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Fire maintained, closed canopy barren communities in western Illinois.

ABSTRACT

Barrens were common in Illinois at the time of European settlement. These communities were fire-maintained, had an open canopy, and a grass-dominated ground layer containing both forest and prairie species. Barrens were commonly underlain by nutrient poor, clayey soils, and were found on rolling topography. Fire suppression following the arrival of European settlers resulted in canopy closure and the loss of many prairie species. The barrens studied are currently being managed by fire. Quercus alba (white oak) dominated the overstory and accounted for more than 65% of the importance value on both barrens. Other overstory species were Carya tomentosa (mockernut hickory), Q. stellata (post oak), and Q. velutina (black oak). Woody seedlings were common, but few shrubs and saplings were present, probably due to recurring fires. Overstory cover at Argyle Hollow Barrens Nature Preserve averaged 77%, while cover at McKee Creek Barrens averaged 85%. Numerous prairie species were found on these barrens but most were restricted to canopy openings.

INTRODUCTION

Barrens were described as open forest communities with a ground layer of prairie grasses and forbs, but also with an unusually high proportion of forest herbs (Ellsworth 1838, Vestal 1936). The woody vegetation of barrens consisted of stunted trees of Quercus stellata (post oak), Q. alba (white oak), Carya spp. (hickories), with the common shrubs being Corylus americana (hazel), Rhus glabra (smooth sumac), and R. copallina (winged sumac) (Peck 1837, Worthen 1868). Grubs, commonly described as brushy trees that had been repeatedly top-killed by recurring fires, were common (Peck 1837). In some instances the grubs were more than 100 years old and consisted of numerous basal branches that sometimes exceeded 3 meters in height (Curtis 1959).

Barrens were fire maintained communities. Bourne (1820) describes the disappearance of barrens after Native Americans left and fires were stopped. Worthen (1868, 1870) also described barrens as fire maintained communities, and Peck (1837) described the growth of vigorous sprouts from grubs once there were no more fires. By the 1860s it was realized that barrens were transient communities and, due to fire suppression, would soon be replaced by forest (Engelmann 1863). Presently few good quality examples of barrens exist in Illinois (Edgin 2000, Taft 2003, Edgin et al. 2005, McClain et al. 2007). Most have been degraded due to fire suppression and currently retain little of the species diversity and community structure that existed in the early 1800s.

In general, the few remaining barrens have been subjected to occasional fires, have very poor quality soils, have been relatively undisturbed by human activity, and are restricted to regions of rolling topography (Bowles and McBride 1994, Bowles et al. 1994, Ebinger et al. 1994, Homoya 1994, Taft 2003, Edgin et al. 2005). As this community is uncommon, attempts are being made to re-establish barrens where they previously existed. The present study was undertaken to determine the composition and structure of the vegetation of two barrens in western Illinois that are presently being managed as barren communities. According to Government Land Office (GLO) survey records, these two areas were barrens in early settlement times (Hutchinson 1988).

DESCRIPTION OF THE STUDY AREA

The barrens examined were located in the Western Forest-Prairie Natural Division, a strongly dissected glacial till plain subjected to the Illinoian stage of Pleistocene Glaciation approximately 125,000 years ago (Schwegman 1973). At the time of European settlement oak-hickory forests, woodlands, and barrens dominated the rugged topography associated with the well developed and extensive drainage systems of this Division (Anderson 1991). Prairies were also common, but mostly restricted to the level uplands (Anderson 1991).

The barrens studied were located on rolling topography and had many floristic similarities to woodlands and forests on dry-mesic to xeric sites. Both barrens were on steep, southwest-facing hillsides, were less than 2 ha in size, and had small canopy openings in which some prairie vegetation was present. Both would presently be classified as mature second-growth dry to dry-mesic upland forests using the classification of White and Madany (1978). The more open canopy, which varied from 77 to 85%, and the open understory were probably the result of recent fire management.

Argyle Hollow Barrens

This dedicated nature preserve is within Lake Argyle State Park about 2 kms north of Colchester, McDonough County, Illinois (NE1/4 S36 T6N R4W). The barrens community was located on the rolling uplands on the east side of Argyle Lake. Sandstone and shale outcrops were common on the steep slope with a sandstone cliff at the edge of the lake. The soils were classified as Hickory loam with 10 to 18% slope (Walker 1997).

McKee Creek Barrens

This natural area was in Siloam Springs State Park about 18 kms south of Clayton, Adams County, Illinois (SE1/4 S24 T2S R5W). The barren overlooks McKee Creek that forms the southern boundary of the park. A cliff at the base of the barrens was about 15 meters high. The soils were classified as Marseilles silt loam with 18 to 35% slope (Tegeler 2003).

The climate is continental, characterized by humid, hot summers and cold winters. Weather records for Quincy, Illinois, gives an average annual precipitation of 98 cm that falls mostly as rain from March through October (www.sws.uiuc.edu 2005). January is the coldest month with an average high temperature of 0[degrees]C and an average low of -9[degrees]C. July is the hottest month with an average low of 19'C and an average high of 30[degrees]C. The frost-free growing period averages 191 days with a low of 166 and a high of 232 days.

MATERIALS AND METHODS

The study sites were visited throughout the growing seasons of 1995 and 1996, and at least once each growing season since that time to complete the vascular plant species list. Voucher specimens of each species were collected, identified, and deposited in the Stover-Ebinger Herbarium of Eastern Illinois University, Charleston, Illinois (EIU). Criteria for designating exotic species followed Mohlenbrock (2002), and Gleason and Cronquist (1991), while nomenclature follows Mohlenbrock (2002).

During the late summer of 1996 the woody overstory was surveyed using a 25 m x 50 m plot. This plot was placed as near the middle of the site as possible to eliminate edge effect. In each quadrate all living woody individuals [greater than or equal to] 10.0 cm dbh were identified and their diameters recorded. From these data, the living-stem density (stems/ha), basal area ([m.sup.2]/ha), relative density, relative dominance, importance value (IV), and average diameter (cm) were calculated for each species. The IV was the sum of the relative density and relative dominance (basal area) (McIntosh 1957). Overstory cover was determined by photographing the canopy and projecting the photo onto a 100 point grid.

To study ground layer vegetation, two transects 25 m long were located randomly in each study area. Along each transect, 1 [m.sup.2] quadrates were located at 1 m intervals (n=25/transect), odd-numbered quadrates to the right even-numbered to the left. A random numbers table was used to determine the number of meters (0 to 9) a quadrate was located from the transect line. Cover was determined by using the Daubenmire cover class system (Daubenmire 1959) as modified by Bailey and Poulton (1968). The modified Daubenmire cover scale is as follows: class 1 = 0 to 1%; class 2 = >1 to 5%; class 3 = >5 to 25%; class 4 = >25 to 50%; class 5 = >50 to 75%; class 6 = >75 to 95%; class 7 = >95 to 100%. Importance value (IV) for ground layer species was determined by summing relative cover and relative frequency.

RESULTS

Within the barrens studied a total of 139 plant species representing 46 families and 98 genera were documented (Appendix I). Only one fern species was encountered, while 40 were monocots in eight families, and 98 were dicots in 37 families. Seven exotic species were found, none being common in the plots. Of the species encountered 114 were found on Argyle Hollow Barrens, 104 were collected from McKee Creek Barrens, and 82 were found on both barrens. Nearly 25% of the herbaceous species collected were associated with prairie vegetation (Appendix I).

Quercus alba dominated the overstories of both barrens, accounting for more than 65% of the IV (Table 1). At Argyle Hollow Barrens Carya tomentosa was second in IV, while on McKee Creek Barrens Q. stellata was second in IV. Quercus velutina (black oak) was third in IV on both barrens. The slightly more mesic McKee Creek Barrens had higher species diversity in the overstory, and a canopy closure of 85%. Very few dead-standing trees were encountered.

As a result of management with periodic fires, both barrens had a park-like appearance; the understory was open. Small saplings averaging between 800 and 1375 stems/ha and large saplings averaged 120 to 288 stems/ha on the barrens (Table 2). Numerous tree seedlings were present, however, and averaged between 15,000 and 17,500 stems/ha. Management fires top-killed most seedlings, but many had an enlarged caudex indicating that they re-sprouted after each fire. These sprouts rarely reached the sapling layer.

Ground layer vegetation was sparse on the study sites. Bare ground and litter had cover values of 62.14 and 67.09 on Argyle Hollow and McKee Creek barrens respectively (Table 3). On Argyle Hollow Barrens Parthenocissus quinquefolia (IV of 54.) dominated with Carex pensylvanica, Rubus flagellaris, Solidago ulmifolia, and Helianthus divaricatus being the common species encountered, having a combined IV of 148.0 (possible 200). Similar results were obtained on McKee Creek Barrens except these species differed somewhat in IV and Rubus flagellaris was replaced by Muhlenbergia sobolifera. On this barren the five dominant species had a combined IV of 110.2. The ground layer of both barrens consisted of a mixture of forest and prairie grasses and forbs (Table 3, Appendix I). Forest species were, by far, the most important, and the few species listed as "others" in the table included many of the prairie species encountered. Other prairie species were growing near the barren edges, or in very low frequencies on the barrens, and did not occurr in the survey quadrates.

DISCUSSION

At the time of European settlement in the early 1800s a broad mosaic of prairie and open-to closed-canopy oak-dominated communities (forest, woodland, savanna, barren) existed in Illinois (Davies 1977, Anderson 1983). Most forested areas persisted on the lee side of topographic and wetland fire breaks. Fire frequency and intensity were important in determining the composition and structure of these wooded areas. Intense and frequent fires created prairie and savanna, less intense and less frequent fires causing barrens and woodlands, while low intensity, infrequent fires allowed closed forests to persist (Ebinger and McClain 1991).

Based on early literature and GLO survey notes, it is evident that most upland forests in Illinois had open canopies (Vestal 1936, Anderson and Anderson 1975, Ebinger and McClain 1991). These open canopy forests (woodlands, barrens, savannas) represented a transition between prairies and closed-canopy forests of the dissected terrain of river valleys. These open woodlands, savannas, and barrens were fashioned by climate, topography, edaphic factors, and periodic fires (Heikens and Robertson 1994, McClain and Elzinga 1994). With the cessation of landscape fires soon after the arrival of European settlers, woody plant encroachment usually resulted in canopy closure except where edaphic factors slowed tree growth. Native aborigines were probably responsible for most of these fires (Williams 1989, Davies 1994, McClain and Elzinga 1994).

During the past 15 years attempts wave been underway to re-create the barren aspect at both Argyle Hollow and McKee Creek barrens. Occasional prescribed fires are being used to slowly open the canopy. Presently the ground layer is dominated by species associated with forest communities. Very few of the herbaceous species presently found in the ground layer of these two barrens are prairie species (Table 3). Also the shrub layer of sumac, hazel, and the stunted trees reported by the early GLO surveyors are lacking, as are the dense oak grubs (Vestal 1936). It is possible that under the present management of occasional prescribed fires, the "barrens of the early 1800s" cannot be attained on these sites. It is likely that more intense, landscape fires are necessary; the slow moving ground fires presently being used are not hot enough or intense enough. These ground fires are not killing the large canopy trees, and the canopy is still mostly closed. The continued management with fire, however, will slowly open the canopy and promote an increase in some prairie species. The year after the 1996 fire at the McKee Creek Barrens, Trifolium reflexum (buffalo clover) was found. Apparently the seeds of this Illinois endangered species had been lying dormant in the soil and the heat from the fire promoted germination.

APPENDIX I.

Vascular taxa encountered at the western Illinois barren communities listed alphabetically by family under major plant groups. An asterisk indicates non-native (exotic) species. For each species the author's collection number (JEE) is given followed by the barrens in which each species was collected (a = Argyle Hollow Barrens, m = McKee Creek Barrens. Nomenclature follows Mohlenbrock (2002).

FERN AND FERN-ALLIES

Ophioglossaceae

Botrychium virginianum (L.) Sw., 29643a, 29652m

MONOCOTS

Amaryllidaceae

Hyposis hirsuta (L.) Coville, 29644a

Araceae

Arisaema dracontium (L.) Schott., 30097a

Arisaema triphyllum (L.) Schott., 30794a, 30776m

Commelinaceae

Tradescantia ohiensis Raf., 30098a, 26181m

Cyperaceae

Carex albicans Willd., 26189m

Carex blanda Dewey, 30795a, 26187m

Carex cristatella Britt., 29651a

Carex hirsutella Mack., 26303a, 26190m

Carex muhlenbergii Schk., 26308a, 26192m

Carex pensylvanica Lam., 29457a, 29662m

Carex rosea Schk., 26191m

Juncuaceae

Juncus tenuis Willd., 30099a, 30106m

Liliaceae

Allium canadense L., 30796a

Smilacina racemosa (L.) Desf., 30797a, 29667m

Trillium recurvatum Beck, 29450a

Poaceae

Agrostis perennans (Walt.) Tuckerm., 30509a, 30491m

Andropogon gerardii Vitman, 30510a, 30492m

Bromus pubescens Muhl., 30100a, 30107m

Cinna arundinacea L., 31001a, 30493m

*Dactylis glomerata L., 30798a, 30777m

Danthonia spicata (L.) Roem. & Schultes, 26297a, 26173m

Dichanthelium acuminatum (Sw.) Gould & Clark, 26305a, 30111m

Dichanthelium latifolium (L.) Gould & Clark, 30104a, 26174m

Dichanthelium linearifolium (Scribn.) Gould, 26299a, 30781m

Elymus hystrix L., 30101a, 30108m

Elymus villosus Muhl., 30511a

Elymus virginicus L., 30102a, 30109m

*Festuca pratensis Huds., 30799a, 30778m

Festuca subverticillata (Pers.) E.B. Alexeev., 30103a, 30110m

Glyceria striata (Lam.) Hitchc., 30801a, 30780m

Muhlenbergia schreberi J.F. Gmel., 30512a, 30494m

Muhlenbergia sobolifera (Muhl.) Trin., 30513a, 30495m

*Poa compressa L. 30105a, 30112m

*Poa pratensis L. 29650a, 26172m

Poa sylvestris Gray, 26176m

Schizachyrium scoparium (Michx.) Nash, 30514a, 30496m

Sphenopholus obtusata (Michx.) Scribn, 30802a, 26175m

Tridens flavus (L.) Hitchc. 31191a, 30497m

Vulpia octoflora (Walt.) Rydb., 30782m

Smilacaceae

Smilax tamnoides L., 30515a, 30783m

DICOTS

Acanthaceae

Ruellia humilis Nutt., 30516a, 30113m

Anacardiaceae

Rhus aromatica Ait., 29663m

Rhus glabra L., 31014m

Toxicodendron radicans (L.) Kuntze, 30803a, 30784m

Apiaceae

Sanicula canadensis L., 30075a, 30114m

Taenidia integerrima (L.) Drude, 30813a, 29658m

Aristolochiaceae

Aristolochia serpentaria L., 31008a

Asclepiaadaceae

Asclepias quadrifolia Jacq., 26300a, 26182m

Asteraceae

Ageratina altissima (L.) King & Robins., 30522a, 30501m

Antennaria plataginifolia (L.) Hook., 29453a, 29653m

Aster anomalus Engelm., 30517a, 30498m

Aster lateriflorus (L.) Britt., 30519a, 30499m

Aster pilosus Willd., 30520a

Aster turbinellus Lindl., 30521a, 30500m

Aster urophyllus Lindl., 30518a

Coreopsis palmata Nutt., 30117m

Erigeron annuus (L.) Pers., 30077a, 30115m

Erigeron philadelphicus L., 30804a, 30785m

Eupatorium serotinum Michx., 31192a

Helianthus divaricatus L., 30076a, 30116m

Hieracium gronovii L., 31005a

Krigia biflora (Walt.) Blake, 29645a

Lactuca canadensis L., 31007a

Lactuca floridana (L.) Gaertn., 31009m

Liatris aspera Michx., 30523a, 30502m

Prenanthes alba L., 30524a

Solidago nemoralis Ait., 30526a, 30503m

Solidago speciosa Nutt., 31193a

Solidago ulmifolia Muhl., 30525a, 30504m

Berberidaceae

Podophyllum peltatum L., 30810a, 29665m

Boraginaceae

Hackelia virginiana (L.) I.M.Johnston, 30078a

Brassicaceae

Arabis canadensis L., 30805a, 29654m

Arabis laevigata (Willd.) Poir., 30079a

Caesalpiniaceae

Chamaecrista fasciculata (Michx.) Greene, 30082a

Campanulaceae

Campanulastrum americanum (L.) Small, 30080a, 31010m

Lobelia inflata L., 30081a, 30118m

Lobelia spicata Lam., 26311a, 30119m

Triodanis perfoliata (L.) Nieuwl., 30807a, 30786m

Caprifoliaceae

*Lonicera maackii (Rupr.) Maxim., 30527a

Viburnum prunifolium L., 30528a, 30505m

Caryophyllaceae

*Cerastium glomeratum Thuill., 26183m

Convolvulaceae

Calystegia spithamaea (L.) Pursh, 30806a

Corylaceae

Corylus americana Walt., 30083a

Elaeagnaceae

*Elaeagnus umbellata Thunb., 30529a, 30506m

Eupohorbiaceae

Acalypha virginica L., 31003a, 30507m

Euphorbia corollata L., 31002a, 31011m

Fabaceae

Amorpha canescens Pursh, 26307a, 30120m

Amphicarpaea bracteata (L.) Fern., 30787m

Dalea candida (Michx.) Willd., 26312a, 30123m

Desmodium glutinosum (Muhl.) A. Wood, 30084a, 30122m

Desmodium nudiflorum (L.) DC., 30121m

Lespedeza intermedia (S. Wats.) Britt., 31196m

Lespedeza virginica (L.) Britt., 31194a, 30608m

Tephrosia virginiana (L.) Pers., 30126m

Trifolium reflexum L., 26186m

Grossulariaceae

Ribes missouriense Nutt., 29449a

Hypericaceae

Hypericum punctatum Lam., 26310a

Lamiaceae

Pycnanthemum pilosum Nutt., 30530a

Pycnanthemum tenuifolium Schrad., 26306a, 30124m

Teucrium canadense L., 30085a

Menispermaceae

Menispermum canadense L., 30808a

Onagraceae

Circaea lutetiana L., 30086a

Oxalidaceae

Oxalis fontana Bunge, 30087a

Oxalis violacea L., 29452a, 29656m

Phrymaceae

Phryma leptostachya L., 30088a, 30125m

Polygalaceae

Polygala sanguinea L., 30089a

Portulacaceae

Claytonia virginica L., 29456a

Ranunculaceae

Anemone virginiana L., 30090a, 30127m

Anemonella thalictroides (L.) Spach, 29455a, 26179m

Ranunculus abortivus L., 30809a

Ranunculus micranthus Torr. & Gray, 29655m

Rhamnaceae

Ceanothus americanus L., 30128m

Rosaceae

Agrimonia gryposepala Wallr., 30091a, 30129m

Geum canadense Jacq., 30092a, 30130m

Potentilla simplex Michx., 26304a, 26184m

Rosa carolina L., 30093a, 30131m

Rubus allegheniensis Porter, 29647a

Rubus flagellaris Willd., 29646a, 29666m

Rubus occidentalis L., 30811a, 30788m

Rubus pensilvanicus Poir., 29670m

Rubiaceae

Galium aparine L., 29648a, 26185m

Galium circaezans Michx., 26301a, 30789m

Galium concinnum Torr. & Gray, 26298a, 30133m

Galium pilosum Ait., 30132m

Galium triflorum Michx., 31006m

Rutaceae

Ptelea trifoliata L., 31013m

Zanthoxylum americanum Mill., 30094a

Santalaceae

Comandra umbellata (L.) Nutt., 30095a

Scrophulariaceae

Agalinus tenuifolia (Vahl) Raf.,30531a, 31197m

Aureolaria grandiflora (Benth.) Pennell, 30532a, S31012m

Penstemon pallidus Small, 29649a, 26178m

*Veronica arvensis L., 30812a, 30790m

Solanaceae

Physalis subglabrata Mack. & Bush, 26180m

Physalis virginiana Mill., 29668m

Violaceae

Viola palmata L., 31004a, 30793m

Viola pedata L., 29454a, 30791m

Viola sororia Willd., 29451a, 30792m

Vitaceae

Parthenocissus quinquefolia (L.) Planch, 30096a, 30134m

ACKNOWLEDGMENTS

The authors would like to thank the Illinois Department of Natural Resources for permission to examine the barrens at Argyle Hollow Nature Preserve at Lake Argyle State Park, and the McKee Creek Barrens at Siloam Springs State Park. Dr. Gorden Tucker, Eastern Illinois University, was very helpful in the identification of the Carex species.

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William E. McClain

Adjunct Research Associate in Botany

Illinois State Museum

Springfield, IL 62706

(fidheall@yahoo.com)

John E. Ebinger

Emeritus Professor of Botany

Department of Biological Sciences

Eastern Illinois University

Charleston, IL 61920

(cfjee@eiu.edu) (corresponding author)
Table 1. Density (#/ha), basal area ([m.sup.2]/ha), relative
values, importance values, and average diameters (cm) of the
tree species encountered at Argyle Hollow Barrens, McDonough
County, and McKee Creek Barrens, Adams County, Illinois.

Species Density Basal Rel.
 (#/ha) Area Den.
 ([m.sup.2/ha)

Argyle Hollow Barrens
Quercus alba 144 20.2 69.2
Carya tomentosa 32 0.7 15.4
Quercus velutina 16 0.2 7.7
Carya ovata 16 0.2 7.7
Totals 208 21.3 100.0

McKee Creek Barrens
Quercus alba 128 21.7 50.0
Querrus stellata 48 3.7 18.8
Querrus velutina 24 0.5 9.4
Acer saccharum 24 0.2 9.4
Juglans nigra 8 0.1 3.1
Ostrya virginiana 8 0.1 3.1
Carya ovata 8 0.1 3.1
Fraxinus americana 8 0.1 3.1
Totals 256 26.5 100.0

Species Rel. I.V. Average
 Dom. Diameter
 (cm)

Argyle Hollow Barrens
Quercus alba 94.6 163.8 41.4
Carya tomentosa 3.3 18.7 16.1
Quercus velutina 1.1 8.8 13.7
Carya ovata 1.0 8.7 13.3
Totals 100.0 200.0

McKee Creek Barrens
Quercus alba 82.0 132.0 45.7
Querrus stellata 14.0 32.8 31.0
Querrus velutina 1.9 11.3 15.8
Acer saccharum 0.8 10.2 10.7
Juglans nigra 0.4 3.5 13.4
Ostrya virginiana 0.3 3.4 10.5
Carya ovata 0.3 3.4 12.0
Fraxinus americana 0.3 3.4 10.5
Totals 100.0 200.0

Table 2. Density (stems/ha) of woody seedlings, shrubs, small
saplings, and large saplings encountered at the Argyle Hollow
Barrens, McDonough County, and the McKee Greek Barrens, Adams
County, Illinois.

Species Seedlings Small Saplings Large Saplings

Argyle Hollow Barrens
Quercus alba 11250 125 --
Quercus imbricaria 1250 -- --
Viburnum prunifolium 1250 -- --
Quercus velutina 833 208 8
Prunus serotina 833 542 --
Amelanchier arborea 833 -- --
Sassfras albidum 833 -- --
Carya tomentosa 417 125 56
Ostrya virginiana -- 125 --
Fraxinus americana -- 125 --
Carya ovata -- 83 40
Cercis canadensis -- 42 --
Acer saccharum -- -- 16
Totals 17499 1375 120

McKee Creek Barrens
Viburnum prunifolium 4000 -- --
Quercus alba 2333 100 --
Quercus stellata 1667 100 8
Carya ovata 1667 133 64
Ostrya virginiana 1667 -- 120
Ulmus rubra 1333 33 8
Quercus velutina 1000 167 32
Sassfras albidum 667 -- --
Acer saccharum 333 167 40
Amelanchier arborea 333 -- 16
Prunus serotina -- 100 --
Totals 15000 800 288

Table 3. Frequency (%), average cover, and importance values
of the ground layer species encountered at the Argyle Hollow
Barrens, McDonough County, and the McKee Greek Barrens, Adams
County, Illinois. (* exotic species)

Species Argyle Hollow Barrens

 Freq. Mean Cover I.V.

Parthenocissus quinquefolia 74 13.76 54.0
Carex pensylvanica 84 10.88 48.5
Rubus flagellaris 42 3.80 19.8
Solidago ulmifolia 30 3.17 15.4
Helianthus divaricatus 26 1.64 10.3
Dichanthelium acuminatum 36 0.33 9.1
Antennaria plantaginifolia 20 0.64 6.2
Vola pedata 20 0.35 5.4
Danthanla spoata 10 0.25 3.0
Toxkoderdron radicans 8 0.38 2.8
Smilacina racemosa 6 0.42 2.5
Rosa carolina 8 0.19 2.3
Potentilla simplex 4 0.36 2.0
Aster turbinellus 6 0.18 1.9
Anemonella thalictroides 8 0.04 1.9
Carex hirsutella 6 0.08 1.6
Ga1lium concinnum 6 0.08 1.6
Hieracium gronovii 6 0.03 1.5
Elymus virginicus 4 0.07 1.2
Galium cricaezans 4 0.07 1.2
Carex muhlenbergii 4 0.02 1.1
Acalypha virginica 4 0.02 1.1
* Poa compressa 4 0.02 1.1
Muhlenberegia sobolifera -- -- --
Desmodium glutinosum -- -- --
Aster anomalus -- -- --
Phlox divaricata -- -- --
Bromus pubescens -- -- --
Festuca subverticillata -- -- --
Sanicula canadensis -- -- --
Amphicarpaea bracteata -- -- --
Ageratina altissima -- -- --
Tradescantia ohiensis -- -- --
Lactuca canadensis -- -- --
Geum canadense -- -- --
Lespedeza intermedia -- -- --
Others -- 0.20 4.5
Totals 36.98 200.0
Bare ground and litter 62.14

Species McKee Creek Barrens

 Freq. Mean Cover I.V.

Parthenocissus quinquefolia 28 2.04 12.0
Carex pensylvanica 62 3.00 21.5
Rubus flagellaris 4 0.60 2.8
Solidago ulmifolia 82 5.93 34.9
Helianthus divaricatus 22 4.54 19
Dichanthelium acuminatum 4 0.02 0.9
Antennaria plantaginifolia -- --
Vola pedata -- --
Danthanla spoata 6 0.08 1.4
Toxkoderdron radicans 2 0.06 0.6
Smilacina racemosa 2 0.06 0.6
Rosa carolina 16 0.48 4.6
Potentilla simplex 4 0.12 1.2
Aster turbinellus -- --
Anemonella thalictroides 10 0.10 2.2
Carex hirsutella 12 0.16 2.8
Ga1lium concinnum 20 1.09 7.3
Hieracium gronovii -- --
Elymus virginicus 8 0.04 1.6
Galium cricaezans 10 0.30 2.8
Carex muhlenbergii 16 0.13 3.4
Acalypha virginica 16 0.08 3.3
* Poa compressa -- --
Muhlenberegia sobolifera 32 5.13 22.8
Desmodium glutinosum 20 1.52 8.7
Aster anomalus 24 1.21 8.4
Phlox divaricata 22 1.10 7.8
Bromus pubescens 14 0.42 4
Festuca subverticillata 16 0.18 3.6
Sanicula canadensis 16 0.18 3.6
Amphicarpaea bracteata 10 0.49 3.5
Ageratina altissima 10 0.49 3.5
Tradescantia ohiensis 12 0.31 3.3
Lactuca canadensis 4 0.31 1.8
Geum canadense 6 0.13 1.5
Lespedeza intermedia 4 0.07 1
Others -- 0.28 3.6
Totals 30.61 200.0
Bare ground and litter 67.09
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Author:McClain, William E.; Ebinger, John E.
Publication:Transactions of the Illinois State Academy of Science
Date:Jul 1, 2007
Words:4863
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