 2008

Rhodophyta: Florideophyceae:       Dawson 1960, 1961
Acrochaetiales:
Acrochaetiaceae

Rhodophyta: Florideophyceae:       Dawson 1960, 1961
Ceramiales: Ceramiaceae
                                   Dawson 1960, 1961

Rhodophyta: Florideophyceae:       Taylor 1945, Dawson 1961
Ceramiales: Delesseriaceae

Rhodophyta: Florideophyceae:       Cortes 2001
Ceramiales: Rhodomelaceae

                                   Taylor 1945, Dawson 1961

                                   Taylor 1945, Dawson 1957

Rhodophyta: Florideophyceae:       Dawson 1961
Ceramiales: Spyridiaceae

                                   Dawson & Beaudette 1959

Rhodophyta: Florideophyceae:       Dawson & Beaudette 1959
Corallinales: Corallinaceae
                                   Dawson 1957

                                   Tsuda 1968

Rhodophyta: Florideophyceae:       Taylor 1945, Dawson 1957
Corallinales: Hapalidiaceae

Rhodophyta: Florideophyceae:       Taylor 1945, Dawson 1957
Gigartinales: Areschougiaceae

                                   Dawson 1961

Rhodophyta: Florideophyceae:       Dawson 1961
Gracilariales: Gracilariaceae

                                   Dawson 1957

                                   Dawson 1957, 1961

                                   Taylor 1945, Dawson 1961

Rhodophyta: Florideophyceae:       Dawson 1960, 1961
Halymeniales: Halymeniaceae

                                   Dawson 1960, 1961

Rhodophyta: Florideophyceae:       Taylor 1945
Nemaliales: Galaxauraceae

Rhodophyta: Florideophyceae:       Dawson & Beaudette 1959
Nemaliales: Liagoraceae

Rhodophyta: Florideophyceae:       Taylor 1945, Dawson 1961
Nemaliales: Scinaiaceae

                                   Dawson 1960

                                   Taylor 1945, Dawson 1957

Rhodophyta: Florideophyceae:       Taylor 1945, Dawson 1961
Rhodymeniales: Lomentariaceae

                                   Dawson 1960, 1961

Ochrophyta: Phaeophyceae:          Taylor 1945, Dawson 1961
Dictyotales: Dictyotaceae

                                   Taylor 1945, Dawson 1957

                                   Taylor 1945

                                   Dawson & Beaudette 1959

Ochrophyta: Phaeophyceae:          Tsuda 1968
Ectocarpales: Actinetosporaceae

Ochrophyta: Phaeophyceae:          Tsuda 1968
Scytothamnales: Scytothamnaceae

Magnoliophyta: Liliopsida:         Cortes 2001
Najadales: Ruppiaceae

Magnoliophyta: Liliopsida:         Cortes 2001
Alismatales: Hydrocharitaceae

Magnoliophyta: Magnoliopsida:      Cabrera-Pena et al. 1994
Lamiales: Verbenaceae:

Magnoliophyta: Magnoliopsida:      Cabrera-Pena et al. 1994
Myrtales: Combretaceae

Magnoliophyta: Magnoliopsida:      Cabrera-Pena et al. 1994
Rhizophorales: Rhizophoraceae

Protozoa: Sarcodina:               Bandy & Arnal 1957
Foraminiferida: Bolivinitidae

                                   Cushman & McCulloch 1942

Protozoa: Sarcodina:               Cushman & McCulloch 1948
Foraminiferida: Buliminidae

                                   Bandy & Arnal 1957

Protozoa: Sarcodina:               Bandy & Arnal 1957
Foraminiferida: Cibicididae

Protozoa: Sarcodina:               Cushman & McCulloch 1940
Foraminiferida: Elphidiidae

Protozoa: Sarcodina:               Crouch & Poag 1987
Foraminiferida: Eponididae

Protozoa: Sarcodina:               Bandy & Arnal 1957
Foraminiferida: Fusenkoinidae

Protozoa: Sarcodina:               Bandy & Arnal 1957
Foraminiferida: Globigerinidae

                                   Bandy & Arnal 1957

                                   Bandy & Arnal 1957

                                   Bandy & Arnal 1957

Protozoa: Sarcodina:               Bandy & Arnal 1957
Foraminiferida:
Globorotaliidae

Protozoa: Sarcodina:               Bandy & Arnal 1957
Foraminiferida:
Haplophragmoididae

Protozoa: Sarcodina:               Cushman & McCulloch 1942
Foraminiferida: Loxostomatidae

Protozoa: Sarcodina:               Bandy & Arnal 1957
Foraminiferida: Miliolidae

                                   Bandy & Arnal 1957

                                   Crouch & Poag 1987

                                   Crouch & Poag 1987

                                   Crouch & Poag 1987

Protozoa: Sarcodina:               Bandy & Arnal 1957
Foraminiferida: Nonionidae

Protozoa: Sarcodina:               Cushman & McCulloch 1939
Foraminiferida: Nouriidae

Protozoa: Sarcodina:               Bandy & Arnal 1957
Foraminiferida:
Planorbulinidae

Protozoa: Sarcodina:               Cushman & McCulloch 1948
Foraminiferida: Reussellidae

                                   Cushman & McCulloch 1948

Protozoa: Sarcodina:               Bandy & Arnal 1957
Foraminiferida:
Sphaeroidinidae

Protozoa: Sarcodina:               Bandy & Arnal 1957
Foraminiferida:
Spiroloculinidae

Protozoa: Sarcodina:               Crouch & Poag 1987
Foraminiferida: Textulariidae
                                   Lalicker & McCulloch 1940

                                   Lalicker & McCulloch 1940,
                                   Bandy & Arnal 1957

                                   Lalicker & McCulloch 1940

                                   Bandy & Arnal 1957

Protozoa: Sarcodina:               Cushman & McCulloch 1939
Foraminiferida: Trochamminidae

Cnidaria: Anthozoa:                Jimenez 1997
Alcyonacea: Clavulariidae

Cnidaria: Anthozoa:                Breedy & Guzman 2007,
Alcyonacea: Gorgoniidae            Breedy 2009

                                   Breedy & Guzman 2005

                                   Breedy & Guzman 2007

                                   Breedy & Guzman 2003

Cnidaria: Anthozoa:                Breedy & Guzman 2011
Alcyonacea: Plexauridae

Cnidaria: Anthozoa:                Cortes & Murillo 1985
Scleractinia: Agariciidae

                                   Cortes & Murillo 1985

                                   Cortes & Murillo 1985

                                   Cortes & Guzman 1998

                                   Cortes & Guzman 1998

Cnidaria: Anthozoa:                Cortes & Guzman 1998
Scleractinia: Fungiidae

                                   Cortes & Guzman 1998

Cnidaria: Anthozoa:                Cortes & Guzman 1998
Scleractinia: Pocilloporidae

                                   Glynn et al. 1983

                                   Jimenez 1997

                                   Cortes & Murillo 1985

                                   Cortes & Murillo 1985

Cnidaria: Anthozoa:                Cortes & Murillo 1985
Scleractinia: Poritidae
                                   Cortes & Guzman 1998

Cnidaria: Anthozoa:                Cortes & Murillo 1985
Scleractinia: Rhizangiidae

Cnidaria: Anthozoa:                Glynn et al. 1983
Scleractinia: Siderastreidae

                                   Jimenez 1997

Cnidaria: Scyphozoa:               Rodriguez-Saenz & Segura-
Coronateae: Nausithoidae           Puertas 2009

Cnidaria: Scyphozoa:               Rodriguez-Saenz & Segura-
Semaeostomae: Pelagiidae           Puertas 2009

Cnidaria: Hydrozoa:                Fraser 1938
Anthoathecata:
Bougainvilliidae

                                   Rodriguez-Saenz & Segura-
                                   Puertas 2009

                                   Rodriguez-Saenz & Segura-
                                   Puertas 2009

                                   Fraser 1938

                                   Rodriguez-Saenz & Segura-
                                   Puertas 2009

                                   Rodriguez-Saenz & Segura-
                                   Puertas 2009

                                   Fraser 1938

Cnidaria: Hydrozoa:                Rodriguez-Saenz & Segura-
Anthoathecata: Bythotiaridae       Puertas 2009

                                   Rodriguez-Saenz & Segura-
                                   Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz & Segura-
Anthoathecata: Corymorphidae       Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz & Segura-
Anthoathecata: Corynidae           Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Anthoathecata: Euphysidae          Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

Cnidaria: Hydrozoa:                Fraser 1938
Anthoathecata: Hydractiniidae

                                   Fraser 1938

Cnidaria: Hydrozoa:                Fraser 1938
Anthoathecata: Oceaniidae

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Anthoathecata: Pandeidae           Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

Cnidaria: Hydrozoa:                Fraser 1938, Kelmo & Vargas
Anthoathecata: Pennariidae         2002

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Anthoathecata: Porpitidae          Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Anthoathecata: Protiaridae         Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Anthoathecata: Trichydridae        Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Anthoathecata: Tubulariidae        Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz & Gasca 2009
Calycophorae: Abylidae

                                   Rodriguez-Saenz & Gasca 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz & Gasca 2009
Calycophorae: Dyphidae

                                   Rodriguez-Saenz & Gasca 2009

                                   Rodriguez-Saenz & Gasca 2009

                                   Rodriguez-Saenz & Gasca 2009

                                   Rodriguez-Saenz & Gasca 2009

                                   Rodriguez-Saenz & Gasca 2009

                                   Rodriguez-Saenz & Gasca 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz & Gasca 2009
Calycophorae: Prayidae

                                   Rodriguez-Saenz & Gasca 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Leptothecata: Aequoridae           Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Leptothecata: Campanulariidae      Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Leptothecata:                      Segura-Puertas 2009
Dipleurosomatidae

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Leptothecata: Eutimidae            Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

Cnidaria: Hydrozoa:                Fraser 1938
Leptothecata: Haleciidae

                                   Fraser 1938

Cnidaria: Hydrozoa:                Rodriguez-Saenz & Segura-
Leptothecata: Laodiceidae          Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Leptothecata: Lovenellidae         Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Leptothecata: Malagazziidae        Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Leptothecata: Melicertidae         Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Limnomedusae: Olindiidae           Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Leptothecata: Phialellidae         Segura-Puertas 2009

Cnidaria: Hydrozoa:                Fraser 1938
Leptothecata: Plumulariidae

Cnidaria: Hydrozoa:                Fraser 1938
Leptothecata: Sertulariidae
                                   Fraser 1948

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Narcomedusae: Aeginidae            Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Narcomedusae: Cuninidae            Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz & Gasca 2009
Physonectae: Agalmidae

                                   Rodriguez-Saenz & Gasca 2009

                                   Rodriguez-Saenz & Gasca 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz & Gasca 2009
Physonectae: Forskaliidae

Cnidaria: Hydrozoa:                Rodriguez-Saenz &
Trachymedusae: Geryoniidae         Segura-Puertas 2009

Cnidaria: Hydrozoa:                Rodriguez-Saenz & Segura-
Trachymedusae:                     Puertas 2009
Rhopalonematidae
                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

                                   Rodriguez-Saenz &
                                   Segura-Puertas 2009

Ctenophora                         Bednarski & Morales-Ramirez
                                   2004

Platyhelminthes: Cestoda:          Marques et al. 1996
Rhinebothriidea:
Rhinobothriidae

Platyhelminthes: Cestoda:          Marques et al. 1996
Tetraphyllidae:
Onchobothriidae

Platyhelminthes: Monogenea:        Meserve 1938, Rodriguez-Ortiz
Polyopisthocotylea:                et al. 2004
Protomicrocotylidae

Platyhelminthes: Trematoda:        Manter 1940, Rodriguez-Ortiz
Plagiorchiida: Bucephalidae        et al. 2004

Platyhelminthes: Trematoda:        Perez-Ponce de Leon et al.
Plagiorchiida: Didymozoidae        1998, Rodriguez-Ortiz et al.
                                   2004

Platyhelminthes: Trematoda:        Manter 1940, Rodriguez-Ortiz
Plagiorchiida: Fellodistomidae     et al. 2004

Platyhelminthes: Trematoda:        Perez-Ponce de Leon et al.
                                   1998,

Plagiorchiida: Lepocreadiidae      Rodriguez-Ortiz et al. 2004

Platyhelminthes: Trematoda:        Manter 1940, Rodriguez-Ortiz
Plagiorchiida: Monorchiidae        et al. 2004

Platyhelminthes: Trematoda:        Manter 1940, Rodriguez-Ortiz
Plagiorchiida: Opecoelidae         et al. 2004

Acanthocephala:                    Van Cleave 1940, Monks et al.
Palaeacanthocephala:               1997
Echinorhynchida: Illiosentidae

                                   Van Cleave 1940

Nematoda: Secernentea:             Hoberg et al. 1998
Spirurida: Gnathostomatidae

Annelida: Polychaeta:              Augener 1922, Dean 2004
Amphinomida: Amphinomidae

                                   Hartman 1940, Dean 2004

Annelida: Polychaeta:              Hartman 1944b, Dean 2004
Eunicida: Lumbrineridae

Annelida: Polychaeta:              Hartman 1944a, Dean 2004
Sabellida: Sabellariidae

Sipunculida:                       Spongberg 2006
Phascolosomatidea:

Phascolosomatida:                  Spongberg 2006
Phascolosomatidae

Mollusca: Bivalvia: Arcoida:       Rost 1955
Arcidae
                                   Rost 1955

                                   Hertlein & Strong 1943

                                   Hertlein & Strong 1943

                                   Hertlein & Strong 1943, Rost
                                   1955

                                   Hertlein & Strong 1943

                                   Hertlein & Strong 1943

                                   Hertlein & Strong 1943, Rost
                                   1955

Mollusca: Bivalvia: Arcoida:       Hertlein & Strong 1943
Glycymerididae

Mollusca: Bivalvia: Myoida:        Hertlein & Strong 1950
Corbulidae

Mollusca: Bivalvia: Myoida:        Soot-Ryen 1955
Mytilidae

                                   Soot-Ryen 1955

                                   Soot-Ryen 1955

                                   Soot-Ryen 1955

                                   Soot-Ryen 1955

                                   Soot-Ryen 1955

                                   Soot-Ryen 1955

Mollusca: Bivalvia: Veneroida:     Hertlein & Strong 1946b
Cardiidae

                                   Hertlein & Strong 1947

                                   Hertlein & Strong 1947

Mollusca: Bivalvia: Veneroida:     Hertlein & Strong 1949b
Donacidae
                                   Hertlein & Strong 1949b

                                   Hertlein & Strong 1949b

Mollusca: Bivalvia: Veneroida:     Skoglund 2001
Lasaeidae
                                   Skoglund 2001

                                   Skoglund 2001

Mollusca: Bivalvia: Veneroida:     Hertlein & Strong 1950
Psammobiidae

Mollusca: Bivalvia: Veneroida:     Hertlein & Strong 1950
Solecurtidae

                                   Hertlein & Strong 1950

                                   Hertlein & Strong 1950

                                   Hertlein & Strong 1950

Mollusca: Bivalvia: Veneroida:     Hertlein & Strong 1949a
Tellinidae

                                   Hertlein & Strong 1949a

                                   Hertlein & Strong 1949a

Mollusca: Bivalvia: Veneroida:     Hertlein & Strong 1947
Ungulinidae

                                   Hertlein & Strong 1947

Mollusca: Bivalvia: Veneroida:     Hertlein & Strong 1948
Veneridae

                                   Hertlein & Strong 1948

                                   Hertlein & Strong 1948

                                   Hertlein & Strong 1948

                                   Hertlein & Strong 1948

                                   Hertlein & Strong 1948

                                   Hertlein & Strong 1948

                                   Hertlein & Strong 1948

                                   Hertlein & Strong 1948

                                   Hertlein & Strong 1948

                                   Hertlein & Strong 1948

Mollusca: Bivalvia: Ostreoida:     Hertlein & Strong 1946a
Ostreidae

Mollusca: Bivalvia: Pectinoida:    Hertlein & Strong 1946a
Pectinidae

                                   Hertlein & Strong 1946a

                                   Hertlein & Strong 1946a

Mollusca: Bivalvia:                Hertlein & Strong 1943
Pectinoida: Spondylidae

Mollusca: Bivalvia:                Cortes 2001
Pterioidea: Pinnidae               Dawson & Beaudette 1959

Mollusca: Bivalvia:                Hertlein & Strong 1943
Pterioidea: Pteriidae

Mollusca: Gastropoda:              Valdes y Camacho-Garcia 2004
Cephalaspidea: Bullidae

Mollusca: Gastropoda:              Valdes y Camacho-Garcia 2004
Cephalaspidea: Cylichnidae

Mollusca: Gastropoda:              Valdes y Camacho-Garcia 2004
Cephalaspidea: Cylichnidae

Mollusca: Gastropoda:              Valdes y Camacho-Garcia 2004
Cephalaspidea: Haminoeidae

Mollusca: Gastropoda:              Valdes y Camacho-Garcia 2004
Cephalaspidea: Retusidae

Mollusca: Gastropoda:              Valdes y Camacho-Garcia 2004
Cephalaspidea: Rhizoridae

Mollusca: Gastropoda:              Valdes y Camacho-Garcia 2004
Cephalaspidea: Rhizoridae

Mollusca: Gastropoda:              Skoglund 2002
Littorinimorpha: Caecidae

                                   Skoglund 2002

Mollusca: Gastropoda:              Reid 2002
Littorinimorpha: Littorinidae

Mollusca: Gastropoda:              Cortes 2001
Littorinimorpha: Strombidae

Mollusca: Gastropoda:              Skoglund 2002
Littorinimorpha: Tornidae

Mollusca: Gastropoda:              Jung 1989
Neogastropoda: Columbellidae

                                   Jung 1989

                                   Jung 1989

                                   Jung 1989

                                   Jung 1989

                                   Jung 1989

Mollusca: Gastropoda:              Hanna & Strong 1949
Neogastropoda: Conidae
                                   Hanna & Strong 1949

                                   Hanna & Strong 1949

                                   Hanna & Strong 1949

                                   Hanna & Strong 1949

                                   Hanna & Strong 1949

                                   Hanna & Strong 1949

                                   Hanna & Strong 1949

                                   Hanna & Strong 1949

                                   Hanna & Strong 1949

                                   Hertlein & Strong 1951

Mollusca: Gastropoda:              Hertlein & Strong 1951
Neogastropoda: Fasciolariidae

Mollusca: Gastropoda:              Cernohorsky 1976
Neogastropoda: Mitridae

Mollusca: Gastropoda:              Berry 1958
Neogastropoda: Muricidae

Mollusca: Gastropoda:              Madrigal-Castro et al. 1984,
Neogastropoda: Muricidae           Gravel et al. 2006

Mollusca: Gastropoda:              Dawson & Beaudette 1959
Neogastropoda: Terebridae

Mollusca: Gastropoda:              McLean & Poorman 1971
Neogastropoda: Turridae

                                   Hertlein & Strong 1951

Mollusca: Gastropoda               Cortes 2001
Neotaenioglossa: Litiopidae

Mollusca: Gastropoda:              Ajtai et al. 2003
Nudibranchia: Scyllaeidae

Arthropoda: Crustacea:             Barnard 1954
Malacostraca: Amphipoda:
Ampeliscidae

Arthropoda: Crustacea:             Barnard & Barnard 1982
Malacostraca: Amphipoda:
Phoxocephalidae

Arthropoda: Crustacea:             Vargas & Cortes 2006
Malacostraca: Decapoda:
Anomura: Diogenidae
                                   Vargas & Cortes 2006

Arthropoda: Crustacea:             McLaughlin 1981
Malacostraca: Decapoda:
Anomura: Paguridae

Arthropoda: Crustacea:             Haig 1960
Malacostraca: Decapoda:
Anomura: Porcellanidae
                                   Haig 1960

                                   Haig 1960

Arthropoda: Crustacea:             Haig 1968
Malacostraca: Decapoda:
Brachyura: Aethridae
                                   Haig 1968

                                   Haig 1960, 1968

Arthropoda: Crustacea:             Haig 1968, Gore 1982
Malacostraca: Decapoda:
Brachyura: Calappidae
                                   Haig 1960, 1968

                                   Haig 1960

                                   Haig 1960

                                   Haig 1960

                                   Haig 1960, 1968

                                   Haig 1960, 1968

                                   Haig 1960, 1968

                                   Rathbun 1937

                                   Garth 1966

                                   Rathbun 1937

Arthropoda: Crustacea:             Crane 1947
Malacostraca: Decapoda:
Brachyura: Dairidae

Arthropoda: Crustacea:             Rathbun 1937
Malacostraca: Decapoda:
Brachyura: Dromiidae

Arthropoda: Crustacea:             Garth 1958, 1959
Malacostraca: Decapoda:
Brachyura: Epialtidae
                                   Garth 1958

                                   Garth 1958

                                   Garth 1958

Arthropoda: Crustacea:             Garth 1966
Malacostraca: Decapoda:
Brachyura: Ethusidae               Rathbun 1937

Arthropoda: Crustacea:             Turkay 1970
Malacostraca: Decapoda:
Brachyura: Gecarcinidae

                                   Cabrera-Pena et al. 1994

Arthropoda: Crustacea:             Crane 1947
Malacostraca: Decapoda:
Brachyura: Grapsidae
                                   Crane 1947

                                   Crane 1947

                                   Crane 1947

Arthropoda: Crustacea:             Garth 1958
Malacostraca: Decapoda:
Brachyura: Inachidae
                                   Garth 1958

                                   Garth 1958, 1959

Arthropoda: Crustacea:             Garth 1958
Malacostraca: Decapoda:
Brachyura: Inachoididae

                                   Garth 1958

                                   Garth 1958

                                   Garth 1958, 1959

Arthropoda: Crustacea:             Rathbun 1937
Malacostraca: Decapoda:
Brachyura: Leucosiidae
                                   Rathbun 1937

                                   Rathbun 1937

                                   Rathbun 1937

                                   Rathbun 1937

                                   Garth 1966

Arthropoda: Crustacea:             Garth 1958
Malacostraca: Decapoda:
Brachyura: Majidae                 Garth 1958

                                   Garth 1958

                                   Garth 1958

                                   Garth 1958

                                   Garth 1958

                                   Garth 1958

                                   Garth 1958

Arthropoda: Crustacea:             Crane 1941b
Malacostraca: Decapoda:
Brachyura: Ocypodidae
                                   Crane 1941b

                                   Crane 1975

                                   Crane 1941a, 1975

Arthropoda: Crustacea:             Crane 1947
Malacostraca: Decapoda:
Brachyura: Oziidae

Arthropoda: Crustacea:             Garth 1958
Malacostraca: Decapoda:
Brachyura: Parthenopidae
                                   Garth 1958

                                   Garth 1959

                                   Garth 1958

                                   Garth 1958

                                   Garth 1958

Arthropoda: Crustacea:             Crane 1947
Malacostraca: Decapoda:
Brachyura: Panopeidae
                                   Crane 1947

                                   Garth 1940, 1961

                                   Garth 1961, 1966

                                   Crane 1947

                                   Crane 1947

Arthropoda: Crustacea:             Crane 1947
Malacostraca: Decapoda:
Brachyura: Pilumnidae              Crane 1947

Arthropoda: Crustacea:             Garth 1961, 1966
Malacostraca: Decapoda:
Brachyura: Portunidae
                                   Garth 1961, 1966

                                   Garth 1961, 1966

                                   Garth 1961, 1966

Arthropoda: Crustacea:             Rathbun 1937
Malacostraca: Decapoda:
Brachyura: Raninidae

Arthropoda: Crustacea:             Crane 1947, Castro 1996
Malacostraca: Decapoda:
Brachyura: Trapeziidae
                                   Castro 1996

                                   Castro 1996

Arthropoda: Crustacea:             Garth 1961, 1966
Malacostraca: Decapoda:
Brachyura: Xanthidae
                                   Crane 1947

                                   Crane 1947

                                   Crane 1947

                                   Crane 1947

Arthropoda: Crustacea:             Kim & Abele 1988
Malacostraca: Decapoda:
Caridea: Alpheidae

Arthropoda: Crustacea:             Wicksten 1983
Malacostraca: Decapoda:
Caridea: Palaemonidae
                                   Wehrtmann & Vargas 2003

                                   Wicksten & Hendrickx 1992

                                   Cabrera-Pena & Solano-Lopez
                                   1996

                                   Wicksten & Hendrickx 1992

Arthropoda: Crustacea:             Hendrickx & Wicksten 1989
Malacostraca: Decapoda:
Caridea: Pandalidae

Arthropoda: Crustacea:             Brusca & Iverson 1985
Malacostraca: Isopoda:
Cirolanidae

Arthropoda: Crustacea:             Schmitt 1940, Manning 1971
Malacostraca: Stomatopoda:
Gonodactylidae

                                   Manning 1971, 1974

Arthropoda: Crustacea:             Manning 1971, 1974
Malacostraca: Stomatopoda:
Squillidae
                                   Manning 1971

Arthropoda: Crustacea:             Bednarski & Morales-Ramirez
Maxillopoda: Calanoida:            2004
Acartiidae
                                   Bednarski & Morales-Ramirez
                                   2004

Arthropoda: Crustacea:             Bednarski & Morales-Ramirez
Maxillopoda: Calanoida:            2004
Calanidae

Arthropoda: Crustacea:             Wilson 1937
Maxillopoda: Calanoida:
Caligidae                          Bednarski 2001

                                   Bednarski 2001

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda: Calanoida:
Calocalanidae

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda: Calanoida:
Candaciidae

                                   Bednarski 2001

Arthropoda: Crustacea:             Bednarski & Morales-Ramirez
Maxillopoda: Calanoida:            2004
Centropagidae

                                   Bednarski 2001

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda: Calanoida:
Clausocalanidae

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda: Calanoida:
Eucalanidae
                                   Bednarski 2001

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda: Calanoida:
Euchaetidae
                                   Bednarski 2001

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda: Calanoida:
Lucicutiidae

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda: Calanoida:
Paracalanidae
                                   Bednarski 2001

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda: Calanoida:
Pontellidae                        Bednarski 2001

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda: Calanoida:
Pseudodiaptomidae

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda: Calanoida:
Scolecitrichidae

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda: Calanoida:
Temoridae

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda: Calanoida:
Rhincalanidae

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda: Calanoida:
Subeucalanidae
                                   Bednarski 2001

Arthropoda: Crustacea:             Bednarski & Morales-Ramirez
Maxillopoda: Cyclopoida:           2004
Oithonidae
                                   Bednarski 2001

                                   Bednarski 2001

Arthropoda: Crustacea:             Suarez-Morales & Morales-
Maxillopoda: Mostrilloida:         Ramirez 2003
Monstrillidae

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda:
Poecilostomatoida:
Clausidiidae

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda:
Poecilostomatoida: Corycaeidae     Bednarski 2001

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda:
Poecilostomatoida: Oncaeidae
                                   Bednarski 2001

Arthropoda: Crustacea:             Bednarski 2001
Maxillopoda:
Poecilostomatoida:                 Bednarski 2001
Sapphirinidae
                                   Bednarski 2001

                                   Bednarski 2001

Arthropoda: Crustacea:             Henry & McLaughlin 1967
Maxillopoda: Sessilia:
Balanidae
                                   Henry & McLaughlin 1975

                                   Henry & McLaughlin 1975

Arthropoda: Crustacea:             Wilson 1937
Maxillopoda:
Siphonostomatoida:
Lernanthropidae

Arthropoda: Crustacea:             Bednarski & Morales-Ramirez
Branchiopoda: Diplostraca:         2004
Cladocera

Arthropoda: Crustacea:             Bednarski & Morales-Ramirez
Ostracoda                          2004

Arthropoda: Chelicerata:           Childs 1979
Pycnogonida: Pantopoda:
Phoxichilidiidae

Bryozoa: Gymnolaemata:             Osburn 1950
Cheilostomatida: Beaniidae

Bryozoa: Gymnolaemata:             Osburn 1952
Cheilostomatida:
Buffonellodidae

Bryozoa: Gymnolaemata:             Osburn 1952
Cheilostomatida:
Bryocryptellidae

Bryozoa: Gymnolaemata:             Osburn 1952
Cheilostomatida:
Mamilloporidae

Bryozoa: Gymnolaemata:             Osburn 1950
Cheilostomatida:
Membraniporidae

Bryozoa: Gymnolaemata:             Osburn 1952
Cheilostomatida:
Microporellidae

Bryozoa: Gymnolaemata:             Osburn 1952
Cheilostomatida: Petraliidae

Bryozoa: Gymnolaemata:             Osburn 1952
Cheilostomatida:
Schizoporellidae

Bryozoa: Gymnolaemata:             Osburn & Soule 1953
Ctenostomatida: Vesiculariidae

Echinodermata: Ophiuroidea:        Ziesenhenne 1940
Ophiurida: Ophiolepididae

Echinodermata: Asteroidea:         Clark 1940
Paxillosida: Astropectinidae

Echinodermata: Echinoidea:         Jimenez 1998
Cidaroida: Cidaridae

Echinodermata: Echinoidea:         Clark 1948
Clypeasteroida: Mellitidae

Echinodermata: Echinoidea:         Alvarado 2008
Diadematoida: Diadematidae

                                   Jimenez 1998

Echinodermata: Echinoidea:         Clark 1948
Temnopleuroida: Toxopneustidae

Echinodermata: Holothuroidea:      Deichmann 1958
Aspidochirota: Holothuriidae

                                   Deichmann 1958

                                   Deichmann 1958

                                   Deichmann 1958

Chaetognatha: Sagittoidea:         Bednarski & Morales-Ramirez
Sagittoidea: Sagittidae            2004

Chordata: Appendicularia:          Castellanos et al. 2009
Copelata: Oikopleuridae

                                   Castellanos et al. 2009

                                   Castellanos et al. 2009

Chordata: Appendicularia:          Castellanos et al. 2009
Copelata: Fritillariidae

Chordata: Thaliacea: Salpida:      Bednarski & Morales-Ramirez
Salpidae                           2004

Chordata: Chondrichthyes:          Dominici-Arosemena et al. 2005
Orectolobiformes:
Ginglymostomatidae

Chordata: Chondrichthyes:          Marques et al. 1996
Rajiformes: Dasyatidae

Chordata: Chondrichthyes:          Dominici-Arosemena et al. 2005
Rajiformes: Myliobatidae

Chordata: Actinopterygii:          Beebe 1942, Pfeiler 2008
Albuliformes: Albulidae

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Anguilliformes: Muraenidae
Beryciformes: Holocentridae        Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Syngnathiformes: Fistulariidae

Chordata: Actinopterygii:          Bussing & Lopez 2009
Syngnathiformes: Syngnathidae

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Acanthuridae

Chordata: Actinopterygii:          Alvarado 2008
Perciformes: Apogonidae
                                   Alvarado 2008

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Blenniidae

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Manter 1940
Perciformes: Carangidae
                                   Manter 1940

                                   Wilson 1937

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Chaetodontidae

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Cirrhitidae

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Ephippidae

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Haemulidae

                                   Dominici-Arosemena et al. 2005

                                   Perez-Ponce de Leon et al.
                                   1998

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Kyphosidae
                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Labridae

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005,
                                   Alvarado 2008

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005
                                   Alpermann 2001

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Lutjanidae            Alvarado 2008

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Malacanthidae

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Pomacanthidae

                                   Dominici-Arosemena et al. 2005,
                                   Alvarado 2008

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Pomacentridae

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Scaridae

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Sciaenidae

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Scombridae

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Perciformes: Serranidae

                                   Alvarado 2008

                                   Dominici-Arosemena et al. 2005

                                   Alvarado, 2008

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Fritzsche 1980
Syngnathiformes: Syngnathidae

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Tetraodontiformes: Balistidae

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Tetraodontiformes: Diodontidae

                                   Dominici-Arosemena et al. 2005

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Tetraodontiformes:
Monocanthidae

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Tetraodontiformes: Ostraciidae

Chordata: Actinopterygii:          Dominici-Arosemena et al. 2005
Tetraodontiformes:
Tetraodontidae
                                   Dominici-Arosemena et al. 2005

                                   Dominici-Arosemena et al. 2005

Chordata: Reptilia: Squamata:      Taylor 1953, Tu 1976
Elapidae

Chordata: Mammalia: Cetacea:       May-Collado 2009
Balaenopteridae

Chordata: Mammalia: Cetacea:       Rodriguez-Saenz & Rodriguez-
Delphinidae                        Fonseca 2004

                                   May-Collado & Morales-Ramirez
                                   2005

                                   May-Collado et al. 2005

as = As reported in the original publication.

(?) = Doubtful record.

(a) = species reported by Crouch & Poag (1987) but not found in any
database.

TABLE 2

Number species per taxonomic group

                         Number of species

Cyanobacteria                    5
Bacillariophyta                  3
Dinoflagellata                  14
Chlorophyta                     10
Rhodophyta                      28
Ochrophyta                       6
Magnoliophyta                    5
Protozoa: Foraminifera          33
Cnidaria                        100
  Anthozoa                      23
  Scyphozoa                      2
  Hydrozoa                      75
Ctenophora                       1
Plathyhelminthes                 9
Acanthocephala                   2
Nematoda                         1
Annelida                         4
Sipunculida                      2
Mollusca                        93
  Bivalvia                      55
  Gastropoda                    38
Crustacea                       147
  Malacostraca                  101
    Decapoda                    94
  Maxillopoda                   44
  Other Crustacea                2
Pycnogonida                      1
Bryozoa                          9
Echinodermata                   11
Chaetognatha                     1
Chordata                        92
  Fishes                        82
TOTAL                           577

Fig. 1. New reports
or species described
of marine species for
Bahia Culebra per decade.

1920-29     1
1930-39    29
1940-49   110
1950-59    85
1960-69    41
1970-79    10
1980-89    28
1990-99    26
2000-09   247

Note: Table made from line graph.
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Article Details
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Title Annotation:articulo en ingles
Author:Cortes, Jorge; Vargas-Castillo, Rita; Nivia-Ruiz, Jaime
Publication:Revista de Biologia Tropical
Date:Dec 1, 2012
Words:12104
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