First record of the trapdoor spider genus Conothele (Araneae, Ctenizidae) from India, with a description of two new species.
The genus Conothele was erected by Thorell in 1878 to accommodate Conothele malayana (Doleschall 1859), which was originally misplaced in the genus Cteniza Latreille 1829. Since then many new species of Conothele have been described (Thorell 1881, 1887, 1890; Pocock 1898, 1899; Kulczyn'ski 1908; Strand 1913; Hogg 1914; Chamberlin 1917; Berland 1938; Saaristo 2002; Platnick 2008). In his revision of the infraorder Mygalomorphae, Raven (1985) synonymized the genus Lechrictenus Chamberlin 1917 with Conothele. Some species of Ummidia Thorell 1875 were transferred to Conothele (Raven 1985; Haupt 2006). The genus Conothele is known from 16 species and has a wide distribution range extending from Myanmar to Australia (Platnick 2008).
In this paper, two new species of Conothele are described based on female specimens collected from Orissa and Arunachal Pradesh. Males of the species could not be found even after repeated searching in the area where females were collected. Considering the significance of this find, we formally report the occurrence of the genus Conothele from India. It also confirms the earlier reports by Gravely (1915, 1935) of the existence of this genus in India. Information on the habitat, behavior, and burrow of the two new species are provided. Interestingly, in the present study, these spiders were found strictly ground dwelling though previously the genus was considered arboreal in habit (Pocock 1900; Gravely 1935). We provide information on the burrow structure of these spiders, which is unique in comparison with other members of this genus.
The two new species from India are considered different from the rest of the known species of Conothele described from various islands of Australasia because of the geographical distribution. Geographically, the species closest in distribution to this new species is Conothele birmanica Thorell 1887 from Myanmar. RR examined the type specimen of C. birmanica at The Natural History Museum, London but spermathecae was not dissected, hence the information is lacking in this paper. Comparison with C. birmanica and C. taiwanensis Tso et al. 2003 is based only on available literature (Pocock 1900; Tso et al. 2003).
Spiders were collected during the theraphosid spider surveys conducted in the year 2005 and 2007 in Arunachal Pradesh and Orissa respectively. The specimens are deposited at Wildlife Information Liaison Development Society, Coimbatore, Tamil Nadu. Measurements of body parts except for the eyes were taken with a MitutoyoTM Vernier Caliper. Eye measurements were done with a calibrated ocular micrometer. All measurements are in mm. Spermathecae were dissected and cleaned in concentrated lactic acid in 100[degrees]C water bath for 15-20 minutes. All illustrations were prepared with the help of camera lucida attached to a CETII[TM] stereomicroscope by MS.
Abbreviations: ALE = anterior lateral eye, AME = anterior median eye, MOQ = median ocular quadrate, PLE = posterior lateral eye, PME = posterior median eye, PLS = posterior later spinnerets, PMS = posterior median spinnerets, WILD = Wildlife Information Liaison Development Society. Abbreviations used for hair and spines count are d = dorsal, fe = femur, mt = metatarsus, p = prolateral, pa = patella, r = retrolateral, ta = tarsus, ti = tibia, v = ventral.
Conothele Thorell 1878
Conothele Thorell 1878:303; Simon 1892:88; Pocock 1900:165
Type.--Conothele malayana (Doleschall 1859) designated based on a female specimen. Not examined.
Diagnosis.--It differs from other known genera of this family by trochanters I and II ventrally not notched (Raven 1985); tibia III consists of saddle-shape depression on the basal upper part (Figs. 8, 23) as seen in Ummidia (Pocock 1900; Raven 1985).
Conothele varvarti new species (Figs. 1-15)
Type specimens.--INDIA: Orissa: holotype female, Barehipani road, Chahala range, Similipal Tiger Reserve, 744 m elev., 21[degrees]57'46.2" N, 86[degrees]20'23.4"E, 31 March 2007, Manoj V. Nair (WILD-07-ARA-163); Orissa, 1 paratype female, Ramthirtha, Jashipur, 419 m elev., 21[degrees]57'08.4"N, 86[degrees]04'14.0"E, 29 August 2007, M. Siliwal (WILD-07-ARA-207).
Other material examined.--INDIA: Orissa: 1 subadult ?, Barehipani road, Chahala range, Similipal Tiger Reserve, 744 m elev., 21[degrees]57'46.2"N, 86[degrees]20'23.4"E, 31 March 2007, M. Nair (WILD-07-ARA-164); 2 immatures, Baniabasa, Udala range, Similipal Tiger Reserve, 545 m elev., 21[degrees]44'03.2"N, 86[degrees]26'51.8"E, 27 April 2007, Manoj V. Nair (WILD-07-ARA-165, 166); 2 immatures, before check gate of Jacum range, periphery of Karlapat Wildlife Sanctuary, 336 m elev., 19[degrees]44'49.3"N, 83[degrees]06'27.4"E, 14 April 2007, M. Siliwal, S. Behera (WILD-07-ARA-167, 168).
Diagnosis.--It differs from other known species of this genus by the posterior row of eyes weakly procurved (Fig. 3), (PRE straight in C. birmanica; PRE recurved C. taiwanensis); abdomen dorsally warty/rough (Fig. 1); spermathecae with bowl-shaped apical lobe on a stalk, which is twisted twice distally (Fig. 12) and carapace slightly shorter than patella and tibia of leg I and IV. It differs from C. brimanica by curved spines on tibiae I-II (Fig. 7).
Etymology.--The name of the species is based on the Sanskrit root of the word wart, referring to the warty appearance of the spider's abdomen.
Description of female holotype.--Total length, 15.52; carapace 5.52 long, 5.1 wide; chelicerae 2.42 intact. Abdomen 10.0 long, 6.84 wide. Spinnerets: PMS, 0.9 long, 0.3 wide, 0.1 apart; PLS, 2.0 total length (1.2 basal, 0.5 middle, 0.3 distal; midwidths 1.1, 0.8, 0.65 respectively). Morphometry of legs and palp is given in Table 1.
Carapace: reddish-brown, glabrous except for five long hairs on caput, few lines of depression along interstitial ridges (Figs. 1, 2), weak crenulations on caput, more conspicuous near eye group and anteriolaterally, elsewhere absent or inconspicuous. Caput with distinct mound between fovea and eyes (Fig. 2). Fovea deep, procurved, U-shaped (Fig. 1).
Eyes (Fig. 3): eight in two rows, both rows procurved, posterior row slightly procurved, ocular group 0.8 long, 1.2 wide, about 0.3 of headwidth; diameter AME 0.3, PME 0.15, ALE 0.4, PLE 0.25; distance between ALE-AME 0.1, AME-AME 0.05, PLE-PME adjacent, PME-PME 0.4, ALE-ALE 0.6, ALE-PLE 0.2; MOQ not square, 0.6 long, 0.6 front width, 0.7 back width; clypeus 0.6 high.
Maxillae (Fig. 4): 1.7 long in front, 2.5 long in back, 1.4 wide; cuspule numbers not same on right and left maxillae, right maxillae with 14 small cuspules on prolateral-proximal corner, 16 large ones ventrally in 3/4 of length, left maxillae with 8 small cuspules on prolateral-proximal corner, 17 large ventrally in % of length. Anterior lobe absent or greatly reduced.
Labium (Fig. 4): 1.0 long, 1.4 wide, labiosternal groove shallow and slightly concave, 5 large cuspules in three rows (2 + 2 + 1) centrally, size of cuspules similar to that on maxillae.
Chelicerae (Fig. 5): 4 large promarginal teeth, 5 large and 1 very small retromarginal teeth, basomesal teeth absent; rastellum conspicuous, raised on a low mound, consisting of 12 thick spines on vertical face and up, of which 10 are in anterior row; many long and short spines present along anterior dorsal surface.
Sternum (Fig. 4): broader posteriorly, reddish-brown, with elevated anterior and lateral sides, sloping posteriorly, 3.68 long, 3.2 wide, covered with long black hair, more dense towards lateral sides, posterior angle blunt and not separating coxae IV. Sigilla large, irregular shape, centrally placed. Nonsigillate area with fine corrugations.
Legs: all legs similar in thickness, reddish-brown above and light yellowish-brown below except tarsi of palp and metatarsi and tarsi of all legs that are black above and reddish-brown below. Tibiae, metatarsi and tarsi of leg I-II and tibiae and tarsi of palp dorsoventrally flattened. Femora III clearly wider than rest. Tibiae III with saddle-shaped depression on basal upper part. Metatarsi of leg I, II, IV longer than tarsi. Metatarsal preening combs absent on all legs. Coxae of legs reddish-brown ventrally. Legs covered with sparsely distributed hair, bristles, and few curved thick thorn-like spines (Figs. 6-8). Two conspicuous hairless bands running over length of femora, patellae, and tibiae. Scopulae and claw tufts absent on tarsi of all legs and palp (Figs. 6-8). Leg formula 4132.
Spines (Figs. 6-8): curved thick thorn-like spines, ti I, p = 38, r = 41; mt I, p = 34, r = 24; ta I, p = 26, r = 15; ti II, p = 26, r = 21; mt II, p = 30, r = 8; ta II, p = 23, r = 7; pa III, p = 10; ti III, p = 3; mt III, d = 4; ta III, p = 9, r = 3; mt IV, p = 3; ta IV, p = 6; palp, pa, p = 1; ti, p = 46 + 2 broken, r = 41; ta, p = 45+2 broken, r = 43.
Trichobothria: mt I-II with 10 filiform trichobothria in two rows in distal half; ta I with 4 clavate trichobothria centrally, 12 filiform in two longitudinal rows; ta II with 7 clavate in basal one fourth, 10 filiform in two longitudinal rows; ta III with 4 clavate in basal one fourth, 16 filiform in two longitudinal rows; ta IV with 2 clavate basal, 5 filiform in two rows in distal half; palp, ti with 10-12 filiform in two curved rows; ta with 5 clavate in center, 12 filiform in 2 longitudinal rows.
Leg coxae: coxa IV slightly wider than III, I and II subequal.
Claws: all legs with three claws, paired claw with single tooth (Fig. 9). Palp with single claw bearing a single unequal bifid tooth (Fig. 10).
Abdomen: grayish-brown, with few yellow small spots, covered with short and long thorn-like setae and several bristles, which gives it a warty appearance (Fig. 1); one in 6-8 thorns is long and is about 2-3 times longer and slightly thicker than short thorns. Ventrally yellowish, faint black patch above book lungs, uniformly covered with short and long bristles.
[FIGURES 1-5 OMITTED]
[FIGURES 6-15 OMITTED]
Spinnerets: PMS digitiform covered with brown hair; PLS, covered with brown hair, apical segment dome-shape (Fig. 11).
Spermathecae: bowl-shape apical lobe on a stalk, which is twisted twice distally (Fig. 12).
Description of female paratype (WILD-07-ARA-204).-- Total length 11.24; carapace 4.52 long 3.44 wide; chelicerae 1.36 long intact, 8 retromarginal and 4 promarginal teeth. Sternum 2.24 long, 2.0 wide. Labium 0.64 long, 0.96.2 wide, 9 large cuspules in 3 rows (5 + 2 + 2). Maxillae 2.06 long back, 1.28 long front, 1.08 wide, cuspules 20-24 of varying size in angular shape. Abdomen 6.72 long and 5.18 wide. Spinnerets: PMS, 0.6 long, 0.2 wide, 0.14 apart; PLS, 1.44 total length (0.72 basal, 0.42 middle, 0.3 distal; midwidths 0.8, 0.56, 0.34 respectively). Morphometry of leg and palp given in Table 1. Rest of the characters are same as holotype (WILD-07-ARA-163).
Distribution.--India, Orissa: Chahala, Upper Barahkamura, Jenabil and Baniabasa ranges in Similipal Tiger Reserve, Maurbhanj district; Ramthirtha in Jashipur; Karlapat Wildlife Sanctuary in Kalahandi district.
Individuals of C varvarti new species were found throughout Similipal Tiger Reserve, where more than 30 burrows of various sizes were located and almost all were found on mossy roadside cuttings close to the ground and up to 1.5 m, both in open and closed canopy areas. Similarly, several burrows of the species were located on roadside bunds in the periphery of Karlapat WLS in southern Orissa suggesting the species to be ground dwelling. This differs from what was previously reported by Pocock (1900) that the spiders of this genus from Myanmar and islands of Australasia were arboreal in habit with their trapdoor retreats on tree trunks. Also, Gravely (1935) reported a male of this genus in a retreat built in deodar Cedrus deodara humus, which he mentions as unusual since these spider retreats were generally found on tree trunks.
The burrow of C varvarti new species consists of a short silken tube about 60-70 mm long and 15-22 mm wide, and mostly aligned perpendicular to the angle of the slope of roadside bund (Figs. 13,14). The entrance of the burrow is closed with a moderately thick D-shaped hinged trapdoor (about 15 mm wide), with the attached side being straight rather than curved (Fig. 15). Also, the door hinge is mostly at the base or on the sides of the entrance but never at the top. The trapdoor is well camouflaged with soil and bits of dried leaves and moss. On the inner end of the burrow a second D-shaped hinged trapdoor exists, which opens into another chamber or retreat devoid of any silk lining (Figs. 13, 14). This unique burrow structure is probably a strategy to escape from intruders.
Conothele vali new species (Figs. 16-27)
Type specimen.--INDIA: Arunachal Pradesh: holotype female, near Shurbi village, Tawang District, 1856 m elev., 27[degrees]33'27.2"N, 91[degrees]53'41.5"E, 13 May 2005, M. Siliwal (WILD-05-ARA-77).
Diagnosis.--It differs from other known species of this genus in having posterior row of eyes weakly procurved (Fig. 18) as seen in C varvarti new species (PRE straight in C birmanica; PRE recurved C taiwanensis); eight cuspules on labium (Fig. 19) (5 cuspules of C brimanica and C varvarti new species; 14 cuspules in C taiwanensis); 3 small and 26-28 large cuspules on maxillae (Fig. 19) (8 small and 16-17 large cuspules in C varvarti; 53 cuspules in C taiwanensis); abdomen heavily wrinkled (Fig. 16) (warty in C varvarti, normal in C birmanica and C taiwanensis); spermathecae with globular apical lobe on a stalk, which is bent twice distally in zigzag manner (Fig. 27) (in C varvarti, stalk is twisted twice and anterior lobe is bowl- shape; stalk right-angled curve at apical globe in C taiwanensis); it differs from C brimanica by curved spines on tibiae I-II (Figs. 21, 22); it differs from C varvarti in leg formula 4123 (4132 in C varvarti, Table 2); paired claws on all the legs with two unequal teeth (Fig. 24) (single tooth on paired claw of legs I-III in C varvarti and C birmanica; row of teeth in C taiwanensis).
Etymology.--The species is derived from the Sanskrit word vali meaning wrinkle, as one of the characteristic features of this spiders is its heavily wrinkled abdomen.
Description of female holotype.--Total length, 11.30; carapace 5.92 long, 5.54 wide; chelicerae 1.76 intact. Abdomen 5.38 long, 4.4 wide. Spinnerets: PMS, 0.8 long, 0.3 wide, 0.5 apart; PLS, 1.5 total length (0.7 basal, 0.3 middle, 0.5 distal; midwidths 1.1, 0.9, 0.7 respectively). Morphometry of legs and palp is given in Table 2.
[FIGURES 16-20 OMITTED]
Carapace: reddish-brown, glabrous; black with net-like design, more prominent on the posterior half; thin black band present at periphery; few lines of depression along the interstitial ridges (Fig. 16), weak crenulations on caput, more conspicuous anteriolaterally and near eye region, elsewhere absent or inconspicuous. Caput with distinct mound between fovea and eyes, about twice higher than eyes (Figs. 16, 17). Fovea deep, procurved, U-shaped (Fig. 16). Bristles: 3 long and 18 short ones on caput; one long at anterior edge of ocular tubercle; 2 long and 2 short ones on clypeal plate.
Eyes (Fig. 18): eight in 2 rows, both rows slightly procurved; ocular group 1.0 long, 1.4 wide, about 0.4 of headwidth. Diameter AME 0.4, PME 0.2, ALE 0.6, PLE 0.3; distance between ALE-AME 0.15, AME-AME 0.1, PLE-PME adjacent, PME-PME 0.5, ALE-ALE 0.7, ALE-PLE 0.2; MOQ not square, 0.8 long, 0.8 front width, 1.0 back width; clypeus 0.7 high.
[FIGURES 21-27 OMITTED]
Maxillae (Fig. 19): 1.8 anterior length, 2.5 posterior length, 1.4 wide; cuspule numbers not same on right and left maxillae, right maxillae with 3 small ones on prolateral-proximal corner, 26 large ones ventrally in % of length, left maxillae with 3 small ones on prolateral-proximal corner, 28 large ones ventrally in 3/4 of length. Anterior lobe absent or greatly reduced.
Labium (Fig. 19): 0.9 long, 1.3 wide, shallow labiosternal groove slightly concave; a pair of rectangular black patches covers basal 1/4 of labium and labiosternal groove; 8 large cuspules in 2 rows (6 in anterior row and 2 behind centrally), size of cuspules similar to that of maxillae.
Chelicerae (Fig. 20): 3 large and one small promarginal teeth, 6 large and 2 very small retromarginal teeth, basomesal teeth absent; rastellum conspicuous, raised on a low mound, consist of 14 thick spines on vertical face and up, with 8 in anterior row; many long and short spines present along entire anterior dorsal surface.
Sternum (Fig. 19): broader posteriorly, reddish-brown, elevated anterior and lateral sides, sloping posteriorly, 3.22 long, 3.34 wide, covered with long black hair, more towards lateral sides, posterior angle blunt and not separating coxae IV. Sigilla large, irregular shape, centrally placed. Nonsigillate area with fine corrugations.
Legs: all legs similar in thickness, reddish-brown above and light yellowish-brown below except tarsi of palp and metatarsi and tarsi of all legs that are dark brown above and reddish-brown below. Tibiae, metatarsi and tarsi of leg I-II and tibiae and tarsi of palp dorsoventrally flattened. Femora III clearly wider than rest. Tibia III with saddle-shape depression on basal upper part. Metatarsi of leg I, II, IV longer than tarsi. Metatarsal preening combs on all legs absent. Coxae of legs yellowish-brown ventrally. Legs covered with sparsely distributed hair, bristles and few curved thick thorn-like spines (Figs. 21-23). Two conspicuous hairless bands running longitudinally on femora, patellae, and tibiae. Scopulae and claw tufts absent on tarsi of all legs and palp. Leg formula 4123.
Spines (Figs. 21-23): curved thick thorn-like spines present on leg I-II and palp, rest normal thick spines, ti I, p = 33, r = 39; mt I, p = 13 + 1 broken, r = 29; ta I, p = 20 + 1 broken, r = 15; ti II, p = 21, r = 13; mt II, p = 23, r = 15, v = 1; ta II, p = 17, r = 9; pa III, p = 10; ti III, p = 2; mt III, p = 2, d = 4, r = 3; ta III, p = 13, r = 6; mt IV, p = 2; ta IV, p = 11, v = 2; palp, pa, p = 1; ti, p = 35, r = 37; ta, p = 44, r = 41.
Trichobothria: mt I-II with 6 filiform trichobothria in two rows in distal half; mt III with 6 short filiform distally; mt IV with 14 short filiform for length; ta I with 5 clavate trichobothria in basal half, 10 filiform in two longitudinal rows; ta II with 5 clavate in basal half, 12 filiform in two longitudinal rows; ta III with 5 clavate in basal 1/4, 18 filiform in two longitudinal rows; ta IV with 4 clavate basal 1/4, 10 filiform in two longitudinal rows; palp, ti with 8 filiform in curved two rows; ta with 9 clavate centrally, 12 filiform in 2 rows in distal half.
Leg coxae: coxa IV slightly wider than III, I and II subequal.
Claws: all legs with three claws, paired claw with two unequal teeth (Fig. 24). Palp with single claw bearing single unequal bifid tooth (Fig. 25).
Abdomen (Fig. 16): grayish-black with few yellow small spots, wrinkled integument, covered with short and long thorn setae and several bristles; one in 6-8 thorns is long and is about twice longer and slightly thicker than the short thorns. Ventrally yellowish-black, uniformly covered with short and long bristles.
Spinnerets (Fig. 26): PMS digitiform covered with brown hair; PLS, covered with brown hair, apical segment dome-shape.
Spermathecae (Fig. 27): globular apical lobe on a stalk, which is bent (almost 45[degrees]) twice distally in zigzag manner.
Distribution.--Known only from the type locality, Tawang, Arunachal Pradesh, India
The spider was found in the open along a narrow trail and on the ground at about 2000 m above sea level. The habitat around was mostly secondary scrub with few pine trees, Pinus walichiana, growing nearby. The burrow of this spider could not be located as the spider was found in the open; probably the burrow was washed away due to the heavy rainfall at that time.
An important character which distinguishes Conothele from Ummidia is the presence of a single short tooth on paired claws of legs I-III (Raven 1985). The transfer of Ummidia taiwanensis Tso et al. 2003 and Ummidia fragaria (Donitz 1887) to the genus Conothele reveals that these species posses more than one dentition pattern on the paired claws of legs I-III, which does not match the generic key. Similarly, C vali new species also possesses two teeth on the paired claws rather than a single tooth as seen in C varvarti new species. The only character which can distinguish Conothele from Ummidia will be the absence of the notch on trochanters I-II. Therefore, there is a need to revise the diagnosis of genera, and the generic key of the family Ctenizidae.
The Rufford Maurice Laing Foundation and DEFRA / FFI Flagship Species Fund (project No. 06/16/02 FLAG) provided financial support to the Indian Tarantula Project, during which survey trips these trapdoor spiders were found. The authors are also grateful to the following personnel: Sally Walker, Zoo Outreach Organisation for her interest in the spider study; PCCF and Mr. C. Loma, Arunachal Pradesh Forest Department for giving the collecting permit and providing help to carry out spider surveys in different protected areas in Arunachal Pradesh; PCCF and Dr. S.K. Kar, Orissa Forest Department for giving the permit to carry out spider surveys in different protected areas in Orissa; Field Director of Similipal Tiger Reserve for providing necessary survey permits and logistics during the survey; Dr. A.S. Dippenaar-Schoeman, Agricultural Research Council, Pretoria and Suresh Kumar, Wildlife Institute of India, Dehra Dun for commenting on the first draft of this paper; Saroj Behera, Ganapati Sahu, Khandu, Dorjie Raptan, and Nima Shiring for their assistance during field work; Prof. M. Ganeshkumar, Tamil Nadu Agriculture University, Coimbatore, for providing technical support; and Varad Giri, for providing much needed literature on trapdoor spiders from the Bombay Natural History Society library.
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Manuscript received 19 November 2007, revised 28 July 2008.
Manju Siliwal: Wildlife Information Liaison Development Society, 9-A, Lal Bahadur Colony, Peelamedu, Coimbatore 641004, Tamil Nadu, India. E-mail: email@example.com
Manoj V. Nair: Similipal Tiger Reserve, Baripada 757001, Orissa, India
Sanjay Molur: Wildlife Information Liaison Development Society/ Zoo Outreach Organisation, 9-A, Lal Bahadur Colony, Peelamedu, Coimbatore 641004, Tamil Nadu, India
Robert Raven: Queensland Museum, Grey Street, PO Box 3300, South Brisbane, 4101, Queensland, Australia
Table 1.--Morphometry of legs and palp of the holotype female (WILD-07-ARA-163) and paratype female (WILD-07-ARA-207) of Conothele varvarti new species. Leg I Leg II Leg III Holo Para Holo Para Holo Para #163 #207 #163 #207 #163 #207 Femur 4.2 2.84 3.30 2.44 3.12 2.24 Patella 2.46 2.10 2.14 1.66 2.0 1.56 Tibia 3.12 2.18 1.9 1.42 1.74 1.78 Metatarsus 1.42 1.12 1.46 1.10 1.26 1.0 Tarsus 1.32 0.9 1.12 0.9 1.96 1.44 Total 12.52 9.14 9.92 7.52 10.08 8.02 Midwidth Femur 1.0 0.64 0.9 0.52 1.42 0.94 Tibia 1.06 0.72 0.96 0.68 1.0 0.72 Leg IV Palp Holo Para Holo Para #163 #207 #163 #207 Femur 4.0 3.12 4.24 3.34 Patella 2.36 2.14 2.16 1.44 Tibia 2.92 2.3 3.0 1.7 Metatarsus 2.42 1.86 -- -- Tarsus 1.72 0.88 2.3 1.8 Total 13.42 10.3 11.7 8.28 Midwidth Femur 1.2 0.64 1.1 0.62 Tibia 1.0 0.72 1.0 0.72 Table 2.--Morphometry of legs and palp of holotype female (WILD-05-ARA-77) of Conotele vali new species. Leg I Leg II Leg III Leg IV Palp Femur 4.12 3.58 3.22 4.1 3.48 Patella 2.30 2.30 2.14 2.24 2.32 Tibia 2.68 2.34 2.08 2.82 2.54 Metatarsus 2.06 1.56 1.74 2.56 -- Tarsus 1.24 1.34 1.88 1.52 2.28 Total 12.4 11.12 11.06 13.24 10.62 Midwidth Femur 0.88 0.9 1.48 0.9 0.9 Tibia 1.0 0.9 0.98 0.9 1.14
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|Author:||Siliwal, Manju; Nair, Manoj V.; Molur, Sanjay; Raven, Robert|
|Publication:||The Journal of Arachnology|
|Date:||Jan 1, 2009|
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