Printer Friendly

Diversity of Albanian Cladoniaceae/Diversidad de las Cladoniaceae de Albania.

Introduction

Albania is a Mediterranean country located in SE Europe and surrounded by the Adriatic Sea and Montenegro in the NW, by the Ionian Sea and Greece in the SW and North Macedonia in the East. Despite its small extension, 28,748 [km.sup.2], it is topographically very diverse. Although Albania lies within the temperate zone the topographic heterogeneity and the combination of Mediterranean and continental influences confer a great climatic and vegetation diversity (Reed et al. 2004).

The main mountain ranges in the country, the Dinaric Alps, run parallel to the coast and they present the highest peak at 2753 m (Mt. Korab). The coastal region has a Mediterranean climate, and further east the mountains act as a barrier to the penetration of temperate air masses. The climate of the mountainous regions is alpine, with strong changes in precipitation and temperature depending on height and exposure. The temperature differences between the coast and the interior regions are more marked in winter, while the precipitations are more intense in summer (Tatli & Turkes 2014).

The karstic soils (limestones) are predominant, although there are locations with serpentine substrates containing a certain amount of silicates, yeilding slightly acid soils (Bego et al. 2018).

Most of the Albanian territory is included in the Illyrian province of the Circumboreal Region with predominant deciduous forest of Quercus ssp. (Q. pubescens, Q. frainetto and Q. cerris) and with an important presence of Carpinus orientalis (Polunin & Walters 1989). In the higher altitudinal zones the grasslands and beech forests are dominant. The Mediterranean vegetation belongs to the Adriatic province and is usually well developed. It occupies a small coastal strip and has sub-Mediterranean and Mediterranean species of evergreen and maquis shrubs (Sadori et al. 2015).

The lichenological studies in Albania are quite scarce as confirmed by Hafellner & Kashta (2003) and Hafellner (2007). Later Svoboda et al. (2012) increased the number of species found up to 333 taxa of lichenized fungi and 11 of them represent Cladonia. The aim of this contribution is to summarize results from new collections in the framework of a project on the family Cladoniaceae in the Mediterranean area of Eurasia.

Materials and Methods

After several recollections in the Albanian territory at different periods during 2009-2017, we studied and analysed 335 Cladonia specimens, which are listed in Annex 1. The new collections were deposited in Madrid (MACB), Helsinki (H) or Budapest (BP). Most of the specimens were analysed by means of thin-layer chromatography (TLC, HPTLC) to identify the major lichen substances according to standardized methods (Orange et al. 2001, 2010; Arup et al. 1993). The main secondary metabolites of Albanian Cladonia are listed in Table 1. Comments on world distribution of species are from Ahti & Stenroos (2013). The distribution maps are presented according to the administrative divisions in counties of Albania (Figure 1). The ITS rDNA region was sequenced in some specimens to confirm the identifications and the GeneBank numbers are provided (Annex 1). The primers and protocols used are described in Pino-Bodas et al. (2015).

Results and discussion

Cladonia is the only genus of Cladoniaceae represented in Albania. Of the 27 analysed species the following 15 are newly reported to the former lichen checklist of Albania (*): Cladonia borealis, C. caespiticia, C. cervicornis, C. coccifera, C. conista, C. cryptochlorophaea, C. cyathomorpha, C. humilis, C. merochlorophaea, C. parasitica, C. peziziformis, C. polycarpoides, C. ramulosa, C. rei and C. subulata. The species studied occur between the sea level and 1800 m of altitude based on our records and the information published by other authors (Figure 2).

The commonest species are: C. cariosa, C. fimbriata, C. foliacea, C. furcata, C. pyxidata and C. rangiformis, while C. caespiticia, C. coccifera, C. digitata, C. macilenta, C. peziziformis, C. polycarpoides and C. ramulosa are restricted to northern Albania, and C. borealis, C. cryptochlorophaea, C. humilis, C. merochlorophaea and C. parasitica have a southern or a south-eastern distribution (Figure 3).

Nine species are chemically variable (Table 1) C. cariosa, C. foliacea, C. furcata, C. polycarpoides, C. pyxidata, C. rangiformis, C. rei and C. symphycarpa. Cladonia cariosa specimens contain three chemotypes: atranorin; atranorin and norstictic acid; atranorin and rangiformic acid, and additional zeorin. Most of the specimens of C. foliacea contain fumarprotocetraric and usnic acids, additionally physodalic acid; but a few contain fumarprotocetraric, psoromic and usnic acids. In C. furcata most of the specimens contain fumarprotocetraric acid, occasionally with physodalic acid; specimens with atranorin and fumarprotocetraric acid or atranorin and bourgeanic acid were also found. Two chemotypes were detected in the specimens of C. polycarpoides atranorin and norstictic acid, and one specimen containing only norstictic acid. Cladonia pyxidata contains fumarprotocetraric acid and additionally physodalic acid, and also some specimens have fumarprotocetraric and psoromic acids. The commonest chemotype in C. rangiformis contains atranorin, fumarprotocetraric and rangiformic acids sometimes with additional physodalic acid, the other chemotype with atranorin and rangiformic acid is very rare in the Albanian specimens. The specimens of C. rei contain fumarprotocetraric and homosekikaic acids and other with only homosekikaic acid. Specimens of C. symphycarpa have atranorin and norstictic acid; and other also atranorin, norstictic and fumarprotocetraric acids.

List of species

*1. Cladonia borealis S. Stenroos, Ann. Bot. Fenn. 26: 160 (1989)

This terricolous species was only found in Korce County growing on sandstone in slopes of beech forest at 1550 m altitude. It is distributed in arctic/antarctic-temperate and circumboreal regions (Ahti & Stenroos 2013).

*2. Cladonia caespiticia (Pers.) Florke, Clad. Comm.: 8 (1828)

It is a very rare species only found in Lezhe County on limestone in slopes of Carpinus orientalis formations at 40 m altitude. This is mainly a temperate species with oceanic tendencies, not continuously circumpolar (Ahti & Stenroos 2013).

3. Cladonia cariosa (Ach.) Spreng., Syst. Veg., Edn 16, 4(1): 272 (1827)

A very common species found in Elbasan, Korce, Kukes, Shkoder and Tirane counties. It was reported by Svoboda et al. (2012) from Diber County on calcareous soil. This is a terricolous species growing on bare soils of limestone, sandstone, serpentine and slate substrates in a wide variability of formations at 120-1550 m altitude. It is a Holarctic species with an outlier in southern South America (Ahti & Stenroos 2013). It is very common and frequent with several chemotypes.

4. Cladonia carneola (Fr.) Fr., Lichenogr. Eur. Reform.: 233 (1831)

This species has only been reported from Shkoder County by Zahlbruckner (1889: 7) (Hafellner 2007). Species with bipolar distribution (Ahti & Stenroos 2013).

*5. Cladonia cervicornis (Ach.) Flot., Jahresber., Schles. Ges. Vaterl. Cult. 27: 105 (1849)

This species is not frequent, only encountered in Durres, Shkoder and Vlore counties. It grows from sea level to 865 m altitude on limestone and sandy and acid soils. It has a Holarctic distribution with uncertain records in South America (Ahti & Stenroos 2013).

6. Cladonia chlorophaea (Florke ex Sommerf.) Spreng., Syst. Veg., Edn 16, 4(1): 273 (1827)

A rare species collected in Gjirokaster and Vlore counties. But it was early reported from Kukes and Shkoder districts (Hafellner & Kastha 2003); from Diber and Tirane districts by Markgraf (1931, 1932) (Hafellner 2007). It is a cosmopolitan species (Ahti & Stenroos 2013).

*7. Cladonia coccifera (L.) Willd., Fl. Berol. Prodr.: 361 (1787)

This is a very rare species only collected in Shkoder County at 735 m altitude on serpentine substrate. It has a cosmopolitan distribution (Ahti & Stenroos 2013).

8. Cladonia coniocraea (Florke) Spreng., Syst. Veg., Edn 16, 4(1): 272 (1827) syn. C. ochrochlora Florke

It was collected from Durres, Lezhe and Vlore counties from sea level until 828 m altitude, on limestone and sandy substrates or stumps of Castanea sativa. It was reported of Diber and Shkoder districts by Svoboda et al. (2012) and Hafellner & Kashta (2003). All of the Albanian specimens belong to C. coniocraea s.str. the more oceanic morphotype often recognized as C. ochrochlora, which is not distinguishable in DNA analyses (Pino-Bodas et al. 2011) is not present here.

*9. Cladonia conista (Nyl.) Robbins, in Allen, Rhodora 32: 92 (1930)

A common species collected in Durres, Elbasan, Fier, Gjirokaster, Korce, Lezhe, Shkoder, Tirana and Vlore counties. It grows on slopes and bare soils of Carpinus orientalis formations and also Quercus coccifera, on sandstone and limestones substrates, from 40-1000 m altitude. Traditionally considered a chemotype of C. humilis with fumarprotocetraric and bourgeanic acids nevertheless, should be considered a different species in base of the results of Pino-Bodas et al. (2012).

*10. Cladonia cryptochlorophaea Asahina, J. Jap. Bot. 16: 711 (1940)

This is a very rare species only collected in Vlore County at 186 m altitude growing on limestones with Erica arborea and deciduous Quercus. Is a terricolous species with arctic/antarctic to northern temperate distribution (Ahti & Stenroos 2013).

*11. Cladonia cyathomorpha Stirt. ex Walt. Watson, J. Bot. 73: 156 (1935)

It was collected from Elbasan, Kukes, Shkoder and Tirane counties growing at 120-540 m altitude on limestone and serpentine substrates with Carpinus orientalis, deciduous Quercus and Arbutus unedo. It has a western European distribution with some outpost in southern South America (Ahti & Stenroos 2013).

12. Cladonia digitata (L.) Hoffm., Deutschl. Fl., Zweiter Theil (Erlangen): 124 (1796)

This species was only reported by Svoboda et al. (2012) from Diber district growing on rotten wood at 1700-1730 m altitude. It is mainly a Holarctic species with some outposts in East Africa (Ahti & Stenroos 2013).

13. Cladonia fimbriata (L.) Fr., Lich. Eur. Reform. (Lund): 222 (1831)

syn. C. major (K. G. Hagen) Sandst., C. minor (K. G. Hagen) Szat.

A very common species encountered in Diber, Durres, Elbasan, Gjirokaster, Korce, Kukes, Lezhe, Shkoder, Tirane and Vlore counties. It grows from sea level to 1750 m altitude on rotten wood, tree bases and humose soil over limestone, sandstone and sandy soil. The species was reported by Hafellner & Kashta (2003) and Svoboda et al. (2012) from Diber, Has, Kukes and Shkoder districts. It is a cosmopolitan species (Ahti & Stenroos 2013).

14. Cladonia foliacea (Huds.) Willd., Fl. Berol. Prodr.: 363 (1787)

incl. C. convoluta (Lam.) Anders

This is a very common species found in Berat, Diber, Durres, Elbasan, Gjirokaster, Korce, Kukes, Lezhe, Shkoder, Tirane and Vlore counties from 78 to 1550 m altitude. It is a terricolous species growing on grasslands and open soil, preferably calcareous. It is a Western Eurasian, Macaronesian and North African species which extends in Asia as far as Mongolia and it grows in temperate to subtropical areas (Ahti & Stenroos 2013).

The specimens containing fumarprotocetraric and usnic acids, additionally physodalic acid are the commonest, but samples with fumarprotocetraric, usnic and psoromic acids were found in Korce, Gjirokaster and Vlore counties on sandstone at 211-953 m altitude.

Most of the early reports (Markgraf 1927, 1931, 1932; Hafellner & Kashta 2003; Svoboda et al. 2012) were published as C. convoluta but we include it in the concept of C. foliacea because there are no molecular differences between these taxa (Pino-Bodas et al. 2010a, 2018), although specimens called C. convoluta are of larger size due to its adaptation to calcareous soil and dry situations.

15. Cladonia furcata (Huds.) Schrad., Spicil. Fl. Germ. 1: 107 (1794)

incl. C. subrangiformis Sandst.

This common species was collected in Durres, Elbasan, Gjirokaster, Korce and Vlore counties from sea level to 1700 m altitude. Most of the samples contain fumarprotocetraric acid and occasionally physodalic acid. The morphotype with atranorin and fumarprotocetraric acid or atranorin and bourgeanic acid, traditionally identified as C. subrangiformis, was collected from Korce, Shkoder and Vlore counties. This taxon is considered to fall into the chemical variability of C. furcata supported by the DNA studies of Pino-Bodas et al. (2015).

C. furcata was reported from Diber, Permet, Shkoder and Tirane districts by Markgraf (1931, 1932), Hafellner & Kashta (2003) and Svoboda et al. (2012) without chemical indications. It grows on humose, shaded or open soil over limestone, serpentine, and conglomerate substrates in a wide varirty of forests. It has a subcosmopolitan distribution in boreal to tropical areas (Ahti & Stenroos 2013).

*16. Cladonia humilis (With.) J.R. Laundon, Lichenologist 16: 220 (1984)

A rare species only collected in Elbasan and Vlore counties from 50 to 540 m altitude. It grows on limestone substrate with Carpinus orientalis, Quercus spp. and Buxus sempervirens. The samples analysed contain atranorin and fumarprotocetraric acid. It has a Holarctic distribution, with several outposts in the Southern Hemisphere, occurring from boreal to arctic areas (Ahti & Stenroos 2013), although its distribution is not well known.

17. Cladonia macilenta Hoffm., Deutschl. Fl., Zweiter Theil (Erlangen): 126 (1796)

This is a rare species only collected in Lezhe County growing on bark of Pinus nigra. It was reported from Vlore district (Hafellner & Kashta 2003). This species has a subcosmopolitan distribution (Ahti & Stenroos 2013).

*18. Cladonia merochlorophaea Asahina, J. Jap. Bot. 6: 713 (1940)

A very rare species only collected in Korce County, on calcareous soil, at 1625 m altitude. This is a subcosmopolitan species but its taxonomic status is not fully clear (Ahti & Stenroos 2013).

*19. Cladonia parasitica (Hoffm.) Hoffm., Deutschl. Fl. 2: 127 (1796)

This is a very rare species only collected in Vlore County at 828 m altitude growing on Castanea sativa stumps. This is a Holarctic species with continental tendencies (Ahti & Stenroos 2013).

*20. Cladonia peziziformis (With.) J.R. Laundon, Lichenologist 16: 223 (1984)

A very rare species only collected in Shkoder County, on serpentine soil at 120 m altitude. This is a subcosmopolitan species easily overlooked due to its small size (Ahti & Stenroos 2013).

*21. Cladonia polycarpoides Nyl. in Zwackh, Lich. Exs. Nos. 626, 626bis (1892)

syn. C. subcariosa auct.

It is a rare species collected in Kukes and Shkoder counties growing on serpentine soil from sea level to 600 m altitude. It is a subcosmopolitan species especially abundant under suboceanic to oceanic conditions (Ahti & Stenroos 2013).

22. Cladonia pyxidata (L.) Hoffm., Deutschl. Fl., Zweiter Theil (Erlangen): 121 (1796)

incl. C. pocillum (Ach.) Grognot; C. monomorpha Aptroot, Sipman & van Herk

A common species collected in Diber, Durres, Elbasan, Fier, Gjirokaster, Korce, Shkoder, Tirane and Vlore counties growing on bare soils, grasslands and rarely on wood. Most of the samples analysed contain fumarprotocetraric acid and often additional physodalic acid. A chemotype with fumarprotocetraric and psoromic acids was collected at 540 m altitude from Elbasan County. Cladonia pyxidata was early reported from Kukes, Lezhe, Shkoder and Vlore districts (Zahlbruckner 1889, Markgraf 1931, Szatala & Timko 1926, Murati 1979, Hafellner & Kashta 2003, Svoboda et al. 2012). Some of these reports refer to C. pocillum but we do not distinguish it as a different entity because molecular studies do not support it (Kotelko & Piercey-Normore 2010). It has a cosmopolitan distribution (Ahti & Stenroos 2013).

*23. Cladonia ramulosa (With.) J.R. Laundon, Lichenologist 16: 225 (1984)

It was collected in Durres and Lezhe counties on sandy soil and limestone from sea level to 40 m altitude. This is a subcosmopolitan species (Ahti & Stenroos 2013).

24. Cladonia rangiformis Hoffm., Deutschl. Fl., Zweiter Theil (Erlangen): 114 (1796)

A very common species collected in Berat, Diber, Durres, Elbasan, Fier, Gjirokaster, Korce, Kukes, Lezhe, Shkoder, Tirane and Vlore counties. It grows on grasslands and open soil of sandy substrate, serpentines, limestones and slates from sea level to 1770 m altitude. Its distribution is mainly restricted to western Eurasia and North Africa with one outpost in eastern Canada (Ahti & Stenroos 2013). The commonest chemotype contains atranorin, fumarprotocetraric and rangiformic acids sometimes with additional physodalic acid, but the chemotype with atranorin and rangiformic acid (common elsewhere) is very rare. One specimen of a sorediate morph ('var. sorediophora') was found in Shkoder.

It was reported from Diber, Gjirokaster, Kukes, Shkoder, Tirane and Vlore counties by Szatala & Timko (1926), Markgraf (1927, 1932) and Svoboda et al. (2012), but without indication of their chemistry.

*25. Cladonia rei Schaer. Lich. Helv. Spicil. 1(1): 34 (1823)

It is an uncommon species collected in Durres, Kukes and Vlore counties. It grows on limestone and sandstone substrates at 90-330 m altitude on limestone and sandstones substrates. It has a Holarctic distribution with continental preferences (Ahti & Stenroos 2013). One sample analysed contains homosekikaic acid only, but other specimens contain homosekikaic and fumarprotocetraric acids. Some time ago C. rei was considered a homosekikaic chemotype of C. subulata (Spier & Aptroot 2007) but molecular studies maintained them as different species (Pino-Bodas et al. 2010b).

*26. Cladonia subulata (L.) F.H. Wigg., Prim. Fl. Holsat.: 90 (1780)

This is a rare species only collected in Kukes and Lezhe counties on limestone susbstrate at 40-330 m altitude. It is a subcosmopolitan species (Ahti & Stenroos 2013).

27. Cladonia symphycarpa (Florke) Fr., Sched. Crit. Lich. Suec. 8-9. 20 (1826)

incl. C. dahliana Kristinsson

A very common species collected in Elbasan, Korce and Shkoder counties growing on limestone substrate at 400 to 1800 m altitude. It has been reported only from Shkoder County by Svoboda et al. (2012). The most frequent chemotype has atranorin and norstictic acid, additionally fatty acid; the other chemotype with atranorin, norstictic and fumarprotocetraric acids is very rare. It has a Holarctic distribution with some outliers in South America (Ahti & Stenroos 2013).

Cladonia sp.

Some specimens collected in Shkoder and Vlore were difficult to identify when the podetia are not well developed and do not bear split proliferations in the margin of the scyphi. These specimens belong to the Cladonia humilis group (Pino-Bodas et al. 2013) but molecular studies are necessary to clarify the identity of these specimens.

Acknowledgements

A. R. Burgaz received financial support from the project CGL2013-41839-P, Ministry of Economy and Competitiveness, Spain. Chromatographic studies (HPTLC) carried in Hungary were supported by the Hungarian National Research Development and Innovation Fund [NKFI K 124341].

References

Ahti, T. & Stenroos, S. 2013. Cladoniaceae. In: T. Ahti, S. Stenroos & R. Moberg (Eds.) Nordic Lichen Flora 5: 1-117. Uppsala: Museum of Evolution, Uppsala University.

Arup, U., Ekman, S., Lindblom, L., Mattsson, J.-E. 1993. High performance thin layer chromatography (HPTLC), an improved technique for screening lichen substances. Lichenologist 25(1):61-71.

Bego, F., Krystukek, B., Paspali, G. & Rogozi, E. 2018. Small terrestrial mammals of Albania: distribution and diversity (Mammalia, Eulipotyphla, Rodentia). ZooKeys 742: 127-163.

Hafellner, J. 2007. Checklist and bibliography of lichenized and lichenicolous fungi so far reported from Albania (version 05-2007). Fritschiana 59: 1-18.

Hafellner, J. & Kashta, L. 2003. Miscellaneous records of lichens and lichenicolous fungi from Albania. Herzogia 16: 135-142.

Kotelko, R. & Piercey-Normore, M.D. 2010. Cladonia pyxidata and C. pocillum; genetic evidence to regard them as conspecific. Mycologia 102: 534-545.

Markgraf, F. 1927. An den Grenzen des Mittelmeergebietes. Pflanzengeographie von Mittelalbanien. Repertorium Specierum Novarum Regni Vegetabilis, Beiheft 45: 1-217.

Markgraf, F. 1931. Pflanzen aus Albanien 1928. Denkschriften der Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse 102: 317-360.

Markgraf, F. 1932. Pflanzengeographie von Albanien. Ihre Bedeutung fur Vegetation und Flora der Mittelmeerlander. Bibliotheca Botanica 26(105): 1-132.

Murati, M. 1979. Kontribut rreth njohjes se likenoflores te Butrintit. Buletini i Shkencave te Natyres, Tirane 3: 97-99.

Orange, A., James, P.W. & White, F.J. 2001. Microchemical methods for the identification of lichens. British Lichen Society, London.

Orange, A., James, P.W. & White, F.J. 2010. Microchemical methods for the identification of lichens, second ed. British Lichen Society, London.

Pino-Bodas, R., Martin, M.P. & Burgaz, A.R. 2010a. Insight into the Cladonia convoluta-C. foliacea (Ascomycotina) complex and related species revealed through morphological, biochemical and phylogenetic analyses. Syst. Biodivers. 8(4): 575-586.

Pino-Bodas, R., Martin, M.P. & Burgaz, A.R A. R. 2010b. Elucidating the taxonomic rank of the lichens Cladonia subulata and C. rei (Cladoniaceae). Mycotaxon 113: 311-326.

Pino-Bodas, R., Burgaz, A.R., Martin, M.P. & Lumbsch, H.T. 2011. Phenotypical plasticity and homoplasy complicate species delimitation in the Cladonia gracilis group (Cladoniaceae). Org. Divers. Evol. 11: 343-355.

Pino-Bodas, R., Ahti, T., Stenroos, S., Martin, M.P. & Burgaz, A.R. 2012. Cladonia conista and C. humilis (Cladoniaceae) are different species. Biblioth. Lichenol. 108: 161-176.

Pino-Bodas, R., Ahti, T., Stenroos, S., Martin, M. P. & Burgaz, A.R. 2013. Multilocus approach to species recognition in the Cladonia humilis complex (Cladoniaceae, Ascomycota). Am. J. Bot. 100(4): 664-678.

Pino-Bodas, R., Burgaz, A.R., Martin, M.P., Ahti, T., Stenroos, S., Wedin, M. & Lumbsch, H.T. 2015. The morphological features used for distinguishing species within the Cladonia furcata complex are highly homoplasious. Lichenologist 47(5): 287-303.

Pino-Bodas, R., Burgaz, A. R., Ahti, T. & Stenroos, S. 2018. Taxonomy of Cladonia angustiloba and related species. Lichenologist 50(3): 267-282.

Polunin, O. & Walters, M. 1989. Guia de la vegetacion de Europa. Barcelona: Ediciones Omega.

Reed, J.M., Krystufek, B. & Eastwood, W.J. 2004. The physical geography of the Balkans and nomenclature of place names. In: Griffiths H.I., Krystufek B. and Reed J.M. (eds.), Balkan Biodiversity, Kluwer Academic Publishers, Dordrecht: 9-22.

Sadori, L., Giardini, M., Gliozzi, E., Mazzini, I., Sulpizio, R., Van-Welden, A. & Zanchetta, G. 2015. Vegetation, climate and environmental history of the last 4500 years at lake Shkodra (Albania/Montenegro). The Holocene 25(3): 435-444.

Spier, L. & Aptroot, A. 2007. Cladonia rei is a chemotype and synonym of Cladonia subulata. Lichenologist 39: 57-60.

Szatala, O. & Timko, Gy. 1926. Adatok Albania florajahoz. Additamenta ad floram Albaniae. IV. Lichenes. Magyar Tudomanyos Akademia Balkan-Kutatasainak Tudomanyos Eredmenyei 3: 159-179.

Svoboda, D., Bouda, F., Malicek, J. & Hafellner, J. 2012. A contribution to the knowledge of lichenized and lichenicolous fungi in Albania. Herzogia 25: 149-165.

Tatli, H. & Turkes, M. 2014. Climatological evaluation of Haines forest fire weather index over the Mediterranean Basin. Meteorol. Appl. 21: 545-552.

Zahlbruckner, A. 1889. Lichenes. In: G. Beck & I. Szyszylowiczi: Plantas a Dr. Ign. Szyszylowicz in itinere per Cernagoram et in Albania adiacenti anno 1886 lectas. Rozprawy i Sprawozdania z Posiedzen Wydzialu Matematyczno-Przyrodniczego Akademii Umiejetnosci 19: 6-14.

Annex 1. Selected specimens examined

Cladonia borealis S. Stenroos

Barbatic acid: KORCE: District of Korce, Dardhe, SH90 Boboshtice-Dardhe, 40[grados]31'26"N 20[grados]48'06"E, Fagus sylvatica, Juniperus communis, Abies sp., sandstone, 1550 m, 23-IV-2017, A.R. Burgaz (MACB 111457).

Cladonia caespiticia (Pers.) Florke

Fumarprotocetraric acid: LEZHE: District of Kurbin, Milot, left side Mat river, SH25 Milot-Zogu Bridge, 41[grados]41'54"N 19[grados]45'10"E, Carpinus orientalis, Erica arborea, limestone, 40 m, 21-IV-2017, A.R. Burgaz (MACB 111430, GenBank accession no: MK543953).

Cladonia cariosa (Ach.) Spreng.

Atranorin chemotype: KUKES: District of Kukes, Kukes, Rruga Tirana, 42[grados]04'46"N 20[grados]24'45"E, Quercus cerris, deciduous Quercus, limestone, 330 m, 21-IV-2017, A.R. Burgaz (MACB 111436). SHKODER: Vau i Dejes, SH25 Vau i Dejes-Koman, 42[grados]00'55"N 19[grados]39'37"E, Pinus pinaster, Carpinus orientalis, serpentine soil, 120 m, 20-IV-2017, A.R. Burgaz (MACB 111420). Vau i Dejes, Karme, SH25 Vau i Dejes-Koman, 42[grados]04'02"N 19[grados]40'59"E, Quercus macrolepis, Q. pubescens, Juniperus oxycedrus, Erica arborea, slates, 130 m, 20-IV-2017, A.R. Burgaz (MACB 111429). District of Malesi e Madhe, Boge, SH21 Koplik-Boge, 42[grados]21'46"N 19[grados]37'56"E, Pinus nigra, Juniperus oxycedrus, limestone, 865 m, 20-IV-2017, AR. Burgaz (MACB 111298). Atranorin and norstictic acid chemotype: ELBASAN: Prrenjas, Urake, SH9 Qafe Thane, 41[grados]04'24"N 20[grados]36'18"E, Quercus macrolepis, deciduous Quercus, Juniperus oxycedrus, limestone, 1000 m, 22-IV-2017, A.R. Burgaz (MACB 111446). District of Elbasan, Bradashesh, SH3 Bradashesh-Graze, 41[grados]06'33"N 19[grados]59'49"E, Carpinus betulus Juniperus oxycedrus, limestone karst, 530 m, 15-IV-2017, A.R. Burgaz (MACB 111402). KORCE: District of Korce, Dardhe, SH90 Boboshtice-Dardhe, 40[grados]31'18"N 20[grados]49'22"E, Fagus sylvatica, Pinus nigra, Juniperus communis, Betula pendula, Abies sp., limestone, 1390 m, 23-IV-2017, A.R. Burgaz (MACB 111459). District of Korce, Dardhe, SH90 Boboshtice-Dardhe, 40[grados]31'26"N 20[grados]48'06"E, Fagus sylvatica, Juniperus communis, Abies sp., sandstone, 1550 m, 23-IV-2017, A.R. Burgaz (MACB 111455). District of Korce, Boboshtice, SH90 Boboshtice-Dardhe, 40[grados]32'37"N 20[grados]47'11"E, Juniperus oxycedrus, pastureland, sandstone, 1200 m, 23-IV-2017, A.R. Burgaz (MACB 111453). District of Pogradec, Udenisht, SH3 Piskutat-Pogradec, shore Ohrid lake, 40[grados]58'55"N 20[grados]38'21"E, Pinus halepensis, Juniperus oxycedrus, Buxus sempervirens, sandstone, 710 m, 22-IV-2017, A.R. Burgaz (MACB 111452). KUKES: District of Kukes, Kolsh, SH5 close Drin river, 42[grados]04'18"N 20[grados]20'32"E, deciduous Quercus, Juniperus oxycedrus, Buxus sempervirens, serpentine soil, 510 m, 21-IV-2017, A.R. Burgaz (MACB 111433). District of Kukes, Rruga Tirana, 42[grados]04'46"N 20[grados]24'45"E, Quercus cerris, deciduous Quercus, limestone, 330 m, 21-IV-2017, A.R. Burgaz (MACB 111435, 111434). Atranorin and rangiformic acid chemotype: ELBASAN: Prrenjas, Urake, SH9 Qafe Thane, 41[grados]04'24"N 20[grados]36'18"E, Quercus macrolepis, deciduous Quercus, Juniperus oxycedrus, limestone, 1000 m, 22-IV-2017, A.R. Burgaz (MACB 111447). KORCE: District of Kolonje (Rrethi i Kolonjes), c. 1.9 km W of village Pepellash, 40[grados]28'56.18"N 20[grados]40'21.27"E, on acid soil in moist road slope, 549 m, 18-X-2011, Z. Barina, L. Lokos, D. Pifko (BP 96585). District of Korce, Dardhe, SH90 Boboshtice-Dardhe, 40[grados]31'18"N 20[grados]49'22"E, Fagus sylvatica, Pinus nigra, Juniperus communis, Betula pendula, Abies sp., limestone, 1390 m, 23-IV-2017, A.R. Burgaz (MACB 111460). District of Korce, Dardhe, SH90 Boboshtice-Dardhe, 40[grados]31'26"N 20[grados]48'06"E, Fagus sylvatica, Juniperus communis, Abies sp., sandstone, 155 m, 23-IV-2017, A.R. Burgaz (MACB 111456). District of Korce (Rrethi i Korces), on the ridge of Mts Mali i Moraves above village Dardhe, near peak Buzeja e Boboshtices (1701 m), 40[grados]32'23.97"N 20[grados]49'14.20"E, on acid soil in beech forest on gravel conglomerate, 1550 m, 10-VIII-2010, L. Lokos (BP 96563). KUKES: District of Kukes, Kolsh, SH5 close Drin river, 42[grados]04'18"N 20[grados]20'32"E, deciduous Quercus, Juniperus oxycedrus, Buxus sempervirens, serpentine soil, 510 m, 21-IV-2017, A.R. Burgaz (MACB 111462). SHKODER: Vau i Dejes, SH25 Vau i Dejes-Koman, 42[grados]03'01"N 19[grados]39'45"E, Pinus pinaster, Juniperus oxycedrus, Quercus cerris, Erica arborea, Cotinus coggygria, serpentine soil, 130 m, 20-IV-2017, A.R. Burgaz (MACB 111426). TIRANE: District of Tirane, Mushqeta, SH3 Mushqeta-Gracen, 41[grados]12'38"N 19[grados]57'09"E, Arbutus unedo, Erica arborea, Platanus orientalis, boulders, 480 m, 22-IV-2017, A.R. Burgaz (MACB 111440). Atranorin, rangiformic acid and zeorin chemotype: SHKODER: Vau i Dejes, SH25 Vau i Dejes-Koman, 42[grados]03'01"N 19[grados]39'45"E, Pinus pinaster, Juniperus oxycedrus, Quercus cerris, Erica arborea, Cotinus coggygria, serpentine soil, 130 m, 20-IV-2017, A.R. Burgaz (MACB 111425). Vau i Dejes, Karme, SH25 Vau i Dejes-Koman, 42[grados]04'02"N 19[grados]40'59"E, Quercus macrolepis, Q. pubescens, Juniperus oxycedrus, Erica arborea, slates, 130 m, 20-IV-2017, A.R. Burgaz (MACB 111428).

Cladonia cervicornis (Ach.) Flot.

Fumarprotocetraric acid: DURRES: District of Durres, Fushe-Draci, San Pietro beach track, 41[grados]31'01"N 19[grados]31'12"E, Pinus pinea, sandy soil, 1 m, 19-IV-2017, A.R. Burgaz (MACB 111225). SHKODER: District of Malesi e Madhe, Poica, SH21 Koplik-Boge, 42[grados]15'53"N 19[grados]29'47"E, Juniperus oxycedrus, Cupressus sempervirens, boulders, 320 m, 20-IV-2017, A.R. Burgaz (MACB 111292). District of Malesi e Madhe, Dedaj, SH21 Koplik-Boge, 42[grados]18'04"N 19[grados]32'32"E, Carpinus betulus, Juniperus oxycedrus, limestone rocks, 495 m, 20-IV-2017, A.R. Burgaz (MACB 111299). District of Malesi e Madhe, Boge, SH21 Koplik-Boge, 42[grados]21'46"N 19[grados]37'56"E, Pinus nigra, Juniperus oxycedrus, limestone, 865 m, 20-IV-2017, A.R. Burgaz (MACB 111303). Puke District (Rrethi i Pukes), Mount Rras' e Qerretit above village Qerret, 42[grados]03'36.98"N 19[grados]49'53.45"E, in serpentine rocky area, on acid soil in pine woodland, 735 m, 6-V-2015, L. Lokos (BP 96620). District of Shkoder (Rrethi i Shkodres), Qafa e Celes above village Mirdice, 42[grados]0'11.98"N 19[grados]43'33.16"E, on soil in roadside dry rocky grassland, 470 m, 12-X-2011, L. Lokos (BP 96756). VLORE: Finiq, Muzine, Muzine, 39[grados]54'48"N 20[grados]11'20"E, deciduous Quercus, Erica arborea, limestone, 186 m, 17-IV-2017, A.R. Burgaz (MACB 111199).

Cladonia chlorophaea (Florke ex Sommerf.) Spreng.

Fumarprotocetraric and physodalic acids: GJIROKASTER: District of Tepelene, Kelcyre, Kelcyre gorge, left side Vjosa river, 40[grados]17'52>>N 20[grados]10'05>>E, Carpinus betulus, Pinus halepensis, sandstone, 180 m, 18-IV-2017, A.R. Burgaz, (MACB 111411). LEZHE: District of Mirdite (Rrethi i Mirdites), in the valley of stream Fani i Vogel above village Thirre, in serpentine rocky area, on bark (Pinus nigra), 42[grados]0'20.98"N 20[grados]14'33.25"E, 905 m, 6-V-2015, L. Lokos (BP 96627). VLORE: Finiq, Bistrice, Muzine, 39[grados]55'08"N 20[grados]08'48"E, Quercus coccifera, Carpinus betulus, Pinus halepensis, Cercis siliquastrum, limestone slopes, 125 m, 17-IV-2017, A.R. Burgaz (MACB 111193).

Cladonia coccifera (L.) Willd.

Usnic acid and zeorin: SHKODER: Puke District (Rrethi i Pukes), Mount Rras' e Qerretit above village Qerret, 42[grados]03'36.98"N 19[grados]49'53.45"E, in serpentine rocky area, on acid soil in pine woodland, 735 m, 6-V-2015, L. Lokos (BP 96621).

Cladonia coniocraea (Florke) Spreng.

Fumarprotocetraric acid: DURRES: District of Durres, Fushe-Draci, San Pietro beach track, 41[grados]31'01"N 19[grados]31'12"E, Pinus pinea, sandy soil, 1 m, 19-IV-2017, A.R. Burgaz (MACB 111224). LEZHE: District of Kurbin, Milot, left side Mat river, SH25 Milot-Zogu Bridge, 41[grados]41'54"N 19[grados]45'10"E, Carpinus orientalis, Erica arborea, limestone, 40 m, 21-IV-2017, A.R. Burgaz (MACB 111305). VLORE: District of Delvine (Rrethi i Delvines), Mali i Gjere, on the southern slope of 1331 m high mount above Delvine, 39[grados]57'42.12"N 20[grados]07'59.66"E, in chestnut woodland, on bark (Castanea sativa), 828 m, 1-V-2009, Z. Barina & D. Pifko (BP 96542).

Cladonia conista (Nyl.) Robbins

Fumarprotocetraric and bourgeanic acids: DURRES: District of Durres, Fushe-Draci, road cape of Rodon, 41[grados]32'50"N 19[grados]31'08"E, Carpinus betulus, Arbutus unedo, Pistacia lentiscus, Cistus salviifolius, sandstone, 90 m, 19-IV-2017, A.R. Burgaz (MACB 111412). District of Durres (Rrethi i Durresit), west of village Shenpjeter, at the southern foot of hill M. e Multit (159.9 m), near the coast of the Adriatic Sea, 41[grados]32'41.0"N 19[grados]29'16.0"E, on soil in grasslands, 0 m, 26-IV-2009, L. Lokos (BP 96551). ELBASAN: Prrenjas, Urake, SH9 Qafe Thane, 41[grados]04'24"N 20[grados]36'18"E, Quercus macrolepis, deciduous Quercus, Juniperus oxycedrus, limestone, 1000 m, 22-IV-2017, A.R. Burgaz (MACB 111449). District of Librazhd (Rrethi i Librazhdit), Polis Mts, above village Polis, on the ridge between the valley of brook "Sut Kukit" and river Shkumbin, 41[grados]09'16.50"N 20[grados]15'08.79"E, in bushland, on conglomeratic flysh sediments, 555 m, 12-X-2010, L. Lokos (BP 96571). Elbasan county, in open boxwood scrubland above village Kuterman, 41[grados]13'40.99"N 20[grados]14'41.04"E, on acid soil in rocky serpentine area, 510 m, 16-V-2016, L. Lokos (BP 96660). FIER: Fier County, Pylli i Levanit, above village Levan, 40[grados]40'48.99"N 19[grados]30'11.70"E, on calcareous soil in oak woodland, 95 m, 5-V-2017, L. Lokos (BP 96678). GJIROKASTER: Gjirokaster District (Rrethi i Gjirokastres), Mali i Nikollaqit above village Sofratike, 39[grados]59'41.22"N 20[grados]12'40.05"E, on calcareous soil in rocky grasslands, 260 m, 8-V-2015, L. Lokos (BP 96631). District of Gjirokaster (Rrethi i Gjirokastres), Kurvelesh i Siperm uplands above village Picar, 40[grados]11'0.39"N 20[grados]02'31.51"E, on calcareous soil in rocky grasslands, 750 m, 29-III-2015, L. Lokos (BP 96600). District of Permet (Rrethi i Permetit); left part of river Vjose, between town Kelcyre and village Mbrezhdan, 40[grados]19'11.24"N 20[grados]05'43.54"E, in scrubland, on flysh soil, 205 m, 9-V-2015, L. Lokos (BP 96633). District of Tepelene (Rrrethi i Tepelenes), c. 500 m N of village Mirne, 40[grados]21'53.91"N 19[grados]58'15.44"E, in dry grassland, on flysh soil, 160 m, 14-X-2012, L. Lokos (BP 96595). KORCE: Kolonje District (Rrethi i Kolonjes), between town Leskovik and village Vrepcke, 40[grados]10'30.62"N 20[grados]38'22.95"E, on acid soil in rocky serpentine grasslands, 1030 m, 10-V-2015, Z. Barina, L. Lokos & D. Pifko (BP 96637). District of Korce (Rrethi i Korces), above the water reservoir c 2.5 km WSW of village Ujebardhe, 40[grados]31'17.31"N 20[grados]39'50.62"E, on acid soil in dry rocky grassland, 1105 m, 18-X-2011, L. Lokos (BP 96586). LEZHE: Kurbin, Milot, left side Mat river, SH25 Milot-Zogu Bridge, 41[grados]41'54"N 19[grados]45'10"E, Carpinus orientalis, Erica arborea, limestone, 40 m, 21-IV-2017, A.R. Burgaz (MACB 111431). SHKODER: Puke District (Rrethi i Pukes), Mount Rras' e Qerretit above village Qerret, 42[grados]03'36.98"N 19[grados]49'53.45"E, in serpentine rocky area, on bark (Juniperus), 735 m, 6-V-2015, L. Lokos (BP 96622). TIRANE: District of Kavaje (Rrethi i Kavajes), by the coast of the Adriatic Sea between villages Spille and Rreth (Rreth-Greth), 41[grados]05'07.6"N 19[grados]27'31.5"E, on sandy soil in sand dune area, 10 m, 27-IV-2009, L. Lokos (BP 96552). VLORE: Finiq, Bistrice, Muzine, 39[grados]55'08"N 20[grados]08'48"E, Quercus coccifera, Carpinus betulus, Pinus halepensis, Cercis siliquastrum, limestone slopes, 125 m, 17-IV-2017, A.R. Burgaz (MACB 111409). District of Delvine (Rrethi i Delvines), Mali i Gjere, on the southern slope of 1331 m high mount above Delvine, 39[grados]57'42.12"N 20[grados]07'59.66"E, in chestnut woodland, on bark (Castanea sativa), 828 m, 1-V-2009, Z. Barina, D. Pifko (BP 96543). District of Vlore (Rrethi i Vlores), between village Ilias and manastir Shen Theodor, 40[grados]08'07.8"N 19[grados]40'26.4"E, on soil in olive plantation, 211 m, 29-IV-2009, L. Lokos (BP 96557).

Cladonia cryptochlorophaea Asahina

Cryptochlorophaeic acid: VLORE: Finiq, Muzine, Muzine, 39[grados]54'48"N 20[grados]11'20"E, deciduous Quercus, Erica arborea, limestone, 186 m, 17-IV-2017, A.R. Burgaz (MACB 111410).

Cladonia cyathomorpha Stirt. ex Walt. Watson

Fumarprotocetraric acid: ELBASAN: Prrenjas, Prrenjas, SH3 Qukes-Prrenjas, 41[grados]04'24"N 20[grados]30'15"E, Carpinus orientalis, Quercus macrolepis, deciduous Quercus, Buxus sempervirens, limestone slopes, 540 m, 22-IV-2017, A.R. Burgaz (MACB 111443). KUKES: District of Kukes, Kukes, Rruga Tirana, 42[grados]04'46"N 20[grados]24'45"E, Quercus cerris, deciduous Quercus, limestone, 330 m, 21-IV-2017, A.R. Burgaz (MACB 111438). SHKODER: Vau i Dejes, SH25 Vau i Dejes-Koman, 42[grados]00'55"N 19[grados]39'37"E, Pinus pinaster, Carpinus orientalis, serpentine soil, 120 m, 20-IV-2017, A.R. Burgaz (MACB 111421). TIRANE: District of Tirane, Mushqeta, SH3 Mushqeta-Gracen, 41[grados]12'38"N 19[grados]57'09"E, Arbutus unedo, Erica arborea, Platanus orientalis, boulders, 480 m, 22-IV-2017, A.R. Burgaz (MACB 111442).

Cladonia fimbriata (L.) Fr.

Fumarprotocetraric acid: DIBER: District of Mat (Rrethi i Matit), Mali i Skenderbeut, on the northern forehills of Mt. "maja e Mellezit", near village Shkopet, 41[grados]41'38.1"N 19[grados]49'16.4"E, on soil in Ostrya carpinifolia scrubland, 75 m, 24-IV-2009, L. Lokos (BP 96549). DURRES: District of Durres, Fushe-Draci, San Pietro beach track, 41[grados]31'01"N 19[grados]31'12"E, Pinus pinea, sandy soil, 1 m, 19-IV-2017, A.R. Burgaz (MACB 111413). District of Kruje, Kruje, SH38 to Qafe-Shtame N. P., 41[grados]31'03"N 19[grados]49'27"E, Carpinus betulus, pasturelands, limestone, 720 m, 19-IV-2017, A.R. Burgaz (MACB 111414). ELBASAN: Prrenjas, SH3 Qukes-Prrenjas, 41[grados]04'24"N 20[grados]30'15"E, Carpinus orientalis, Quercus macrolepis, deciduous Quercus, Buxus sempervirens, limestone slopes, 540 m, 22-IV-2017, A.R. Burgaz (MACB 111445). Prrenjas, Urake, SH9 Qafe Thane, 41[grados]04'24"N 20[grados]36'18"E, Quercus macrolepis, deciduous Quercus, Juniperus oxycedrus, limestone, 1000 m, 22-IV-2017, A.R. Burgaz (MACB 111448). GJIROKASTER: Gjirokaster County, Bredhi i Sotires above village Sotire, 39[grados]48'53.21"N 20[grados]21'41.98"E, in dry scrubland area, on bark (Platanus orientalis), 550 m, 6-V-2017, L. Lokos (BP 96680). District of Tepelene, Kelcyre, Kelcyre gorge, left side Vjosa river, 40[grados]17'52"N 20[grados]10'05"E, Carpinus betulus, Pinus halepensis, sandstone, 180 m, 18-IV-2017, A.R. Burgaz (MACB 111350). KORCE: District of Korce, Dardhe, SH90 Boboshtice-Dardhe, 40[grados]31'18"N 20[grados]49'22"E, Fagus sylvatica, Pinus nigra, Juniperus communis, Betula pendula, Abies sp., limestone, 1390 m, 23-IV-2017, A.R. Burgaz (MACB 111458). District of Korce, Dardhe, SH90 Boboshtice-Dardhe, 40[grados]31'26"N 20[grados]48'06"E, Fagus sylvatica, Juniperus communis, Abies sp., sandstone, 1550 m, 23-IV-2017, A.R. Burgaz (MACB 111454). District of Korce (Rrethi i Korces), Mali i Strugarit above village Belorta, 40[grados]39'56.56"N 20[grados]50'9.79"E, open grasslands on conglomerate rocks, 1010 m, 11-V-2015, Z. Barina, L. Lokos, D. Pifko (BP 96642). District of Pogradec, Udenisht, SH3 Piskutat-Pogradec, shore Ohrid lake, 40[grados]58'55"N 20[grados]38'21"E, Pinus halepensis, Juniperus oxycedrus, Buxus sempervirens, sandstone, 710 m, 22-IV-2017, A.R. Burgaz (MACB 111451). KUKES: District of Kukes, Kukes, Rruga Tirana, 42[grados]04'46"N 20[grados]24'45"E, Quercus cerris, deciduous Quercus, limestone, 330 m, 21-IV-2017, A.R. Burgaz (MACB 111437). LEZHE: District of Lezhe (Rrethi i Lezhes), on the slope of Mt Guri i Kuq above village Fregen, 41[grados]51'41.58"N 19[grados]44'11.22"E, on acid soil in open serpentine grasslands, 575 m, 30-III-2015, L. Lokos (BP 96606). SHKODER: Vau i Dejes, SH25 Vau i Dejes-Koman, 42[grados]03'01"N 19[grados]39'45"E, Pinus pinaster, Juniperus oxycedrus, Quercus cerris, Erica arborea, Cotinus coggygria, serpentine soil, 130 m, 20-IV-2017, A.R. Burgaz (MACB 111424). District of Malesi e Madhe, Boge, SH21 Koplik-Boge, 42[grados]21'46"N 19[grados]37'56"E, Pinus nigra, Juniperus oxycedrus, limestone, 865 m, 20-IV-2017, A.R. Burgaz (MACB 111417). TIRANE: Rrogozhine, Grethi i Mesem, 41[grados]05'00"N 19[grados]27'35"E, Pinus pinaster, Juniperus oxycedrus, sandy soil, 0 m, 15-IV-2017, A.R. Burgaz (MACB 111346). District of Tirane, Mushqeta, SH3 Mushqeta-Gracen, 41[grados]12'38"N 19[grados]57'09"E, Arbutus unedo, Erica arborea, Platanus orientalis, boulders, 480 m, 22-IV-2017, A.R. Burgaz (MACB 111441). VLORE: District of Sarande, Pasqirave, Butrint N. P., SH81, Shengjergj monastery, 39[grados]48'42"N 20[grados]00'43"E, Quercus coccifera, limestone slopes, 50 m, 17-IV-2017, A.R. Burgaz (MACB 111347). Finiq, Bistrice, Muzine, 39[grados]55'08"N 20[grados]08'48"E, Quercus coccifera, Carpinus betulus, Pinus halepensis, Cercis siliquastrum, limestone slopes, 125 m, 17-IV-2017, A.R. Burgaz (MACB 111348). Finiq, Muzine, 39[grados]54'48"N 20[grados]11'20"E, deciduous Quercus, Erica arborea, limestone, 186 m, 17-IV-2017, A.R. Burgaz (MACB 111349). District of Delvine (Rrethi i Delvines), Mali i Gjere, on the southern slope of 1331 m high mount above Delvine, 39[grados]57'42.12"N 20[grados]07'59.66"E, in chestnut woodland, on bark (Castanea sativa), 828 m, 1-V-2009, Z. Barina & D. Pifko (BP 96544).

Cladonia foliacea (Huds.) Willd.

Fumarprotocetraric and usnic acids chemotype: BERAT: District of Berat (Rrethi i Beratit), Tomor Mountains (Mali i Tomorrit), c. 700 m north of village Karkanjoz, 40[grados]41'55.61"N 20[grados]03'28.69"E, in Mediterranean scrubland, on calcareous soil in rocky grassland, 490 m, 25-V-2004, Z. Barina & Cs. Nemeth (BP 96540). District of Skrapar (Rrethi i Skraparit), above the gorge of river Osum next to village Dhores, 40[grados]28'27.79"N 20[grados]15'16.11"E, in open bushland, on calcareous soil, 420 m, 13-X-2010, L. Lokos (BP 96575). DIBER: Diber county, on the mount between villages Fushe Stude and Zabzun, 41[grados]19'54.43"N 20[grados]24'38.25"E, on calcareous soil in rocky grasslands, 1295 m, 18-V-2016, L. Lokos (BP 96668). DURRES: District of Durres, Fushe-Draci, San Pietro beach track, 41[grados]31'01"N 19[grados]31'12"E, Pinus pinea, sandy soil, 1 m, 19-IV-2017, A.R. Burgaz (MACB 111288). Prrenjas, Urake, SH9 Qafe Thane, 41[grados]04'24"N 20[grados]36'18"E, Quercus macrolepis, deciduous Quercus, Juniperus oxycedrus, limestone, 1000 m, 22-IV-2017, A.R. Burgaz (MACB 111313). ELBASAN: District of Elbasan, Bradashesh, SH3 Bradashesh-Graze, 41[grados]06'05"N 20[grados]00'26"E, Olea europea, limestone karst, 390 m, 15-IV-2017, A.R. Burgaz (MACB 111278). District of Elbasan, Bradashesh, SH3 Bradashesh-Graze, 41[grados]06'33"N 19[grados]59'49"E, Carpinus betulus, Juniperus oxycedrus, limestone karst, 530 m, 15-IV-2017, A.R. Burgaz (MACB 111279). Elbasan County (Qarku i Elbasanit), on the mount above bridge Ura e Murrashit, between Mirake and Librazhd, 41[grados]11'1.64"N 20[grados]16'33.36"E, on acid soil in serpentine scrublands, 290 m, 3-IV-2016, L. Lokos (BP 96657). Elbasan county, in village Floq, 41[grados]16'1.19"N 20[grados]14'19.17"E, on soil by the roadside, 650 m, 17-V-2016, L. Lokos (BP 96664). GJIROKASTER: Dropull, Grapsh, SH78, 39[grados]56'50"N 20[grados]14'31"E, deciduous Quercus, limestone slope rocks, 440 m, 17-IV-2017, A.R. Burgaz (MACB 111285). Dropull, Goranxi, to Kishae Ravenes, 40[grados]00'56"N 20[grados]11'15"E, limestone rocks, 268 m, 17-IV-2017, A.R. Burgaz (MACB 111286). District of Gjirokaster (Rrethi i Gjirokastres), Kurvelesh i Siperm uplands above village Picar, 40[grados]10'21.66"N 20[grados]02'40.64"E, on calcareous soil in rocky grasslands, 390 m, 29-III-2015, L. Lokos (BP 96602). District of Permet (Rrethi i Permetit), c. 1.3 km SE of village Kosine, along river Vjose, 40[grados]15'40.11"N 20[grados]19'08.84"E, on calcareous conglomerate rocks, 215 m, 17-X-2011, Z. Barina, L. Lokos & D. Pifko (BP 96580). District of Permet (Rrethi i Permetit), right part of river Vjose between villages Golemas and Kutal, 40[grados]15'22.93"N 20[grados]19'27.80"E, on conglomerate rocks, 220 m, 10-V-2015, Z. Barina, L. Lokos & D. Pifko (BP 96641). District of Tepelene, Dukaj, SH76 Tepelene-Sinanaj, 40[grados]18'45"N 19[grados]57'46"E, Quercus coccifera, limestone, 280 m, 18-IV-2017, A.R. Burgaz (MACB 111287). District of Tepelene (Rrrethi i Tepelenes), c. 1.4 km S of village Vasjar, 40[grados]21'27.84"N 19[grados]55'57.13"E, on calcareous soil in exposed rocky grasslands, 155 m, 14-X-2012, L. Lokos (BP 96596). KORCE: Kolonje District (Rrethi i Kolonjes), between town Leskovik and village Vrepcke, 40[grados]10'30.62"N 20[grados]38'22.95"E, on acid soil in rocky serpentine grasslands, 1030 m, 10-V-2015, Z. Barina, L. Lokos, D. Pifko (BP 96638). District of Korce (Rrethi i Korces), above the water reservoir c 2.5 km WSW of village Ujebardhe, 40[grados]31'17.31"N 20[grados]39'50.62"E, on acid soil in dry rocky grassland, 1105 m, 18-X-2011, L. Lokos (BP 96587). District of Devoll (Rrethi i Devollit), on the hills above village Poncare, 40[grados]31'17.63"N 20[grados]57'54.23"E, in dry grasslands, on flysh soil, 1035 m, 11-V-2015, Z. Barina, L. Lokos & D. Pifko (BP 96646). District of Korce, Dardhe, SH90 Boboshtice-Dardhe, 40[grados]31'26"N 20[grados]48'06"E, Fagus sylvatica, Juniperus communis, Abies sp., sandstone, 550 m, 23-IV-2017, A.R. Burgaz (MACB 111319). District of Korce (Rrethi i Korces), on the ridge of Mts Mali i Moraves above village Dardhe, near peak Buzeja e Boboshtices (1701 m), 40[grados]32'23.97"N 20[grados]49'14.20"E, on acid soil in beech forest on gravel conglomerate, 1550 m, 10-VIII-2010, L. Lokos (BP 96561). Korce, Boboshtice, SH90 Boboshtice-Dardhe, 40[grados]32'37"N 20[grados]47'11"E, Juniperus oxycedrus, pastureland, sandstone, 1200 m, 23-IV-2017, A.R. Burgaz (MACB 111317). District of Korce (Rrethi i Korces), Mali i Strugarit above village Belorta, 40[grados] 39' 56.56"N 20[grados]50'9.79"E, open grasslands on conglomerate rocks, 1010 m, 11-V-2015, Z. Barina, L. Lokos, D. Pifko (BP 96643). District of Devoll (Rrethi i Devollit), on the mount between village Tren and lake Liqeni i Prespes se Vogel, 40[grados]40'10.55"N 20[grados]59'12.62"E, on calcareous soil in rocky grasslands, 910 m, 12-V-2015, Z. Barina, L. Lokos & D. Pifko (BP 96648). Korce County, Mali i Lenies at village Gopesh, 40[grados]43'37.56"N 20[grados]23'56.77"E, on calcareous soil in rocky pastures, 1005 m, 3-V-2017, L. Lokos (BP 96676). Korce County, Mt "Golines" at village Rakicke, 40[grados]43'8.88"N 20[grados]59'0.97"E, on calcareous soil in rocky grasslands, 1105 m, 1-V-2017, L. Lokos (BP 96672). District of Pogradec, Udenisht, SH3 Piskutat-Pogradec, shore Ohrid lake, 40[grados]58'55"N 20[grados]38'21"E, Pinus halepensis, Juniperus oxycedrus, Buxus sempervirens, sandstone, 710 m, 22-IV-2017, A.R. Burgaz (MACB 111316). KUKES: District of Kukes, Kolsh, SH5 close Drin river, 42[grados]04'18"N 20[grados]20'32"E, deciduous Quercus, Juniperus oxycedrus, Buxus sempervirens, serpentine soil, 510 m, 21-IV-2017, A.R. Burgaz (MACB 111308). LEZHE: District of Lezhe (Rrethi i Lezhes), near the pass between Mt. "Sheshi i Veles" (1049.2 m) and Mt. "Veles" (1170.1 m), above village Kallmet, 41[grados]50'28.1"N 19[grados]42'54.5"E, on soil in rocky forest, 961 m, 23-IV-2009, L. Lokos (BP 96548). SHKODER: Shkoder District (Rrethi i Shkodres), c. 1.5 km NE of village Gomsiqe e Eperme, near road SH5, 41[grados]59'31.61"N 19[grados]48'12.99"E, on acid soil in dry serpentine grasslands, 533 m, 6-V-2015, L. Lokos (BP 96618). District of Shkoder (Rrethi i Shkodres), Qafa e Celes above village Mirdice, 42[grados]0'11.98"N 19[grados]43'33.16"E, on soil in roadside dry rocky grassland, 470 m, 12-X-2011, L. Lokos (BP 96578). Puke District (Rrethi i Pukes), Mount Rras' e Qerretit above village Qerret, 42[grados]03'36.98"N 19[grados]49 53.45"E, in serpentine rocky area, on acid soil in pine woodland, 735 m, 6-V-2015, L. Lokos (BP 96623). Shkoder District (Rrethi i Shkodres), in the valley of stream Perroi i Cibunit E of village Prekal, 42[grados]10'44.69"N 19[grados]43'18.46"E, on calcareous soil in scrubland, 220 m, 5-V-2015, L. Lokos (BP 96617). Shkoder District (Rrethi i Shkodres), between villages Linaj-Egc-Lepurosh and Kurtaj, 42[grados]11'10.22"N 19[grados]33'19.74"E, on calcareous soil in dry scrubland, 260 m, 4-V-2015, Z. Barina, L. Lokos & D. Pifko (BP 96613). District of Shkoder (Rrethi i Shkodres), in the valley of river Perroi i Thate at village Lashaj, along the road nr. E 762, 42[grados]14'19.70"N 19[grados]25'24.82"E, on calcareous soil in open dry rocky grasslands, 78 m, 30-III-2015, L. Lokos (BP 96603). District of Malesi e Madhe, Poica, SH21 Koplik-Boge, 42[grados]15>53>>N 19[grados]29>47>>E, Juniperus oxycedrus, Cupressus sempervirens, boulders, 320 m, 20-IV-2017, A.R. Burgaz (MACB 111293). District of Malesi e Madhe, Dedaj, SH21 Koplik-Boge, 42[grados]18'04"N 19[grados]32'32"E, Carpinus betulus, Juniperus oxycedrus, limestone rocks, 495 m, 20-IV-2017, A.R. Burgaz (MACB 111294). District of Malesi e Madhe, Boge, SH21 Koplik-Boge, 42[grados]21'46"N 19[grados]37'56"E, Pinus nigra, Juniperus oxycedrus, limestone, 865 m, 20-IV-2017, A.R. Burgaz (MACB 111302). Malesi e Madhe District (Rrethi i Malesise se Madhe), on Mount Bukovik above village Rrapshe-Stare, 42[grados]23'6.12"N 19[grados]29'25.56"E, on calcareous soil in rocky grasslands, 715 m, 3-V-2015, L. Lokos (BP 96608). TIRANE: District of Kavaje (Rrethi i Kavajes), on hills east of village Sterbeg-Bragash, 41[grados]06'36.2"N 19[grados]32'07.0"E, on soil in poplar woodland, 107 m, 28-IV-2009, L. Lokos (BP 96554). VLORE: Himare, Ilias, Himare, 40[grados]08'29"N 19[grados]40'11"E, Quercus coccifera, Cupressus sempervirens, Cistaceae shrubs, limestone, 300 m, 16-IV-2017, A.R. Burgaz (MACB 111283). District of Vlore (Rrrethi i Vlores), on the E ridge of Mt maja e Thanasit (1352.1 m) above Llogora pass, 40[grados]11'51.54"N 19[grados]35'27.82"E, on calcareous soil in mixed forest, 1075 m, 12-VIII-2010, L. Lokos (BP 96568). District of Vlore, Dukat i Ri, SH3 Orikum-Dhermi, Llogora N. P., 40[grados]12'00"N 19[grados]35'26"E, Pinus nigra, Juniperus oxycedrus, Buxus sempervirens, limestone, 1030 m, 16-IV-2017, A.R. Burgaz (MACB 111282). District of Vlore, Dukat i Ri, SH3 Orikum-Dhermi, Llogora N. P., 40[grados]15'11"N 19[grados]32'14"E, Quercus coccifera, Pinus halepensis, limestone, 320 m, 16-IV-2017, A.R. Burgaz (MACB 111280). Fumarprotocetraric, usnic and physodalic acids chemotype: KORCE: District of Korce, Dardhe, SH90 Boboshtice-Dardhe, 40[grados]31'18"N 20[grados]49'22"E, Fagus sylvatica, Pinus nigra, Juniperus communis, Betula pendula, Abies sp., limestone, 1390 m, 23-IV-2017, A.R. Burgaz (MACB 111320). KUKES: District of Kukes, Kukes, Rruga Tirana, 42[grados]04'46"N 20[grados]24'45"E, Quercus cerris, deciduous Quercus, limestone, 330 m, 21-IV-2017, A.R. Burgaz (MACB 111310). TIRANE: District of Tirane, Mushqeta, SH3 Mushqeta-Gracen, 41[grados]12'38"N 19[grados]57'09"E, Arbutus unedo, Erica arborea, Platanus orientalis, boulders, 480 m, 22-IV-2017, A.R. Burgaz (MACB 111311). Fumarprotocetraric, usnic and psoromic acids chemotype: KORCE: District of Kolonje (Rrethi i Kolonjes), Mt. Melesin (1410.8 m), 40[grados]08'50.1"N 20[grados]35'37.0"E, on limestone rocks, 953 m, 3-V-2009, L. Lokos (BP 96559). VLORE: District of Vlore (Rrethi i Vlores), between village Ilias and monastir Shen Theodor, 40[grados]08'07.8"N 19[grados]40'26.4"E, on soil in olive plantation, 211 m, 29-IV-2009, L. Lokos (BP 96558).

Cladonia furcata (Huds.) Schrad.

Fumarprotocetraric acid chemotype: DIBER: Diber county, on the mount between villages Fushe Stude and Zabzun, 41[grados]19'54.43"N 20[grados]24'38.25"E, on calcareous soil in rocky grasslands, 1295 m, 18-V-2016, L. Lokos (BP 96669). DURRES: District of Durres, Fushe-Draci, road cape of Rodon, 41[grados]32'50"N 19[grados]31'08"E, Carpinus betulus, Arbutus unedo, Pistacia lentiscus, Cistus salviifolius, sandstone, 90 m, 19-IV-2017, A.R. Burgaz (MACB 111220). ELBASAN: District of Librazhd (Rrethi i Librazhdit): Polis Mts, between villages Gurreshpat and Vilan, 41[grados]05'38.45"N 20[grados]16'15.72"E, on acid soil among serpentine rocks, 1270 m, 12-X-2010, L. Lokos (BP 96574). District of Elbasan, Bradashesh, SH3 Bradashesh-Graze, 41[grados]06'33"N 19[grados]59'49"E, Carpinus betulus Juniperus oxycedrus, limestone karst, 530 m, 15-IV-2017, A.R. Burgaz (MACB 111403). District of Librazhd (Rrethi i Librazhdit): Polis Mts; above village Polis, on the ridge between the valley of brook "Sut Kukit" and river Shkumbin, in bushland, 41[grados]09'16.50"N 20[grados]15'08.79"E, on conglomeratic flysh sediments, 555 m, 12-X-2010, L. Lokos (BP 96572). Elbasan County (Qarku i Elbasanit), on the mount above bridge Ura e Murrashit, between Mirake and Librazhd, 41[grados]11'1.64"N 20[grados]16'33.36"E, on acid soil in serpentine scrublands, 290 m, 3-IV-2016, L. Lokos (BP 96658). Elbasan county, in village Floq, 41[grados]16'1.19"N 20[grados]14'19.17"E, on soil by the roadside, 650 m, 17-V-2016, L. Lokos (BP 96665). GJIROKASTER: District of Gjirokaster (Rrethi i Gjirokastres), between settlements Picar and Andon Poci in the valley of river Vjose near road SH4, 40[grados]10'41.12"N 20[grados]04'45.55"E, in grazed grassland, on flysh soil, 170 m, 28-III-2015, L. Lokos (BP 96598). District of Tepelene, Kelcyre, Kelcyre gorge, left side Vjosa river, 40[grados]17'52"N 20[grados]10'05"E, Carpinus betulus, Pinus halepensis, sandstone, 180 m, 18-IV-2017, A.R. Burgaz (MACB 111215). KORCE: Kolonje District (Rrethi i Kolonjes), between town Leskovik and village Vrepcke, 40[grados]10'30.62"N 20[grados]38'22.95"E, on acid soil in rocky serpentine grasslands, 1030 m, 10-V-2015, Z. Barina, L. Lokos & D. Pifko (BP 96639). District of Kolonje (Rrethi i Kolonjes), c. 1.9 km W of village Pepellash, 40[grados]28'56.18"N 20[grados]40'21.27"E, on acid soil in moist road slope, 549 m, 18-X-2011, Z. Barina, L. Lokos & D. Pifko (BP 96582). District of Korce (Rrethi i Korces), on the ridge of Mts Mali i Moraves above village Dardhe, near peak Buzeja e Boboshtices, 40[grados]31'52.6"N 20[grados]48'36.1"E, on acid soil in beech forest on gravel conglomerate, 1625 m, 10-VIII-2010, L. Lokos (BP 96565). District of Korce, Dardhe, SH90 Boboshtice-Dardhe, 40[grados]31'18"N 20[grados]49'22"E, Fagus sylvatica, Pinus nigra, Juniperus communis, Betula pendula, Abies sp., limestone, 1390 m, 23-IV-2017, A.R. Burgaz (MACB 111344). District of Devoll (Rrethi i Devollit), on the mount between village Tren and lake Liqeni i Prespes se Vogel, 40[grados]40'10.55"N 20[grados]59'12.62"E, on calcareous soil in rocky grasslands, 910 m, 12-V-2015, Z. Barina, L. Lokos & D. Pifko (BP 96649). Korce County, Mali i Lenies at village Gopesh, 40[grados]43'37.56"N 20[grados]23'56.77"E, on calcareous soil in rocky pastures, 1005 m, 3-V-2017, L. Lokos (BP 96677). LEZHE: Puke District (Rrethi i Pukes); Mount Rras' e Qerretit above village Qerret, 42[grados]03'36.98"N 19[grados]49'53.45"E, in serpentine rocky area, on acid soil in pine woodland, 735 m, 6-V-2015, L. Lokos (BP 96624). SHKODER: District of Shkoder (Rrethi i Shkodres), Qafa e Celes above village Mirdice, 42[grados]0'11.98"N 19[grados]43'33.16"E, on soil in roadside dry rocky grassland, 470 m, 12-X-2011, L. Lokos (BP 96579). District of Shkoder (Rrethi i Shkodres), in the valley of river Perroi i Thate at village Lashaj, along the road nr. E 762, 42[grados]14'19.70"N 19[grados]25'24.82"E, on calcareous soil in open dry rocky grasslands, 78 m, 30-III-2015, L. Lokos (BP 96604). TIRANE: Tirana County (Qarku i Tiranes); at the foot of Mt Kalaja e Dorezit, between villages Peze and Dorez, 41[grados]13'30.08"N 19[grados]43'53.26"E, in grasslands, on flysh soil, 180 m, 2-IV-2016, L. Lokos (BP 96652). VLORE: District of Sarande (Rrrethi i Sarandes), on a hill between villages Kullurice and Koder, 39[grados]48'43.85"E 20[grados]06'34.02"E, in dry grassland, on flysh soil, 55 m, 13-X-2012, L. Lokos (BP 96591). Finiq, Bistrice, Muzine, 39[grados]55'08"N 20[grados]08'48"E, Quercus coccifera, Carpinus betulus, Pinus halepensis, Cercis siliquastrum, limestone slopes, 125 m, 17-IV-2017, A.R. Burgaz (MACB 111197). District of Vlore (Rrethi i Vlores), Mts. Mali i Cikes, in the valley below Mt. "Maja e Pandelejmonit" (805.8 m), north of village Dhermi, 40[grados]09'32.2"N 19[grados]38'49.4"E, in dry rocky grassland, on calcareous soil, 260 m, 29-IV-2009, L. Lokos (BP 96555). Fumarprotocetraric and physodalic acids chemotype: DURRES: District of Durres, Fushe-Draci, San Pietro beach track, 41[grados]31'01"N 19[grados]31'12"E, Pinus pinea, sandy soil, 1 m, 19-IV-2017, A.R. Burgaz (MACB 111228). ELBASAN: Prrenjas, Urake, SH9 Qafe Thane, 41[grados]04'24"N 20[grados]36'18"E, Quercus macrolepis, deciduous Quercus, Juniperus oxycedrus, limestone, 1000 m, 22-IV-2017, A.R. Burgaz (MACB 111314). GJIROKASTER: Dropull, Goranxi, to Kishae Ravenes, 40[grados]00'56"N 20[grados]11'15"E, limestone rocks, 268 m, 17-IV-2017, A.R. Burgaz (MACB 111206). GJIROKASTER: District of Permet, Piskove, left side Lumica river, K75 to Frasher, 40[grados]16'44"N 20[grados]17'07"E, Juniperus oxycedrus, Paliurus spinachristi, Arbutus unedo, Pistacia terebinthus, sandstone, 230 m, 18-IV-2017, A.R. Burgaz (MACB 111218). District of Tepelene, Kelcyre, Kelcyre gorge, left side Vjosa river, 40[grados]17'52"N 20[grados]10'05"E, Carpinus betulus, Pinus halepensis, sandstone, 180 m, 18-IV-2017, A.R. Burgaz (MACB 111213). District of Tepelene, Dukaj, SH76 Tepelene-Sinanaj, 40[grados]18'45"N 19[grados]57'46"E, Quercus coccifera, limestone, 280 m, 18-IV-2017, A.R. Burgaz (MACB 111221). KORCE: District of Korce, Dardhe, SH90 Boboshtice-Dardhe, 40[grados]31'18"N 20[grados]49'22"E, Fagus sylvatica, Pinus nigra, Juniperus communis, Betula pendula, Abies sp., limestone, 1390 m, 23-IV-2017, A.R. Burgaz (MACB 111321). District of Pogradec, Udenisht, SH3 Piskutat-Pogradec, shore Ohrid lake, 40[grados]58'55"N 20[grados]38'21"E, Pinus halepensis, Juniperus oxycedrus, Buxus sempervirens, sandstone, 710 m, 22-IV-2017, A.R. Burgaz (MACB 111315). KUKES: District of Kukes, Kolsh, SH5 close Drin river, 42[grados]04'18"N 20[grados]20'32"E, deciduous Quercus, Juniperus oxycedrus, Buxus sempervirens, serpentine soil, 510 m, 21-IV-2017, A.R. Burgaz (MACB 111307). District of Kukes, Kukes, Rruga Tirana, 42[grados]04'46"N 20[grados]24'45"E, Quercus cerris, deciduous Quercus, limestone, 330 m, 21-IV-2017, A.R. Burgaz (MACB 111309). LEZHE: District of Kurbin, Milot, left side Mat river, SH25 Milot-Zogu Bridge, 41[grados]41'54"N 19[grados]45'10"E, Carpinus orientalis, Erica arborea, limestone, 40 m, 21-IV-2017, A.R. Burgaz (MACB 111306). SHKODER: District of Malesi e Madhe, Poica, SH21 Koplik-Boge, 42[grados]15'53"N 19[grados]29'47"E, Juniperus oxycedrus, Cupressus sempervirens, boulders, 320 m, 20-IV-2017, A.R. Burgaz (MACB 111291). District of Malesi e Madhe, Dedaj, SH21 Koplik-Boge, 42[grados]18'04"N 19[grados]32'32"E, Carpinus betulus, Juniperus oxycedrus, limestone rocks, 495 m, 20-IV-2017, A.R. Burgaz (MACB 111297). District of Malesi e Madhe, Boge, SH21 Koplik-Boge, 42[grados]21'46"N 19[grados]37'56"E, Pinus nigra, Juniperus oxycedrus, limestone, 865 m, 20-IV-2017, A.R. Burgaz (MACB 111301). VLORE: Finiq, Muzine, Muzine, 39[grados]54'48"N 20[grados]11'20"E, deciduous Quercus, Erica arborea, limestone, 186 m, 17-IV-2017, A.R. Burgaz (MACB 111200). Finiq, Bistrice, Muzine, 39[grados]55'08"N 20[grados]08'48"E, Quercus coccifera, Carpinus betulus, Pinus halepensis, Cercis siliquastrum, limestone slopes, 125 m, 17-IV-2017, A.R. Burgaz (MACB 111195). Atranorin and fumarprotocetraric acid: KORCE: District of Korce (Rrethi i Korces), on the ridge of Mts Mali i Moraves above village Dardhe, near peak Buzeja e Boboshtices (1701 m), 40[grados]32'23.97"E 20[grados]49'14.20"E, on acid soil in beech forest on gravel conglomerate, 1550 m, 10-VIII-2010, L. Lokos (BP 96562). Atranorin, fumarprotocetraric and physodalic acids chemotype: KORCE: District of Korce, Dardhe, SH90 Boboshtice-Dardhe, 40[grados]31'26"N 20[grados]48'06"E, Fagus sylvatica, Juniperus communis, Abies sp., sandstone, 1550 m, 23-IV-2017, A.R. Burgaz (MACB 111318). SHKODER: District of Malesi e Madhe, Poica, SH21 Koplik-Boge, 42[grados]15'53"N 19[grados]29'47"E, Juniperus oxycedrus, Cupressus sempervirens, boulders, 320 m, 20-IV-2017, A.R. Burgaz (MACB 111290). District of Malesi e Madhe, Dedaj, SH21 Koplik-Boge, 42[grados]18'04"N 19[grados]32'32"E, Carpinus betulus, Juniperus oxycedrus, limestone rocks, 495 m, 20-IV-2017, A.R. Burgaz (MACB 111295, 111296). Malesi e Madhe, Boge, SH21 Koplik-Boge, 42[grados]21'46"N 19[grados]37'56"E, Pinus nigra, Juniperus oxycedrus, limestone, 865 m, 20-IV-2017, A.R. Burgaz (MACB 111300). VLORE: Finiq, Muzine, Muzine, 39[grados]54'48"N 20[grados]11'20"E, deciduous Quercus, Erica arborea, limestone, 186 m, 17-IV-2017, A.R. Burgaz (MACB 111284). District of Vlore, Dukat i Ri, SH3 Orikum-Dhermi, Llogora N. P., 40[grados]15'11"N 19[grados]32'14"E, Quercus coccifera, Pinus halepensis, limestone, 320 m, 16-IV-2017, A.R. Burgaz (MACB 111281). Atranorin, bourgeanic and fumarprotocetraric acids: KORCE: Korce County, Mt Golines at village Rakicke, 40[grados]43'42.80"N 20[grados]59'28.06"E, on calcareous soil in open, rocky scrublands, 1295 m, 1-V-2017, L. Lokos (BP 96674).

Cladonia humilis (With.) J.R. Laundon

Atranorin and fumarprotocetraric acid: ELBASAN: Prrenjas, Prrenjas, SH3 Qukes-Prrenjas, 41[grados]04'24"N 20[grados]30'15"E, Carpinus orientalis, Quercus macrolepis, deciduous Quercus, Buxus sempervirens, limestone slopes, 540 m, 22-IV-2017, A.R. Burgaz (MACB 111444). VLORE: District of Sarande, Pasqirave, Butrint N. P., SH81, Shengjergj monastery, 39[grados]48'42"N 20[grados]00'43"E, Quercus coccifera, limestone slopes, 50 m, 17-IV-2017, A.R. Burgaz (MACB 111407).

Cladonia macilenta Hoffm.

Barbatic and thamnolic acids: LEZHE: District of Mirdite (Rrethi i Mirdites); in the valley of stream Fani i Vogel above village Thirre; in serpentine rocky area, on bark (Pinus nigra). 42[grados] 0' 20.98"N 20[grados]14'33.25"E, 905 m, 6-V-2015, L. Lokos (BP 96626).

Cladonia merochlorophaea Asahina

Fumarprotocetraricandmerochlorophaeic acids: KORCE: District of Korce (Rrethi i Korces), on the ridge of Mts Mali i Moraves above village Dardhe, near peak Buzeja e Boboshtices, on calcareous soil, 40[grados]31'52.6"N 20[grados]48'36.1"E, 1625 m, 10-VIII-2010, L. Lokos (BP 96566).

Cladonia parasitica (Hoffm.) Hoffm.

Barbatic and thamnolic acids: VLORE: District of Delvine (Rrethi i Delvines), Mali i Gjere, on the southern slope of 1331 m high mount above Delvine; in chestnut woodland, on bark (Castanea sativa), 39[grados]57'42.12"N 20[grados] 07'59.66"E, 828 m, 1-V-2009, Z. Barina & D. Pifko (BP 96545).

Cladonia peziziformis (With.) J.R. Laundon

Fumarprotocetraric acid: SHKODER: Vau i Dejes, SH25 Vau i Dejes-Koman, 42[grados]00'55"N 19[grados]39'37"E, Pinus pinaster, Carpinus orientalis, serpentine soil, 120 m, 20-IV-2017, A.R. Burgaz (MACB 111418).

Cladonia polycarpoides Nyl. (= C. subcariosa auct.)

Norstictic acid chemotype: SHKODER: Vau i Dejes, SH25 Vau i Dejes-Koman, 42[grados]03'01"N 19[grados]39'45"E, Pinus pinaster, Juniperus oxycedrus, Quercus cerris, Erica arborea, Cotinus coggygria, serpentine soil, 130 m, 20-IV-2017, A.R. Burgaz (MACB 111427). Atranorin and norstictic acid chemotype: KUKES: District of Kukes, Kolsh, SH5 close Drin river, 42[grados]04'18"N 20[grados]20'32"E, deciduous Quercus, Juniperus oxycedrus, Buxus sempervirens, serpentine soil, 510 m, 21-IV-2017, A.R. Burgaz (MACB 111432).

Cladonia pyxidata (L.) Hoffm.

Fumarprotocetraric acid chemotype: DIBER: Diber county, Golloborde, in the dry valley Gropa Gjugarica above village Sebisht, 41[grados]23'34.22"N 20[grados]25'16.33"E, on calcareous soil in rocky grasslands, 1015 m, 19-V-2016, L. Lokos (BP 96670). Diber county, Mt Serakoli above villae Ploshtan, 41[grados]50'44.99"N 20[grados]26'43.85"E, on calcareous soil in rocky pastures, 1200 m, 21-V-2016, L. Lokos (BP 96671). DURRES: District of Kruje, Kruje, SH38 to Qafe-Shtame N. P., 41[grados]31'03"N 19[grados]49'27"E, Carpinus betulus, pasturelands, limestone, 720 m, 19-IV-2017, A.R. Burgaz (MACB 111229). ELBASAN: Elbasan county, in open boxwood scrubland above village Kuterman, 41[grados]13'40.99"N 20[grados]14'41.04"E, on acid soil in rocky serpentine area, 510 m, 16-V-2016, L. Lokos (BP 96661). FIER: Fier District (Rrethi i Fierit), Mali i Kalivacit, S of village Kalivac, above river Vjose, 40[grados]24'14.86"N 19[grados]48'4.08"E, on calcareous soil in Quercus coccifera scrubland, 100 m, 8-V-2015, L. Lokos (BP 96629). GJIROKASTER: Gjirokaster County, Bredhi i Sotires above village Sotire, 39[grados]48'53.21"N 20[grados]21'41.98"E, on mossy rocks in dry scrublands, 550 m, 6-V-2017, L. Lokos (BP 96681). Gjirokaster County, at village Dervician, 40[grados]01'24.45"N 20[grados]10'39.21"E, on calcareous soil in rocky grasslands, 260 m, 7-V-2017, L. Lokos (BP 96684). District of Permet (Rrethi i Permetit), c. 1.3 km SE of village Kosine, along river Vjose, 40[grados]15'40.11"N 20[grados]19'8.84"E, on calcareous conglomerate rocks, 215 m, 17-X-2011, Z. Barina, L. Lokos, D. Pifko (BP 96581). District of Permet (Rrethi i Permetit), left part of river Vjose, between town Kelcyre and village Mbrezhdan, 40[grados]18'17.42"N 20[grados]12'17.52"E, in scrubland, on flysh soil, 205 m, 9-V-2015, L. Lokos (BP 96686). Tepelene District (Rrethi i Tepelenes), in village Mezhgoran, 40[grados]19'11.24"N 20[grados]05'43.54"E, on calcareous soil, 660 m, 9-V-2015, Z. Barina, L. Lokos, D. Pifko (BP 96634). KORCE: District of Pogradec, Udenisht, SH3 Piskutat-Pogradec, shore Ohrid lake, 40[grados]58'55"N 20[grados]38'21"E, Pinus halepensis, Juniperus oxycedrus, Buxus sempervirens, sandstone, 710 m, 22-IV-2017, A.R. Burgaz (MACB 111341). District of Korce (Rrethi i Korces); Mali i Strugarit above village Belorta, 40[grados]39'56.56"N 20[grados] 50' 9.79"E, open grasslands on conglomerate rocks, 1010 m, 11-V-2015, Z. Barina, L. Lokos, D. Pifko (BP 96644). District of Devoll (Rrethi i Devollit), on the mount between village Tren and lake Liqeni i Prespes se Vogel, 40[grados]40'10.55"N 20[grados]59'12.62"E, on calcareous soil in rocky grasslands, 910 m, 12-V-2015, Z. Barina, L. Lokos, D. Pifko (BP 96650). Korce County, Mt Golines at village Rakicke, 40[grados]43'8.88"N 20[grados]59'0.97"E, on calcareous soil in rocky grasslands, 1105 m, 1-V-2017, L. Lokos (BP 96673). District of Korce (Rrethi i Korces), Thate Mountains (Mali i Thate), c. 3.0 km north of village Zvezde, c. 1.85 km south-southeast of the peak of Mount "maja e Shengjergjit" (1921 m), 40[grados]44'37.16"N 20[grados]52'35.53"E, on calcareous soil in rocky grasslands, 1165 m, 18-X-2011, L. Lokos (BP 96589). SHKODER: Vau i Dejes, SH25 Vau i Dejes-Koman, 42[grados]03'01"N 19[grados]39'45"E, Pinus pinaster, Juniperus oxycedrus, Quercus cerris, Erica arborea, Cotinus coggygria, serpentine soil, 130 m, 20-IV-2017, A.R. Burgaz (MACB 111423). Shkoder District (Rrethi i Shkodres), in the valley of stream Perroi i Cibunit E of village Prekal, 42[grados]10'50.54"N 19[grados]43'42.57"E, on calcareous soil in scrubland, 250 m, 5-V-2015, L. Lokos (BP 96615). District of Malesi e Madhe, Dedaj, SH21 Koplik-Boge, 42[grados]18'04"N 19[grados]32'32"E, Carpinus betulus, Juniperus oxycedrus, limestone rocks, 495 m, 20-IV-2017, A.R. Burgaz (MACB 111326). District of Malesi e Madhe, Boge, SH21 Koplik-Boge, 42[grados]21'46"N 19[grados]37'56"E, Pinus nigra, Juniperus oxycedrus, limestone, 865 m, 20-IV-2017, A.R. Burgaz (MACB 111327). Malesi e Madhe District (Rrethi i Malesise se Madhe), on Mount Bukovik above village Rrapshe-Stare, 42[grados]23'6.12"N 19[grados]29'25.56"E, on calcareous soil in rocky grasslands, 715 m, 3-V-2015, L. Lokos (BP 96609). TIRANE: District of Tirane, Mushqeta, SH3 Mushqeta-Gracen, 41[grados]12'38"N 19[grados]57'09"E, Arbutus unedo, Erica arborea, Platanus orientalis, boulders, 480 m, 22-IV-2017, A.R. Burgaz (MACB 111337). Tirana County (Qarku i Tiranes), between village Pellumbas and cave Shpella e Pellumbasit, 41[grados]15'4.10"E 19[grados]57'34.16"E, on calcareous soil in Carpinus orientalis woodland, 445 m, 3-IV-2016, L. Lokos (BP 96654). Tirana County (Qarku i Tiranes), near cave Shpella e Pellumbasit above village Pellumbas, 41[grados]15'21.11"N 19[grados]57'50.31"E, on calcareous soil in Carpinus orientalis woodland, 450 m, 3-IV-2016, L. Lokos (BP 96656). VLORE: District of Sarande (Rrrethi i Sarandes), on the eastern slope of Mt Maja e Milletit, above village Sopik, 39[grados]46'12.49"N 20[grados]06'20.58"E, on calcareous soil in rocky grasslands, 160 m, 13-X-2012, L. Lokos (BP 96594). District of Sarande (Rrrethi i Sarandes), on a hill between villages Kullurice and Koder, 39[grados]48'43.85"N 20[grados] 06' 34.02"E, in dry grassland, on flysh soil, 55 m, 13-X-2012, L. Lokos (BP 96592). Ilias, Himare, 40[grados]08'29"N 19[grados]40'11"E, Quercus coccifera, Cupressus sempervirens, Cistaceae shrubs, limestone, 300 m, 16-IV-2017, A.R. Burgaz (MACB 110961). District of Vlore (Rrrethi i Vlores); on the E ridge of Mt maja e Thanasit (1352.1 m) above Llogora pass, 40[grados]11'51.54"N 19[grados]35'27.82"E, on calcareous soil in mixed forest, 1075 m, 12-VIII-2010, L. Lokos (BP 96569). Fumarprotocetraric and physodalic acids chemotype: GJIROKASTER: Dropull, Grapsh, SH78, 39[grados]56'50"N 20[grados]14'31"E, deciduous Quercus, limestone slope rocks, 440 m, 17-IV-2017, A.R. Burgaz (MACB 111201, 111203). Dropull, Goranxi, to Kishae Ravenes, 40[grados]00'56"N 20[grados]11'15"E, limestone rocks, 268 m, 17-IV-2017, A.R. Burgaz (MACB 111205). District of Tepelene, Koder, left Vjosa river bank, 40[grados]17'20"N 20[grados]04'25"E, Paliurus spina-christi, Spartium junceum, Phlomis shrubs, limestone slopes, 160 m, 18-IV-2017, A.R. Burgaz (MACB 111208, 111209). District of Tepelene, Kelcyre, Kelcyre gorge, left side Vjosa river, 40[grados]17'52"N 20[grados]10'05"E, Carpinus betulus, Pinus halepensis, sandstone, 180 m, 18-IV-2017, A.R. Burgaz (MACB 111212, 111214). VLORE: Finiq, Bistrice, Muzine, 39[grados]55'08"N 20[grados]08'48"E, Quercus coccifera, Carpinus betulus, Pinus halepensis, Cercis siliquastrum, limestone slopes, 125 m, 17-IV-2017, A.R. Burgaz (MACB 111192). District of Vlore, Dukat i Ri, SH3 Orikum-Dhermi, Llogora N. P., 40[grados]12'00"N 19[grados]35'26"E, Pinus nigra, Juniperus oxycedrus, Buxus sempervirens, limestone, 1030 m, 16-IV-2017, A.R. Burgaz (MACB 110957, 110958). District of Vlore, Dukat i Ri, SH3 Orikum-Dhermi, Llogora N. P., 40[grados]15'11"N 19[grados]32'14"E, Quercus coccifera, Pinus halepensis, limestone, 320 m, 16-IV-2017, A.R. Burgaz (MACB 110954, 110955). Fumarprotocetraric and psoromic acids chemotype: BERAT: District of Berat (Rrethi i Beratit), Tomor Mountains (Mali i Tomorrit), c. 1800 m SSW of village Tomor; in Mediterranean scrubland, on calcareous soil in rocky grassland, 40[grados]40'53.98"N 20[grados] 07'06.64"E, 1080 m, 25-V-2004, Z. Barina & Cs. Nemeth (BP 96539). ELBASAN, Prrenjas, Prrenjas, SH3 Qukes-Prrenjas, 41[grados]04'24"N 20[grados]30'15", Carpinus orientalis, Quercus macrolepis, deciduous Quercus, Buxus sempervirens, limestone slopes, 540 m, 22-IV-2017, A.R. Burgaz (MACB 111312).

Cladonia ramulosa (With.) J.R. Laundon

Fumarprotocetraric acid: DURRES: District of Durres, Fushe-Draci, San Pietro beach track, 41[grados]31'01"N 19[grados]31'12"E, Pinus pinea, sandy soil, 1 m, 19-IV-2017, A.R. Burgaz (MACB 111226). LEZHE: Kurbin, Milot, left side Mat river, SH25 Milot-Zogu Bridge, 41[grados]41'54"N 19[grados]45'10"E, Carpinus orientalis, Erica arborea, limestone, 40 m, 21-IV-2017, A.R. Burgaz (MACB 111332, GenBank accession no: MK543954).

Cladonia rangiformis Hoffm.

Atranorin and rangiformic acid chemotype: BERAT: District of Berat (Rrethi i Beratit), Tomor Mountains (Mali i Tomorrit), c. 700 m north of village Karkanjoz, 40[grados]41'55.61"N 20[grados]03'28.69"E, in Mediterranean scrubland, on calcareous soil in rocky grassland, 490 m, 25-V-2004, Z. Barina, Cs. Nemeth (BP 96541). ELBASAN: Elbasan county, in open boxwood scrubland above village Kuterman, 41[grados]13'40.99"N 20[grados]14'41.04"E, on acid soil in rocky serpentine area, 510 m, 16-V-2016, L. Lokos (BP 96662). GJIROKASTER: District of Permet (Rrethi i Permetit), left part of river Vjose, between town Kelcyre and village Mbrezhdan, 40[grados]18'17.42"N 20[grados]12'17.52"E, in scrubland, on flysh soil, 205 m, 9-V-2015, L. Lokos (BP 96636). KORCE: Kolonje District (Rrethi i Kolonjes); between town Leskovik and village Vrepcke, 40[grados]10'30.62"N 20[grados]38'22.95"E, on acid soil in rocky serpentine grasslands, 1030 m, 10-V-2015, Z. Barina, L. Lokos & D. Pifko (BP 96640). District of Devoll (Rrethi i Devollit), on the hills above village Poncare, 40[grados]31'17.63"N 20[grados]57'54.23"E, in dry grasslands, on flysh soil, 1035 m, 11-V-2015, Z. Barina, L. Lokos & D. Pifko (BP 96647). Korce County, Mal Morave above village Hocisht, 40[grados]36'12.59"N 20[grados]54'33.39"E, on calcareous soil in oak woodland, 1030 m, 2-V-2017, L. Lokos (BP 96675). District of Korce (Rrethi i Korces), Mali i Strugarit above village Belorta, 40[grados]39'56.56"N 20[grados]50'9.79"E, open grasslands on conglomerate rocks, 1010 m, 11-V-2015, Z. Barina, L. Lokos & D. Pifko (BP 96645). District of Devoll (Rrethi i Devollit), on the mount between village Tren and lake Liqeni i Prespes se Vogel, 40[grados]40'10.55"N 20[grados]59'12.62"E, on calcareous soil in rocky grasslands, 910 m, 12-V-2015, Z. Barina, L. Lokos & D. Pifko (BP 96651). District of Pogradec, Udenisht, SH3 Piskutat-Pogradec, shore Ohrid lake, 40[grados]58'55"N 20[grados]38'21"E, Pinus halepensis, Juniperus oxycedrus, Buxus sempervirens, sandstone, 710 m, 22-IV-2017, A.R. Burgaz (MACB 111342). KUKES: District of Kukes, Kolsh, SH5 close Drin river, 42[grados]04'18"N 20[grados]20'32"E, deciduous Quercus, Juniperus oxycedrus, Buxus sempervirens, serpentine soil, 510 m, 21-IV-2017, A.R. Burgaz (MACB 111334). LEZHE: District of Lezhe (Rrethi i Lezhes), on the slope of Mt Guri i Kuq above village Fregen, 41[grados]51'41.58"N 19[grados]44'11.22"E, on acid soil in open serpentine grasslands, 575 m, 30-III-2015, L. Lokos (BP 96607). SHKODER: Shkoder District (Rrethi i Shkodres), c. 1.5 km NE of village Gomsiqe e Eperme, near road SH5, 41[grados]59'31.61"N 19[grados]48'12.99"E, on acid soil in dry serpentine grasslands, 533 m, 6-V-2015, L. Lokos (BP 96619). Puke District (Rrethi i Pukes); Mount Rras' e Qerretit above village Qerret, 42[grados]03'36.98"N 19[grados]49'53.45"E, in serpentine rocky area, on acid soil in pine woodland, 735 m, 6-V-2015, L. Lokos (BP 96625). District of Shkoder (Rrethi i Shkodres), in the valley of river Perroi i Thate at village Lashaj, along the road nr. E 762, 42[grados]14'19.70"N 19[grados]25'24.82"E, on calcareous soil in open dry rocky grasslands, 78 m, 30-III-2015, L. Lokos (BP 96605). VLORE: District of Vlore (Rrrethi i Vlores), at seashore rruget e Bardha west of village Palase, 40[grados]10'30.46"N 19[grados]36'9.32"E, on calcareous soil in dry roadside scrubland, 225 m, 12-X-2012, L. Lokos (BP 96590). Atranorin, fumarprotocetraric and rangiformic acids chemotype: BERAT: District of Skrapar (Rrethi i Skraparit), above the gorge of river Osum next to village Dhores, 40[grados]28'27.79"N 20[grados]15'16.11"E, in open bushland, on calcareous soil, 420 m, 13-X-2010, L. Lokos (BP 96576). DIBER: District of Mat (Rrethi i Matit), Mali i Skenderbeut, on the northern forehills of Mt. "maja e Mellezit", near village Shkopet, 41[grados]41'28.5"N 19[grados]49'14.8"E, on soil, 222 m, 24-IV-2009, L. Lokos (BP 96550). ELBASAN: Prrenjas, SH3 Qukes-Prrenjas, 41[grados]04'24>>N 20[grados]30'15>>E, Carpinus orientalis, Quercus macrolepis, deciduous Quercus, Buxus sempervirens, limestone slopes, 540 m, 22-IV-2017, A.R. Burgaz (MACB 111339). Prrenjas, Urake, SH9 Qafe Thane, 41[grados]04'24"N 20[grados]36'18"E, Quercus macrolepis, deciduous Quercus, Juniperus oxycedrus, limestone, 1000 m, 22-IV-2017, A.R. Burgaz (MACB 111340). District of Elbasan, Bradashesh, SH3 Bradashesh-Graze, 41[grados]06'05"N 20[grados]00'26"E, Olea europea, limestone karst, 390 m, 15-IV-2017, A.R. Burgaz (MACB 111401). District of Elbasan, Bradashesh, SH3 Bradashesh-Graze, 41[grados]06'33"N 19[grados]59'49"E, Carpinus betulus, Juniperus oxycedrus, limestone karst, 530 m, 15-IV-2017, A.R. Burgaz (MACB 111404). District of Librazhd (Rrethi i Librazhdit): Polis Mts, above village Polis, on the ridge between the valley of brook "Sut Kukit" and river Shkumbin, 41[grados]09'16.50"N 20[grados]15'08.79"E, in bushland, on conglomeratic flysh sediments, 555 m, 12-X-2010, L. Lokos (BP 96573). Elbasan County (Qarku i Elbasanit), on the mount above bridge Ura e Murrashit, between Mirake and Librazhd, 41[grados]11'1.64"N 20[grados]16'33.36"E, on acid soil in serpentine scrublands, 290 m, 3-IV-2016, L. Lokos (BP 96659). Elbasan county, in village Floq, 41[grados]16'1.19"N 20[grados]14'19.17"E, on soil by the roadside, 650 m, 17-V-2016, L. Lokos (BP 96666). FIER: District of Fier (Rrrethi i Fierit), Pishe-Poro Nature Reserve, between village Pishe and the Adriatik Sea, 40[grados]40'3.67"N 19[grados]22'17.76"E, on soil in stone pine forest, 5 m, 12-VIII-2010, L. Lokos (BP 96567). Lushnje District (Rrethi i Lushnjes), Golda e Kullarit W of village Kular, ca 4 km NW of Divjake, 41[grados]01'6.54"N 19[grados]29'24.54"E, in pine woodland on sandy seacoast area, 5 m, 7-V-2015, L. Lokos (BP 96628). Fier District (Rrethi i Fierit), Mali i Kalivacit, S of village Kalivac, above river Vjose, 40[grados]24'14.86"N 19[grados]48'4.08"E, on calcareous soil in Quercus coccifera scrubland, 100 m, 8-V-2015, L. Lokos (BP 96630). Fier County, Pylli i Levanit, above village Levan, 40[grados]40'48.99"N 19[grados]30'11.70"E, on calcareous soil in oak woodland, 95 m, 5-V-2017, L. Lokos (BP 96679). GJIROKASTER: Gjirokaster County, Bredhi i Sotires above village Sotire, 39[grados]48'53.21"N 20[grados]21'41.98"E, on mossy rocks in dry scrublands, 550 m, 6-V-2017, L. Lokos (BP 96682). Gjirokaster County, N of village Koshovice, in Platanus woodland, 39[grados]50'11.52"N 20[grados]23'51.90"E, on soil in rocky grasslands, 430 m, 6-V-2017, L. Lokos (BP 96683). Dropull, Grapsh, SH78, 39[grados]56'50"N 20[grados]14'31"E, deciduous Quercus, limestone slope rocks, 440 m, 17-IV-2017, A.R. Burgaz (MACB 111204). Dropull, Goranxi, to Kishae Ravenes, 40[grados]00'56"N 20[grados]11'15"E, limestone rocks, 268 m, 17-IV-2017, A.R. Burgaz (MACB 111207). Gjirokaster District (Rrethi i Gjirokastres), Mount Maja e Lalucit above village Saraqinishte, 40[grados]06'57.10"N 20[grados]14'20.54"E, on calcareous soil in rocky grasslands, 1055 m, 9-V-2015, L. Lokos (BP 96632). District of Gjirokaster (Rrethi i Gjirokastres), between settlements Picar and Andon Poci in the valley of river Vjose near road SH4, 40[grados]10'41.12"N 20[grados]04'45.55"E, in grazed grassland, on flysh soil, 170 m, 28-III-2015, L. Lokos (BP 96599). District of Gjirokaster (Rrethi i Gjirokastres); Kurvelesh i Siperm uplands above village Picar, 40[grados]11'0.39"N 20[grados]02'31.51"E, on calcareous soil in rocky grasslands, 750 m, 29-III-2015, L. Lokos (BP 96601). District of Tepelene, Koder, left Vjosa river bank, 40[grados]17'20"N 20[grados]04'25"E, Paliurus spina-christi, Spartium junceum, Phlomis shrubs, limestone slopes, 160 m, 18-IV-2017, A.R. Burgaz (MACB 111210). Tepelene District (Rrethi i Tepelenes), in village Mezhgoran, 40[grados]19'11.24"N 20[grados]05'43.54"E, on calcareous soil, 660 m, 9-V-2015, Z. Barina, L. Lokos, D. Pifko (BP 96635). District of Tepelene (Rrrethi i Tepelenes), c. 1.4 km S of village Vasjar, 40[grados]21'27.84"N 19[grados]55'57.13"E, on calcareous soil in exposed rocky grasslands, 155 m, 14-X-2012, L. Lokos (BP 96597). KORCE: Kolonje District (Rrethi i Kolonjes); between town Leskovik and village Vrepcke, 40[grados]10'30.62"N 20[grados]38'22.95"E, on acid soil in rocky serpentine grasslands, 1030 m, 10-V-2015, Z. Barina, L. Lokos, D. Pifko (BP 96640). District of Kolonje (Rrethi i Kolonjes), c. 1.9 km W of village Pepellash, 40[grados]28'56.18"E 20[grados]40'21.27"N, on acid soil in moist road slope, 549 m, 18-X-2011, Z. Barina, L. Lokos, D. Pifko (BP 96583). District of Devoll (Rrethi i Devollit), on the hills above village Poncare, 40[grados]31'17.63"N 20[grados]57'54.23"E, in dry grasslands, on flysh soil, 1035 m, 11-V-2015, Z. Barina, L. Lokos, D. Pifko (BP 96647). District of Korce (Rrethi i Korces), above the water reservoir c 2.5 km WSW of village Ujebardhe, 40[grados]31'17.31"N 20[grados]39'50.62"E, on acid soil in dry rocky grassland, 1105 m, 18-X-2011, L. Lokos (BP 96588). District of Korce, Boboshtice, SH90 Boboshtice-Dardhe, 40[grados]32'37"N 20[grados]47'11"E, Juniperus oxycedrus, pastureland, sandstone, 1200 m, 23-IV-2017, A.R. Burgaz (MACB 111343). District of Korce (Rrethi i Korces); Mts Mali i Moraves, near the peak of Mt Maja e Shendellise (1365 m) above village Mborje, 40[grados]36'39.81"E 20[grados] 48' 55.00"N, on acid soil in serpentine rocky area, 1330 m, 9-VIII-2010, L. Lokos (BP 96560). KUKES: District of Kukes, Kolsh, SH5 close Drin river, 42[grados]04'18"N 20[grados]20'32"E, deciduous Quercus, Juniperus oxycedrus, Buxus sempervirens, serpentine soil, 510 m, 21-IV-2017, A.R. Burgaz (MACB 111333). District of Kukes, Rruga Tirana, 42[grados]04'46"N 20[grados]24'45"E, Quercus cerris, deciduous Quercus, limestone, 330 m, 21-IV-2017, A.R. Burgaz (MACB 111335). SHKODER: Vau i Dejes, SH25 Vau i Dejes-Koman, 42[grados]00'55"N 19[grados]39'37"E, Pinus pinaster, Carpinus orientalis, serpentine soil, 120 m, 20-IV-2017, A.R. Burgaz (MACB 111422). Vau i Dejes, SH25 Vau i Dejes-Koman, 42[grados]03'01"N 19[grados]39'45"E, Pinus pinaster, Juniperus oxycedrus, Quercus cerris, Erica arborea, Cotinus coggygria, serpentine soil, 130 m, 20-IV-2017, A.R. Burgaz (MACB 111329). Vau i Dejes, Karme, SH25 Vau i Dejes-Koman, 42[grados]04'02"N 19[grados]40'59"E, Quercus macrolepis, Q. pubescens, Juniperus oxycedrus, Erica arborea, slates, 130 m, 20-IV-2017, A.R. Burgaz (MACB 111331). District of Shkoder (Rrethi i Shkodres), in the valley of river Perroi i Thate at village Lashaj, along the road nr. E 762, 42[grados]14'19.70"N 19[grados]25'24.82"E, on calcareous soil in open dry rocky grasslands, 78 m, 30-III-2015, L. Lokos (BP 96605). LEZHE: District of Lezhe (Rrethi i Lezhes), Mali i Rrencit, near the peak of Mt Burga (262 m), above village Shengjin, 41[grados]49'28.2"N 19[grados]35'03.9"E, in open oak woodland, on limestone together with bryophytes, 255 m, 22-IV-2009, L. Lokos (BP 96547). SHKODER: Shkoder District (Rrethi i Shkodres), in the valley of stream Perroi i Cibunit E of village Prekal, 42[grados]10'50.54"N 19[grados]43'42.57"E, on calcareous soil in scrubland, 250 m, 5-V-2015, L. Lokos (BP 96616). Shkoder District (Rrethi i Shkodres), between villages Linaj-Egc-Lepurosh and Kurtaj, 42[grados]11'10.22"N 19[grados]33'19.74"E, on calcareous soil in dry scrubland, 260 m, 4-V-2015, Z. Barina, L. Lokos, D. Pifko (BP 96614). Malesi e Madhe District (Rrethi i Malesise se Madhe), on Mount Bukovik above village Rrapshe-Stare, 42[grados]23'6.12"N 19[grados]29'25.56"E, on calcareous soil in rocky grasslands, 715 m, 3-V-2015, L. Lokos (BP 96610). TIRANE: District of Kavaje (Rrethi i Kavajes), by the coast of the Adriatic Sea between villages Spille and Rreth (Rreth-Greth), 41[grados]05'07.6"N 19[grados]27'31.5"E, on sandy soil in sand dune area, 10 m, 27-IV-2009, L. Lokos (BP 96553). Tirana County (Qarku i Tiranes), at the foot of Mt Kalaja e Dorezit, between villages Peze and Dorez, 41[grados]13'30.08"N 19[grados]43'53.26"E, in grasslands, on flysh soil, 180 m, 2-IV-2016, L. Lokos (BP 96653). Tirana County (Qarku i Tiranes), between village Pellumbas and cave Shpella e Pellumbasit, 41[grados]15'4.10"N 19[grados]57'34.16"E, on calcareous soil in Carpinus orientalis woodland, 445 m, 3-IV-2016, L. Lokos (BP 96655). VLORE: Sarande, Pasqirave, Butrint N. P., SH81, Shengjergj monastery, 39[grados]44'51"N 20[grados]00'24"E, Quercus coccifera, Olea europea, Phlomis sp., limestone, 60 m, 17-IV-2017, A.R. Burgaz (MACB 111408). District of Sarande (Rrethi i Sarandes), Mt maja e Dhema, between village Ksamil and Pasqyre, 39[grados]47'28.03"N 20[grados]0'31.43"E, on calcareous soil, 60 m, 17-X-2010, L. Lokos (BP 96577). District of Sarande (Rrrethi i Sarandes), on a hill between villages Kullurice and Koder, 39[grados]48'43.85"N 20[grados]06'34.02"E, in dry grassland, on flysh soil, 55 m, 13-X-2012, L. Lokos (BP 96593). District of Sarande (Rrethi i Sarandes), north of village Gjashte, towards Delvine, 39[grados]53'34.58"N 20[grados]01'58.26"E, in open rocky grassland, on limestone together with bryophytes, 45 m, 1-V-2009, Z. Barina, L. Lokos & D. Pifko (BP 96546). Finiq, Muzine, Muzine, 39[grados]54'48"N 20[grados]11'20"E, deciduous Quercus, Erica arborea, limestone, 186 m, 17-IV-2017, A.R. Burgaz (MACB 111202). Finiq, Bistrice, Muzine, 39[grados]55'08"N 20[grados]08'48"E, Quercus coccifera, Carpinus betulus, Pinus halepensis, Cercis siliquastrum, limestone slopes, 125 m, 17-IV-2017, A.R. Burgaz (MACB 111194). Himare, Ilias, Himare, 40[grados]08'29"N 19[grados]40'11"E, Quercus coccifera, Cupressus sempervirens, Cistaceae shrubs, limestone, 300 m, 16-IV-2017, A.R. Burgaz (MACB 111406). District of Vlore (Rrethi i Vlores), Mts. Mali i Cikes, in the valley below Mt. "Maja e Pandelejmonit" (805.8 m), north of village Dhermi, 40[grados]09'32.2"N 19[grados]38'49.4"E, in dry rocky grassland, on calcareous soil, 260 m, 29-IV-2009, L. Lokos (BP 96556). District of Vlore (Rrrethi i Vlores), on the E ridge of Mt maja e Thanasit (1352.1 m) above Llogora pass, 40[grados]11'51.54"N 19[grados]35'27.82"E, on calcareous soil in mixed forest, 1075 m, 12-VIII-2010, L. Lokos (BP 96570). Atranorin, fumarprotocetraric, rangiformic and physodalic acids chemotype: DURRES: District of Durres, Fushe-Draci, San Pietro beach track, 41[grados]31'01"N 19[grados]31'12"E, Pinus pinea, sandy soil, 1 m, 19-IV-2017, A.R. Burgaz (MACB 111227). Kruje, Kruje, SH38 to Qafe-Shtame N. P., 41[grados]31'03"N 19[grados]49'27"E, Carpinus betulus, pasturelands, limestone, 720 m, 19-IV-2017, A.R. Burgaz (MACB 111289). District of Durres, Fushe-Draci, road cape of Rodon, 41[grados]32'50"N 19[grados]31'08"E, Carpinus betulus, Arbutus unedo, Pistacia lentiscus, Cistus salviifolius, sandstone, 90 m, 19-IV-2017, A.R. Burgaz (MACB 111223). ELBASAN: District of Elbasan, Bradashesh, SH3 Bradashesh-Graze, 41[grados]06'05"N 20[grados]00'26"E, Olea europaea, limestone karst, 390 m, 15-IV-2017, A.R. Burgaz (MACB 110952). District of Elbasan, Bradashesh, SH3 Bradashesh-Graze, 41[grados]06'33"N 19[grados]59'49"E, Carpinus betulus Juniperus oxycedrus, limestone karst, 530 m, 15-IV-2017, A.R. Burgaz (MACB 110953). GJIROKASTER: District of Tepelene, Koder, left Vjosa river bank, 40[grados]17'20"N 20[grados]04'25"E, Paliurus spina-christi, Spartium junceum, Phlomis shrubs, limestone slopes, 160 m, 18-IV-2017, A.R. Burgaz (MACB 111211). District of Tepelene, Kelcyre, Kelcyre gorge, left side Vjosa river, 40[grados]17'52"N 20[grados]10'05"E, Carpinus betulus, Pinus halepensis, sandstone, 180 m, 18-IV-2017, A.R. Burgaz (MACB 111216). District of Tepelene, Dukaj, SH76 Tepelene-Sinanaj, 40[grados]18'45"N 19[grados]57'46"E, Quercus coccifera, limestone, 280 m, 18-IV-2017, A.R. Burgaz (MACB 111219). District of Permet, Piskove, left side Lumica river, K75 to Frasher, 40[grados]16'44"N 20[grados]17'07"E, Juniperus oxycedrus, Paliurus spina-christi, Arbutus unedo, Pistacia terebinthus, sandstone, 230 m, 18-IV-2017, A.R. Burgaz (MACB 111217). SHKODER: Vau i Dejes, SH25 Vau i Dejes-Koman, 42[grados]00'55"N 19[grados]39'37"E, Pinus pinaster, Carpinus orientalis, serpentine soil, 120 m, 20-IV-2017, A.R. Burgaz (MACB 111304). Karme, SH25 Vau i Dejes-Koman, 42[grados]04'02"N 19[grados]40'59"E, Quercus macrolepis, Q. pubescens, Juniperus oxycedrus, Erica arborea, slates, 130 m, 20-IV-2017, A.R. Burgaz (MACB 111330). District of Malesi e Madhe, Poica, SH21 Koplik-Boge, 42[grados]15'53"N 19[grados]29'47"E, Juniperus oxycedrus, Cupressus sempervirens, boulders, 320 m, 20-IV-2017, A.R. Burgaz (MACB 111324). District of Malesi e Madhe, Dedaj, SH21 Koplik-Boge, 42[grados]18'04"N 19[grados]32'32"E, Carpinus betulus, Juniperus oxycedrus, limestone rocks, 495 m, 20-IV-2017, A.R. Burgaz (MACB 111325). District of Malesi e Madhe, Boge, SH21 Koplik-Boge, 42[grados]21'46"N 19[grados]37'56"E, Pinus nigra, Juniperus oxycedrus, limestone, 865 m, 20-IV-2017 m, A.R. Burgaz (MACB 111328). TIRANE: Rrogozhine, Grethi i Mesem, 41[grados]05'00"N 19[grados]27'35"E, Pinus pinaster, Juniperus oxycedrus, sandy soil, 1 m, 15-IV-2017, A.R. Burgaz (MACB 110951). VLORE: District of Sarande, Pasqirave, Butrint N. P., SH81, Shengjergj monastery, 39[grados]44'51"N 20[grados]00'24"E, Quercus coccifera, Olea europaea, Phlomis sp., limestone, 60 m, 17-IV-2017, A.R. Burgaz (MACB 110964). District of Sarande, Pasqirave, Butrint N. P., SH81, Shengjergj monastery, 39[grados]48'42"N 20[grados]00'43"E, Quercus coccifera, limestone slopes, 50 m, 17-IV-2017, A.R. Burgaz (MACB 110963). Himare, Ilias, Himare, 40[grados]08'29"N 19[grados]40'11"E, Quercus coccifera, Cupressus sempervirens, Cistaceae shrubs, limestone, 300 m, 16-IV-2017, A.R. Burgaz (MACB 110962). Himare, Gjilek, Rruget te bardha, 40[grados]10'29"N 19[grados]36'09"E, Quercus coccifera, Pistacia lentiscus, limestone, 225 m, 16-IV-2017, A.R. Burgaz (MACB 110960). District of Vlore, Dukat i Ri, SH3 Orikum-Dhermi, Llogora N. P., 40[grados]12'00"N 19[grados]35'26"E, Pinus nigra, Juniperus oxycedrus, Buxus sempervirens, limestone, 1030 m, 16-IV-2017, A.R. Burgaz (MACB 110959). Vlore, Dukat i Ri, SH3 Orikum-Dhermi, Llogora N. P., 40[grados]15'11"N 19[grados]32'14"E, Quercus coccifera, Pinus halepensis, limestone, 320 m, 16-IV-2017, A.R. Burgaz (MACB 110956).

Cladonia rei Schaer.

Homosekikaic acid chemotype: VLORE: Finiq, Muzine, Muzine, 39[grados]54'48"N 20[grados]11'20"E, deciduous Quercus, Erica arborea, limestone, 186 m, 17-IV-2017, A.R. Burgaz (MACB 111198). Homosekikaic and fumarprotocetraric acids chemotype: DURRES: District of Durres, Fushe-Draci, road cape of Rodon, 41[grados]32'50"N 19[grados]31'08"E, Carpinus betulus, Arbutus unedo, Pistacia lentiscus, Cistus salviifolius, sandstone, 90 m, 19-IV-2017, A.R. Burgaz (MACB 111222). KUKES: District of Kukes, Kukes, Rruga Tirana, 42[grados]04'46"N 20[grados]24'45"E, Quercus cerris, deciduous Quercus, limestone, 330 m, 21-IV-2017, A.R. Burgaz (MACB 111336).

Cladonia subulata (L.) F.H. Wigg.

Fumarprotocetraric acid chemotype: KUKES: District of Kukes, Rruga Tirana, 42[grados]04'46"N 20[grados]24'45"E, Quercus cerris, deciduous Quercus, limestone, 30 m, 21-IV-2017, A.R. Burgaz (MACB 111439).

Cladonia symphycarpa (Florke) Fr.

Atranorin and norstictic acid chemotype: KORCE: District of Kolonje (Rrethi i Kolonjes), c. 1.9 km W of village Pepellash, 40[grados]28'56.18"N 20[grados]40'21.27"E, on acid soil in moist road slope, 549 m, 18-X-2011, Z. Barina, L. Lokos, D. Pifko (BP 96584). District of Korce, Dardhe, SH90 Boboshtice-Dardhe, 40[grados]31'18"N 20[grados]49'22"E, Fagus sylvatica, Pinus nigra, Juniperus communis, Betula pendula, Abies sp., limestone, 1390 m, 23-IV-2017, A.R. Burgaz (MACB 111461). District of Korce (Rrethi i Korces), on the ridge of Mts Mali i Moraves above village Dardhe, near peak Buzeja e Boboshtices (1701 m), 40[grados]32'23.97"N 20[grados]49'14.20"E, on acid soil in beech forest on gravel conglomerate, 1550 m, 10-VIII-2010, L. Lokos (BP 96564). ELBASAN: Elbasan county, in open boxwood scrubland above village Kuterman, 41[grados]13'40.99"N 20[grados]14'41.04"E, on acid soil in rocky serpentine area, 510 m, 16-V-2016, L. Lokos (BP 96663). SHKODER: District of Malesi e Madhe, Dedaj, SH21 Koplik-Boge, 42[grados]18'04"N 19[grados]32'32"E, Carpinus betulus, Juniperus oxycedrus, limestone rocks, 495 m, 20-IV-2017, A.R. Burgaz (MACB 111415). District of Malesi e Madhe, Boge, SH21 Kopik-Boge, 42[grados]21'46"N 19[grados]37'56"E, Pinus nigra, Juniperus oxycedrus, limestone, 865 m, 20-IV-2017, A.R. Burgaz (MACB 111416). Malesi e Madhe District (Rrethi i Malesise se Madhe), on Mount Bukovik above village Rrapshe-Stare, 42[grados]23'6.12"N 19[grados]29'25.56"E, on calcareous soil in rocky grasslands, 715 m, 3-V-2015, L. Lokos (BP 96611). Malesi e Madhe District (Rrethi i Malesise se Madhe); Rapshe Gorge between villages Grabom and Lagjja e Re, 42[grados]25'12.46"N 19[grados]30'30.02"E, on calcareous soil in rocky grasslands, 225 m, 3-V-2015, L. Lokos (BP 96612). Shkoder County, at the foot of Mt Maja e Sokolit between villages Tamare and Selce, 42[grados]29'0.61"N 19[grados]35'0.98"E, on calcareous soil in rocky grasslands, 385 m, 8-V-2017, L. Lokos (BP 96685). VLORE: District of Vlore, Dukat i Ri, SH3 Orikum-Dhermi, Llogora N. P., 40[grados]12'00"N 19[grados]35'26"E, Pinus nigra, Juniperus oxycedrus, Buxus sempervirens, limestone, 1030 m, 16-IV-2017, A.R. Burgaz (MACB 111405). Atranorin, norstictic acid and fatty acid chemotype: ELBASAN: Elbasan county, in village Floq, 41[grados]16'1.19"E 20[grados]14'19.17"E, on soil by the roadside, 650 m, 17-V-2016, L. Lokos (BP 96667). Atranorin, norstictic and fumarprotocetraric acids chemotype: ELBASAN: Prrenjas, Urake, SH9 Qafe Thane, 41[grados]04'24"N 20[grados]36'18"E, Quercus macrolepis, deciduous Quercus, Juniperus oxycedrus, limestone, 1000 m, 22-IV-2017, A.R. Burgaz (MACB 111450).

Cladonia sp.

Fumarprotocetraric acid chemotype. SHKODER: Vau i Dejes, SH25 Vau i Dejes-Koman, 42[grados]00'55"N 19[grados]39'37"E, Pinus pinaster, Carpinus orientalis, serpentine soil, 120 m, 20-IV-2017, A.R. Burgaz. VLORE: Finiq, Bistrice, Muzine, 39[grados]55'08"N 20[grados]08'48"E, Quercus coccifera, Carpinus orientalis, Pinus halepensis, Cercis siliquastrum, limestone slopes, 125 m, 17-IV-2017, A.R. Burgaz.

Ana Rosa Burgaz (1), Sandra Luna-Gonzalez (1), Blanca Gutierrez-Larruga (1), Raquel Pino-Bodas (2), Laszlo Lokos (3) & Edit Farkas (4)

Recibido: 2019 / Aceptado:

(1) Department of Biodiversity, Ecology and Evolution (Botany Unit), Faculty of Biology, Complutense University, E-28040 Madrid, Spain. Corresponding author, e-mail: arburgaz@ucm.es

(2) Comparative Plant and Fungal Biology Department, Royal Botanic Gardens Kew, TW9 3AB, Richmond, Surrey, UK. r.pino@kew.org

(3) Department of Botany, Hungarian Natural History Museum, H-1431 Budapest, Pf. 137, Hungary. lokos.laszlo@nhmus.hu

(4) Institute of Ecology and Botany, MTA Centre for Ecological Research, Hungarian Academy of Sciences, H-2163 Vacratot, Alkotmany u. 2-4, Hungary. farkas.edit@okologia.mta.hu

http://dx.doi.org/10.5209/BOCM.63970
Table 1. Main secondary metabolites of the Cladonia species chemically
analysed from Albania. ATR: atranorin, BAR: barbatic acid, BOU:
bourgeanic acid, CRY: cryptochlorophaeic acid, FUM: fumarprotocetraric
acid, HSEK: homosekikaic acid, MER: merochlorophaeic acid, NST:
norstictic acid, PHY: physodalic acid, PSO: psoromic acid, RAN:
rangiformic acid, THAM: thamnolic acid, USN: usnic acid, ZEO: zeorin.

      Species         ATR  BAR  BOU  CRY  FUM  HSEK  MER  NST  PHY  PSO

C. borealis                +
C. caespiticia                            +
C. cariosa 1          +
C. cariosa 2          +                                   +
C. cariosa 3          +
C. cariosa 4          +
C. cervicornis                            +
C. chlorophaea                            +                    +
C. coccifera
C. coniocraea                             +
C. conista                      +         +
C. cryptochlorophaea                 +
C. cyathomorpha                           +
C. fimbriata                              +
C. foliacea 1                             +
C. foliacea 2                             +                    +
C. foliacea 3                             +                         +
C. furcata 1                              +
C. furcata 2                              +                    +
C. furcata 3          +                   +
C. furcata 4          +                   +                    +
C. furcata 5          +         +         +
C. humilis            +                   +
C. macilenta               +
C. merochlorophaea                        +          +
C. parasitica              +
C. peziziformis                           +
C. polycarpoides 1                                        +
C. polycarpoides 2    +                                   +
C. pyxidata 1                             +
C. pyxidata 2                             +                    +
C. pyxidata 3                             +                        +
C. ramulosa                               +
C. rangiformis 1      +                        +
C. rangiformis 2      +                   +    +
C. rangiformis 3      +                   +                    +
C. rei 1
C. rei 2                                  +
C. subulata                               +
C. symphycarpa 1      +                                   +
C. symphycarpa 2      +                   +               +
C. symphycarpa 2      +                   +               +

      Species         RAN  THAM  USN  ZEO

C. borealis
C. caespiticia
C. cariosa 1
C. cariosa 2
C. cariosa 3          +
C. cariosa 4          +               +
C. cervicornis
C. chlorophaea
C. coccifera                     +    +
C. coniocraea
C. conista
C. cryptochlorophaea
C. cyathomorpha
C. fimbriata
C. foliacea 1                    +
C. foliacea 2                    +
C. foliacea 3                    +
C. furcata 1
C. furcata 2
C. furcata 3
C. furcata 4
C. furcata 5
C. humilis
C. macilenta               +
C. merochlorophaea
C. parasitica              +
C. peziziformis
C. polycarpoides 1
C. polycarpoides 2
C. pyxidata 1
C. pyxidata 2
C. pyxidata 3
C. ramulosa
C. rangiformis 1      +
C. rangiformis 2      +
C. rangiformis 3      +
C. rei 1
C. rei 2
C. subulata
C. symphycarpa 1
C. symphycarpa 2
C. symphycarpa 2
COPYRIGHT 2019 Universidad Complutense de Madrid
No portion of this article can be reproduced without the express written permission from the copyright holder.
Copyright 2019 Gale, Cengage Learning. All rights reserved.

Article Details
Printer friendly Cite/link Email Feedback
Author:Burgaz, Ana Rosa; Luna-Gonzalez, Sandra; Gutierrez-Larruga, Blanca; Pino-Bodas, Raquel; Lokos, Laszl
Publication:Botanica Complutensis
Date:Jan 1, 2019
Words:16301
Previous Article:Bibliografia Botanica Iberica, 2017. Bryophyta.
Topics:

Terms of use | Privacy policy | Copyright © 2020 Farlex, Inc. | Feedback | For webmasters