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A valid name for the taxon known as Tillandsia bryoides auct. (bromeliaceae)/Un nombre valido para el taxon conocido como Tillandsia bryoides auct. (Bromeliaceae).

Tillandsia bryoides was validly described by J. G. Baker (1878) using A. Grisebach's name which appears on the specimen label of Lorentz 128. Four more additional vouchers appear in a separate paragraph following the original description: Gillies s.n. "On trees and bushes at the foot of the Cordillera, near Mendoza" (= T. pedicellata (Mez) A. Cast.; W. Lobb s.n. "South Chili" (= T. tricholepis Baker); Glaziou 3124 "South Brazil" (= T. tricholepis) and Christie s.n. "Parana" (= T. tricholepis), all of them housed in K except for Lorentz 128 which Baker had seen in BM (Till, pers. comm.). One year later, Grisebach (1879) modified the description of T. bryoides based on material from "near Cordoba", "near Tucuman" ( = Lorentz 73), and "near Jujuy", but describing longer leaves (8,5-10 mm vs 6,3-8,5 mm), shorter bracts (4,2-5,3 mm vs. 6,3-8,5 mm) and dismissing the possibility of 1-flowered inflorescences (23-flowered vs. 1-3-flowered). Nomenclatorically he created a younger and invalid homonym to Baker's name. About half a century later, Smith (1935) amended the description of T. bryoides because it "was described from a mixed sheet and the original description unfortunately contained both elements. One element had already been described as T. tricholepis, so that the description is amended here to fit the previously undescribed second element. The only definite correction is that the spikes never have more than a single flower instead of: 'Flowers 1-3' ". Based on Smith's (1935) statements and the data published in his monograph (Smith & Downs, 1977: 879) the original material on which Baker had described T. bryoides was a mixture of individuals of different taxa, and Smith established what part of the original material corresponded to his amended description of T. bryoides, being the lectotype of that name (see McNeill et al., 2006: Article 9.12, Ex. 7), and what part corresponds to T. tricholepis. Smith and Downs (1977) stated for the first time that this lectoptype (as "holotype") is housed in BM with one duplicate ("isotype") in M. These two specimens as well as further duplicates in G, GH, and WU all turned out to be T. tricholepis.

Based on the amendment and commentaries made by Smith (Smith, 1935; Smith & Downs, 1977), it can be inferred that at least part of the voucher P. Lorentz 128 housed in BM included elements of T. bryoides, in agreement with the original description published by Grisebach in Baker (1878) and amended by Smith (1935). But when Till (1989) looked for the lectotype of T. bryoides in BM, he only found half a sheet labelled as P. Lorentz 128 with elements of T. tricholepis. After a recent consultation (John Hunnex, pers. comm.), no other sheet of P. Lorentz 128 could be found in BM. The study of the duplicate kept in M (M-0111455) and another one found in MPU revealed that they both include only elements of T. tricholepis. The herbaria K and particularly CORD (Hunziker, 1960) and GOET, where P. Lorentz deposited his collections, do not possess duplicates of P. Lorentz 128. This search for isotypes of T. bryoides only produced two old specimens, K000321909 and GOET000424, labelled as probable isotypes of T. bryoides, but without information about the collector and locality, and they only include elements of T. tricholepis. All these facts support the inclusion of T. bryoides into the synonymy of T. tricholepis, already suggested by Till (1989). However, there is still a taxon of Tillandsia subgen. Diaphoranthema from central and northwestern Argentina, central Peru, Bolivia, and northwestern Paraguay, with yellow, sessile, solitary flowers and a glabrous floral bract, that has been known as Tillandsia bryoides (cf. Castellanos, 1945; Schinini et al., 2008; Subils, 2009), that Till (1989) distinguished as T. bryoides auct. This paper provides precisely a valid name for that taxon, an illustration, and a discussion of the relationships with its morphologically closest species.

Tillandsia minutiflora Donadio, sp. nov. TYPE: Argentina, San Juan, Depto. Valle Fertil, San Agustin de Valle Fertil, ladera del cerro, frente camping ACA, 850 m s.m., sobre Acacia, 14-XII1987, M. E. Mulgura, C. Antuna & E. A. Ulibarri 704 (holotype, SI). Fig. 1.

Tillandsiae tricholepidi similis, sed caulibus brevioribus, 1,5-3,5(-6) cm longis, foliis 3-nervatis, angustioribus, 0,5 mm latis; pedunculis semper reductis et 1-floris et bractea florali glabra, 1nervia, mediae sepalorum aequans bene differt.

Epiphytic, perennial, tiny herbs, with adhesive, reduced roots. Stems short, 1.5-3.5(-6) x 0.3 cm (leaves included). Leaves densely imbricated, spiral, appressed, erect, 5.5-6 x 0.5-0.7 mm; sheath thin, translucent, 2 x 1.7 mm, 3-nerved, blade shortly subulate, 3.5-4 x 0.5-0.7 mm. Inflorescence reduced to 1 flower partially hidden among the apical leaves. Flowering peduncle very short or reduced, which does not elongate by the fruit ripening. Peduncle bracts 2, similar to the leaves (in shape), but 1-nerved (exceptionally 3-nerved) and without foliose apex, 2.5-3.5 x 1.2-1.4 mm. Floral bract subulate, thin, glabrous (exceptionally with 2 or 3 apical trichomes), 1-nerved, half the length of the sepals, 2.5-3 x 1.5 mm. Flowers minute, 6.5 x 1.5 mm (calix tube diameter). Sepals lanceolate, (4-)5(-7)-nerved, 4.8-5.5 x 1.2 mm densely lepidote on the distal half. Petals linearoblong, 5.5-6.5 x 1 mm, yellow. Stamens 2.6 mm, deeply included, filament 1.5 mm, anthers at the level of the stigma, 1.1 mm. Gynoecium 2 mm, ovary as long as the style, stigma subcapitate. Ovules numerous. Capsule cylindrical, oblong, 20 x 0.5-1 mm, valves straight. Seeds many, comose.

Distribution and habitat. Tillandsia minutiflora grows from central Peru (Till, 1989, sub T. bryoides auct.) through Bolivia up to central Argentina and northwestern Paraguay, with a disjunct area in southern Buenos Aires (Sierra de la Ventana). Its geographical distribution includes areas with altitudes between 600 and 2600 m (Till, 1989, sub T. bryoides auct.). Tillandsia minutiflora is an epiphytic species usually growing on bushes and small trees in "del Monte" phytogeographical province, but it can also colonize rocks.


Observations. Tillandsia minutiflora is morphologically close to T. tricholepis, and differs by the shorter, slender stems, and narrower leaves with 3-nerved sheath, 1-nerved bracts (both peduncle and flower), inflorescence always 1flowered, with a very reduced peduncle which does not elongate with fruit ripening, and a glabrous floral bract, shorter than the sepals. After the study of numerous vouchers, and cultivation of abundant material of Tillandsia tricholepis, there is a strong morphological support to consider T. pedicellata as a valid independent taxon from T. tricholepis because the former has deep purple flowers, sepals glabrous and 3nerved, bracts 1-nerved, and leaves with 3nerved sheath. To clarify the limits among Tillandsia tricholepis, T. pedicellata, and the new species T. minutiflora, an additional comparative table is here presented (Table 1).

Additional material examined

Tillandsia minutiflora sp. nov. (paratypes)

ARGENTINA. Buenos Aires. Depto. Tornquist, Sierra de la Ventana, Hicken s.n. (SI-31944). Catamarca. Depto. Ambato, Rodeo, Quebrada del Nogal Marcado, 15-II-1959, Carenzo 896 (LIL). Depto. Santa Maria, 8 km. from Santa Maria towards Tafi del Valle, 23-II-1994, Leuenberger 4326 (CORD); La Hoyada, 25-I-1908, sine colector s.n. (SI-31951). Chaco. Depto. General Guemes, a 22 km del cruce de la ruta Juana Azurduy en direccion N hacia Nueva Pompeya, 16-XI-1990, Fortunato 1368 (SI). Cordoba. Sine data, Kurtz 11990 (SI). Depto. Calamuchita, Valle de los Reartes, Castellanos 46 (SI). Depto. Colon, Jesus Maria, eastern side of the Sierra Chica, 11 km W of Ascochinga along the street to La Cumbre below Las Tres Cascadas, 04-II-1990, Till 5014 (SI). Depto. Ischilin, unos 10 km E de Villa Quilino, rumbo a San Pedro Norte, 10-XI-1972, Hunziker 22017 (CORD); Los Tartagos, 31-III-1944, O'Donell 667 (LIL). Depto. Punilla, Cerca de la orilla del Lago San Roque, 09-IX-1945, Hunziker 6079 (CORD); Sierra Chica, Falda W, Cavalango, 25-VI-1950, Hunziker 8410 (CORD); En quebrada vecina al Lago San Roque, 02-IX-1945, Hunziker 6065 (LIL). Depto. San Alberto, Villa Cura Brochero, along the street from Mina Clavero to the Cumbre de Achala, 13 km above Nina Paula towards Piedra La Tortuga, 4 km before Pje. Rio los Sauces, 16-II-1990, Till 5220 (SI). Depto. San Javier, Yacanto, 1925, Vattuone 1 (SI). Depto. Totoral, Cerro de Totoral, General Mitre, 26-II1936, Giardelli 285 (SI). La Rioja. Depto. General Juan Facundo Quiroga, Malanzan, subiendo al cerro al E del cerro "El Elefante", 12-I-2009, Donadio 185 (SI). Depto. San Blas de los Sauces, Quebrada de los Sauces, frente de la casa OchoaHuasi, IX-1931, Hicken s.n. (SI-31958). Salta. Depto. Anta, J. V. Gonzalez, Campos del Norte, 07-II-1987, Palaci 882 (MCNS). Depto. Cafayate, Cerro Amarillo, en granito al E de Cafayate, 02 VI-1987, Palaci 954 (MCNS). Depto. Capital, Parque San Martin, 28-XI-1985, Palaci 256 (MCNS). Depto. Rivadavia, Mision La Paz, sobre Rio Pilcomayo, 01-I-2003, Scarpa 536 (SI); Riva davia banda sur, Canada Honda, 28-III-1986, Palaci 551 (MCNS). San Juan. Depto. Valle Fertil, alrededores del dique del Valle Fertil, 21-IV1995, Biurrun 4114 (SI); San Agustin de Valle Fertil, 23-XI-1986, Haene 468 (SI); San Agustin de Valle Fertil, 20-I-1989, Haene 788 (SI). Santiago del Estero. Depto. Guasayan, de Santa Catalina a Guampacha, 14-IV-1979, Rotman 207 (SI). Tucuman. Depto. Tafi del Valle, Rincon de Quilmes, Valles Calchaquies, 17-II-1996, Isasmendi s.n. (LIL-600299). Depto. Trancas,Vipos, 08-XII-1921 (fl), Schreiter 2029 (LIL); Vipos, 03 XII-1922 (fl), Venturi 1978 (SI).

BOLIVIA. Chuquisaca. Prov. Oropeza, entre Sucre y La Palca, II-1979, Ceballos 352 (SI). Tarija. Prov. Cercado, Barrancas, 09-XI-1977, Coro 258/77 (LIL).

PARAGUAY. Alto Paraguay. Depto. Mayor Pablo Lagerenza, Mayor Pablo Lagerenza, 10-IV 1978, Schinini 15008 (SI).

Tillandsia pedicellata (Mez) A. Cast.

ARGENTINA. Buenos Aires. Depto. Tornquist, El Coral, 20-III-1948, Fabris 27 (SI). Catamarca. Depto. Ambato, Ruta Prov. 1, de Buena Vista a Singuil, 02-XI-2008, Zuloaga 10580 (SI). Depto. Andalgala, Cuesta de la Chilca, 10-XI2008, Slanis 4911-2008 (SI). Depto. Tinogasta, La Mesada 73,5 km N of Fiambala on Ruta Provincial 43, 14-XII-1998, Leuenberger 4695 (CORD). Cordoba. Sine data, II/1925, Lossen 189 (Si). Depto. Capital, Achala, Sierra Grande, 28-I-1920, Castellanos s.n. (SI-31959). Depto. Punilla, San Esteban, 09-I-1938, Nicora 1697 (SI). Depto. San Alberto, Villa Cura Brochero, along the street from Mina Clavero to the Cumbre de Achala, 13 km above Nina Paula towards Piedra La Tortuga, 4 km before Pje. Rio Los Sauces, 16-II-1990, Till 5217 (SI). Depto. San Javier, Yacanto, 1925, Vattuone 1a (SI). Jujuy. Depto. Humahuaca, N de Humahuaca, 03-I-1992, Kiesling 8129 (SI). Depto. Tilcara, Maimara, Ruta 9, 30-III-1993, Deginani 320 (SI); Pucara, 10-IV-1986, Palaci 545 (MCNS); Tilcara, 20-IX-1985, Palaci 188 (MCNS); Tilcara, Cerros a 1 km al W de la Ruta No. 9, 28-XII-1989, Novara 9226 (MCNS). Depto. Tumbaya, El Volcan, 10-VII-1922, Castellanos s.n. (SI-31955); Quebrada de Humahuaca, Laguna de Volcan, 07-XI-1974, Subils 2030 (CORD). La Pampa. Depto. Lihuel Calel, Sierra de Lihuel Calel, Cerro "de la Sociedad", ladera N, 16-X 1979, de Azkue s.n. (SI-75440). La Rioja. Depto. Capital, Estribacion oriental de la Sierra de Velasco, La Lancha, camino a El Cantadero, 04-III 1944, Hunziker 5031 (CORD); Pelagio B. Luna, Sierra Velazco, rancho La Esperanza., 15-IV-1951, Sparre 8689 (LIL). Depto. Coronel Felipe Varela, Cuesta de Miranda, Ruta Nac. 40, de Villa Union a Chilecito, 09-I-2009, Donadio 79 (SI); Rio Miranda, 26-II-1941, Burkart 12278 (SI); Ruta Nacional 40, de Guandacol a Chilecito, Cuesta de Miranda, 12-II-2011, Zuloaga 12817 (SI). Depto. General Belgrano, Cumbre de la Sierra de Olta, 08-II-1940, Castellanos 33503 (LIL). Depto. General Juan Facundo Quiroga, Northern foot of the Sierra de Porongo, ca. 4 km E of Malanzan, 13-II 1990, Till 5181 (SI); Ruta Prov. 28, desde El Portezuelo hacia Malanzan, 12-I-2009, Donadio 176 (SI). Mendoza. Sine data, Carette s.n. (SI31946). Depto. Lujan de Cuyo, Crucecita, XII1906, Carette s.n. (SI-31947); Lunlunta, 24-II1936, Ruiz Leal 3820 (LIL); Potrerillos, XI-1912, Sanzin 30 (SI). Depto. San Rafael, 31 km W de 25 de Mayo, 03-III-1989, Leuenberger 3958 (SI, B). Rio Negro. Dpto. San Antonio, Sierra Grande about 100 km S of San Antonio, 20-XII-1938, Eyerdam 23534 (SI). Salta. Depto. Cachi, Recta de Tintin, 05-IV-1987, Palaci 925 (MCNS). Depto. Cafayate, Cerro Amarillo, al E de Cafayate, 02-IV-1987, Palaci 955 (MCNS). Depto. Capital, Quebrada de San Lorenzo, 20-IX-1972, Subils 1450 (CORD). Depto. La Poma, Ruta Nac. 40, El Cajon, Zuloaga 11243 (SI). Depto. Los Andes, San Antonio de los Cobres, 18-II-1986, Palaci 387 (MCNS). Depto. Molinos, Tacuil, 02-IV-1988, Palaci 1112 (MCNS); Tacuil, 02-IV-1988, Palaci 1113 (MCNS). Depto. Rosario de Lerma, a 3 km de el Golgota, rumbo a Chorrillos, Quebrada del Toro, 11-III-1984, Toledo 947 (MCNS); Santa Rosa de Tastil, Quebrada del Toro, km 97, 04-I 1990, Palaci 1129 (MCNS). San Juan. Depto. Angaco, Sierra de Pie de Palo, subiendo por el camino al Mogote Los Corralitos, en la Quebrada del Molle, 28-XI-1980, Hunziker 23694 (CORD). Depto. Jachal, La Cienaga. Ruta Provincial 491, de Rodeo hacia Huaco, 06-I-2009, Donadio 52 (SI). Depto. Valle Fertil, alrededores del dique del Valle Fertil, 21-IV-1995, Biurrun 4114 b (SI); Astica, 14-XI-1986, Haene 175 (SI); Cerro Morado, 28-V-1959, Cuezzo 2966 (LIL); Laderas de cerros Sa. de Elizondo, 16-XII-1987, Mulgura 801 (SI). Depto. Zonda, El Palque, O de Pachaco, 14I-987, Kiesling 6767 (SI).

BOLIVIA. La Paz. Prov. Murillo, Hacienda Huajchilla, 18 km SE of La Paz (La Florida), along the Rio La Paz, 07-IX-1986, Solomon 15577 (SI).

Tillandsia tricholepis Baker

ARGENTINA. Catamarca. Depto. Ancasti, Alrededores del dique de Ipizca, 17-XI-1984, Subils 3682 (CORD). Depto. Paclin, El Salton, en el Arroyito de los Ovejeros, frente al Lago Sumampa (sobre el camino entre Ruta 67 y La Vina), 20-IX-1970, Hunziker 20665 (CORD); 11 km from La Merced on road over the Cuesta del Totoral towards La Vina, on the La Vina side of pass, 24-II-1994, Leuenberger 4341 (CORD). Chaco. Sine data, Meyer 2 (SI). Depto. San Fernando, Isla Soto, 18-XII-1974, Burkart 30778 (SI). Cordoba. Sine data, 1919, Castellanos 670 (SI). Depto. Colon, Jesus Maria, en muros de la Iglesia Colonial, 21-VII-1942, Burkart 12916 (SI); Jesus Maria, muro de la Iglesia San Isidro, II 1936, Giardelli 284 (SI); La Calera, El Diquecito, 12-X-1960, Ariza Espinar 981 (CORD). Depto. Rio Primero, Laguna de los Luduena, entre Rio Primero y Santa Rosa, 20-IV-1985, Subils 3806 (CORD). Depto. Rio Seco, Ruta Provincial 10, Sebastian Elcano, 20-X-1984, Subils 3313 (CORD). Corrientes. Depto. Capital, Corrientes, II-XI-1934, Burkart 6897 (SI). Depto. Mburu cuya, Lomas de Vallejo, 06-X-1954, Burkart 19312 (SI). Formosa. Sine data, XII-1918, Jorgensen 2800 (SI). Jujuy. Depto. Dr. Manuel Belgrano, Abra de Las Lajitas, 14-II-1995, Deginani 811 (SI); Camino al Cucho, Rio Los Blancos, 19 III-1979, Cabrera 30212 (SI). Depto. Ledesma, Camino de Fraile pintado a El Aibal, 12-XI-1992, Kiesling 8178 (SI); Ledesma, 23-II-1940, Burkart 11431 (SI); "Picada al Arroyo Negrito" Puesto, Parque Nacional Calilegua, 19-VIII-1985, Iudica 36 (SI). Depto. San Pedro, Moralito, 12-VII-1922, Castellanos s.n. (SI-32228); Ruta Prov. 56, 2 km de la Villa Turistica de Palpala camino a La Mendieta, 13-XII-1998, Morrone 3318 (SI). Depto. Santa Barbara, alrededores de Finca La Campana, 12-XII-1998, Ahumada 8953 (SI); de Santa Clara a Abra de los Morteros, 21-XI-1980, Cabrera 32194 (SI). Misiones. Depto. General Manuel Belgrano, Deseado, Chacra de Carlos Macoviak, antigua Ruta 19, ca. 2000 m de la Ruta 101, 16-XII-1997, Mulgura de Romero 1917 (SI). Salta. Depto. Anta, Coronel Olleros, 21-IX-1957, Meyer 21177 (LIL); Parque Nacional El Rey, 10 VII-1979, Chalukian 871 (MCNS); Selva basal, 10-VI-1981, Chalukian 1405 (MCNS); Embalse, 25-I-1979, Rumiz 489 b (MCNS); Ruta provincial no. 20: Parque Nacional El Rey, proximidades de Finca del Rey, 25-IX-1972, Subils 1459 (CORD). Depto. Capital. Ciudad de Salta, 18-V-2003, Novara 11964 (MCNS). Depto. Guachipas, Alemania, camino a Cafayate, km 80, 07-I-1990, Palaci 1138 (MCNS). Depto. La Caldera, Mojotoro, 17-XI-1942, Burkart 13012 (SI). Depto. La Vina, Dique Puerta de Diaz, 5 km al oeste de Coronel Moldes, 04-XII-1983, Novara 4149 (CORD). Depto. Oran, Oran viejo, 16-I-1957, Burkart 20310 (SI); Pichanal, 14-XI-1913, Rodriguez 1119 (SI); Ruta Prov. 132, 29 km de la Ruta Nac. 50 camino a Santa Maria, pasando 6 km la Finca Toto, 16-XII-1998, Morrone 3435 (SI); Ruta 34, km 1290, 3 km. al S del arroyo de los Monos, 11-V-1989, Novara 8893 (MCNS); Urundel, 13-II-1944, Soriano 718 (SI). Depto. Rivadavia, Rivadavia banda sur, Canada Honda, 28-III 1986, Palaci 550 (MCNS). Depto. Rosario de la Frontera, La Hollada, 06-XI-1985, Palaci 201 (MCNS); Termas VI-1931, Hicken s.n. (SI 32243). San Juan. Depto. Capital, Capital, III1931, Correa s.n. (SI-32240). Santa Fe. Depto. Castellanos, Colonia Margarita, I-1905, Wolfhugel s.n. (SI-32230). Santiago del Estero. Depto. Choya, Sierra de Guasayan, Quebrada de Moqui jata, 12-X-1989, Ulibarri 1690 (SI). Depto. Guasayan, de Santa Catalina a Guampacha, 14-IV1979, Rotman 207 a (SI); Sierra de Guasayan, Quebrada de Conzo, 21-III-1989, Ulibarri 1648 (SI); Tucuman. Depto. Capital, El Duraznito, 06 I-1924, Venturi 2799 (SI). Depto. Leales, Chanar Pozo, XII-1919, Venturi 738 (SI). Depto. Lules, 15-VII-1911, Lizer 20 (SI). Depto. Trancas, de Ruta Nac. 9 a Ticucho, 2 km de Ruta 9, 30-X 2008, Zuloaga 10475 (SI).

BOLIVIA. Santa Cruz. Prov. Florida, easternmost rocky foothills of the Andes, above narrowest point at La Angostura on highway from Santa Cruz to Samaipata, 9-I-1998, Nee 47868-a (SI). Prov. Vallegrande, Vallegrande, Estrada Mataral Vallegrande, 6 km S de Mataral, 10-XII-2002, Forzza 2319 (SI); Tarija. Prov. Gran Chaco, 30 km N de Yacuiba, Campo de la Tapia, cerca de Caiza, 2 km E de la ruta a Villa Montes, 7-IV-1977, Krapovickas 30975 (SI).

BRASIL. Sine colector s.n. (SI-32224). Mato Grosso do Sul, Munic. Corumba, 18-III-1906,

Etchichury 52 (SI).

Original recibido el 3 de mayo de 2011, aceptado el 28 de junio de 2011.


To Daniela Zappi (K), Hans-Joachim Esser (M), John Hunnex and Steve Cafferty (BM), Elena Reiner Grehwald (GOET), Luis Ariza Espinar (CORD) for their help in the search for duplicates of P. Lorentz 128.

To Raul Pozner, Osvaldo Morrone and Fernando Zuloaga for the nomenclatural advice. To Francisco Rojas and Roberto Kiesling for the figure. To Rosa Subils and Walter Till for the critical review of the manuscript. To my brother Roberto N. Donadio for his help. In loving memory of my beloved mother Silvina Ester Estrada and Osvaldo Morrone a great botanist and dear fellow from the Darwinion.


Baker, J. G. 1878. A synopsis of the species of Diaphoranthema. Journal of Botany 16: 236-241.

Castellanos, A. 1945. Bromeliaceae, en H. R. Descole (ed.), Genera et species plantarum argentinarum 3: 105-378. Guillermo Kraft Ltda., Buenos Aires.

Grisebach, A. 1879. Symbolae ad Floram argentinam. Abhand lungen der koniglichen Gesellschaft der Wissenschaften zu Gottingen 24: 334.

Hunziker, A. T. 1960. Catalogo de los tipos "Grisebachianos" conservados en Cordoba. Boletin de la Academia Nacional de Ciencias 41: 283-421.

McNeill, F.; R. Barrie, H. M. Burdet, V. Demoulin, D. L. Hawkswoth, K. Marhold, D. H. Nicolson, J. Prado, P. C. Silva, J. E. Skog, J. H. Wiersema, N. J. Turland. 2006. International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botanical Congress, Vienna, Austria, July 2005. A. R. G. Gantner Verlag, Ruggell [Regnum Veg. 146].

Schinini, A.; M. G. L. Wanderley, T. Strehl, Z. Martins & B. Moreira. 2008. Bromeliaceae, en F. O. Zuloaga, O. Morrone & M. J. Belgrano (eds.), Catalogo de las Plantas Vasculares del Cono sur (Argentina, sur de Brasil, Chile, Paraguay y Uruguay). Monographs in Systematic Botany from the Missouri Botanical Garden 107: 245-291.

Smith, L. B. 1935. Studies in the Bromeliaceae VI. Proceedings of the American Academy of Arts and Sciences (Contributions from the Gray herbarium of Harvard 106) 70: 153-224.

Smith, L. B. & R. J. Downs. 1977. Tillandsia, en C. T. Rogerson (ed.), Flora Neotropica, monograph 14, part 2, pp. 879880. New York: Hafner Press.

Subils, R. 2009. Bromeliaceae, en R. B. Kiesling (ed.), Flora de San Juan, vol. 4, pp. 340-363. Mendoza: Universidad Nacional de San Juan.

Till, W. 1989. Die Untergattung Diaphoranthema (Beer) C. Koch von Tillandsia Linnaeus. I, Das Tillandsia loliacea Aggregat. Die Bromelie 3: 55-59.

Sabina Donadio Instituto de Botanica Darwinion (CONICET-ANCEFN), Labarden 200, Casilla de Correo 22, B1642HYD San Isidro, Buenos Aires, Argentina;
Table 1. Morphological differences among Tillandsia tricholepis Baker,
T. minutiflora Donadio and T. pedicellata (Mez) Castellanos.

Character                               T. tricholepis

Stems, length                       Long (4-)7-11(-15) cm.

Branches, diameter (leaves          4-6 mm

Leaves (fresh)                      6-9 x 1.5 mm

Leaf blade, position                Appressed, straight (erect),
                                    or patent straight to slightly

Sheath, nervation                   5-nerved

Inflorescence, number of flowers    Few, (1-)2-5(-7)

Inflorescence, peduncle at          Developed

Inflorescence, peduncle at          Developed to reduced
fruit ripening

Bracts (peduncle and                6-7-nerved
flowers), nervation

Bracts (peduncle)                   Clearly distinct from the
differentiation respect to the      leaves (in shape) with
leaves                              foliose apex becoming
                                    shorter towards the distal
                                    end of the peduncle

Bracts (peduncle), number           2-6

Bracts (peduncle at fruit           Dispersed along
ripening), arrangement

Bracts (peduncle),                  Densely lepidote all over
pubescence                          the surface

Bract (flower), relative length     As long as the sepals

Bract (flower), apex                Without foliose apex

Bract (flower), pubescence          Densely lepidote all over
                                    the surface

Sepals, nervation                   7-nerved

Sepals, pubescence                  Loosely lepidote at
                                    the apex

Flower, colour                      Yellow

Character                           T. minutiflora

Stems, length                       Short 1.5-3.5(-6) cm.

Branches, diameter (leaves          3 mm

Leaves (fresh)                      5.5-6 x 0.5-0.7 mm

Leaf blade, position                Appressed, straight (erect)

Sheath, nervation                   3-nerved

Inflorescence, number of flowers    1-flowered

Inflorescence, peduncle at          Reduced

Inflorescence, peduncle at          Reduced
fruit ripening

Bracts (peduncle and                1-nerved
flowers), nervation

Bracts (peduncle)                   Similar to the leaves (in
differentiation respect to the      shape), but 1-nerved and
leaves                              without foliose apex

Bracts (peduncle), number           2

Bracts (peduncle at fruit           Does not correspond
ripening), arrangement

Bracts (peduncle),                  Loosely lepidote at the
pubescence                          margin and apex

Bract (flower), relative length     1/2 the sepal length

Bract (flower), apex                Without foliose apex

Bract (flower), pubescence          Glabrous

Sepals, nervation                   (4-)5(-7)-nerved

Sepals, pubescence                  Densely lepidote in
                                    the distal half

Flower, colour                      Yellow

Character                           T. pedicellata

Stems, length                       Short 1.5-3.5(-6) cm.

Branches, diameter (leaves          5 mm

Leaves (fresh)                      7.5 x 0.8 mm

Leaf blade, position                Appressed, straight (erect)

Sheath, nervation                   3-nerved

Inflorescence, number of flowers    1-flowered

Inflorescence, peduncle at          Reduced

Inflorescence, peduncle at          Developed (sometimes
fruit ripening                      reduced)

Bracts (peduncle and                1-nerved
flowers), nervation

Bracts (peduncle)                   Similar to the leaves (in
differentiation respect to the      shape), but 1-nerved and without
leaves                              foliose apex, becoming shorter
                                    towards the distal end of the

Bracts (peduncle), number           2

Bracts (peduncle at fruit           Only at the base (when the
ripening), arrangement              peduncle develops)

Bracts (peduncle),                  Densely lepidote at
pubescence                          the apex

Bract (flower), relative length     As long as the sepals

Bract (flower), apex                With foliose apex

Bract (flower), pubescence          Densely lepidote at
                                    the apex

Sepals, nervation                   1-3-nerved

Sepals, pubescence                  Glabrous

Flower, colour                      Deep purple
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Title Annotation:Articulo en ingles
Author:Donadio, Sabina
Date:Dec 1, 2011
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