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Floristic records in the Platte and Loup River bottomlands of Platte County, Nebraska.


A recent inventory of vascular plants in the Loup and Platte riverbottoms in Platte County has greatly increased knowledge of the area's flora. Of the 542 species of vascular plants now known to grow in the county, 289 were discovered after 1977, with more than 100 of those after 1990. 425 native and 117 naturalized species are known so far. Nearly 700 native and naturalized alien species are expected to occur in the county, based upon totals from nearby counties. Thus, almost 150 years after European settlement, more than 20% of the species remain unverified; of those, many are undoubtedly nonnative. The riverbottoms of the county have the bestpreserved native flora and are today much richer in species than the uplands, which are more heavily impacted by agriculture.

Ecology and Background

The area of union of the Loup River system with the Platte River basin near Columbus, Nebraska, provides a collective view of the plants found in both river systems. A comprehensive floristic determination for this area could be valuable for evaluation of future land-use impacts in those large watersheds. Growing threats of climate changes could have significant effects on local plant communities. Establishing accurate lists of species is important for detecting signs of change in our local flora.

With ninety-nine percent of the original tall-grass, mixed-grass and wetland prairies plowed for agriculture or used for grazing, populations of many once-common native species are now drastically reduced. Agriculture and urbanization have significantly altered the landscape for many species. As land-use disturbances continue at a rapid rate, bottomland floras are now more diverse, heterogeneous mixes of plants than in presettlement times. The absence of frequent prairie fires and the invasion of many exotics threaten remaining fragments of native prairie and original bottomland forests.

History of Plant Collecting in Nebraska and Platte County

Published reports of Nebraska's flora begin with explorers Lewis and Clark in 1803-6, Nuttall and Bradbury in 1808, James of the Long Expedition in 1820, the Fremont Expedition in the 1840s, and the Warren Expedition of 1857. Among the early attempts to list Nebraska's complete flora are two catalogues, those of Samuel Aughey (1876) and H. J. Webber (1890). Local studies in the state have produced numerous floristic lists for counties and other areas. A detailed list of such publications from 1804 through 1985 is presented by Kantak and Churchill (1986); others are cited by Kaul et al. (2006). The state's vegetation was mapped at 1:1,000,000 by Kaul and Rolfsmeier (1993), and some detail was shown for Platte County. Satellite imagery is depicted at 30-meter resolution by the Nebraska Gap-Analysis Project (1993).

The earliest plant collections in Platte County were by Edwin James of the Long Expedition to the Rocky Mountains in 1820, who traveled the Platte Valley across Nebraska and made many original documentations of Nebraska plant life (Goodman and Lawson 1995). The plants James documented in what is now Platte County are Lithospermum incisum, fringed puccoon; Cirsium sp., a thistle; Vicia americana, American purple vetch (not on our list); Rhus glabra, smooth sumac; Asclepias speciosa, showy milkweed; Toxicodendron radicans, poison ivy; Apocynum cannabinum, hemp dogbane; Helianthus petiolaris, plains sunflower; Amorpha fruticosa, wildindigo; Gaura coccinea, scarlet gaura; Monarda pectinata, a mint not on our list; Potamogeton nodosus, floating-leaf pondweed; Rubus occidentalis, black raspberry; Sparganium eurycarpum, bur-reed; Callirhoe involucrata, purple poppy-mallow; Plantago patagonica, woolly plantain; and Anemone canadensis, meadow anemone.

The junction of the Loup and Platte rivers at Columbus was a landmark for Pawnee Indians in the 1820s (Ducey 2000). Warren (1875) noted that the valley of the Loup Fork was well wooded as far up as the Pawnee villages to the west of present-day Platte County. The Platte River at Columbus was described as having a sand bottom and many sandbars, and the floodplain was said to be from two to ten miles wide, as it is today. In 1844, Carleton reached the fork of the Loup and Platte Rivers and remarked:

The bed of the river is but one wide expanse of quicksand, which is formed in bars and these are continually changing and driving about. The channels are innumerable, but are usually only a foot or so deeper than the surrounding water. The river is filled with beautiful islands. They are all well wooded, but only here and there is there any timber growing upon the main banks. Sometimes we found the channels between the islands and the shore, entirely dry, presenting to the eye a wide extent of sand, which as the wind swept over it, was blown about in clouds, as one would notice on a barren coast of the ocean. The bottomlands are what would be called high river prairie (Carleton 1844-1846 (1983)).

Curry (1950) wrote that when the city of Columbus was founded, prior to 1870, the grass "blue joint" grew thick and matted, was taller than a man's head and common in the area. It was reported to grow between twelve and fifteen feet tall along the sloughs in the area and defied any attempts by primitive plows. (The plant called blue-joint today is Calamagrostis canadensis, which never grows that tall, and perhaps Curry was describing Phragmites australis, common reed, which reaches such height in moist habitats. Calamagrostis canadensis is not on our list of vouchered species, but it is known from several adjacent counties and is undoubtedly in Platte County today.)

The Loup River drains much of the Sandhills, which are less disturbed than most areas of the state, but the substitution of cattle for bison has altered grazing patterns and affected native flora there. The Platte River originates in Colorado and has numerous diversions, with significant irrigation usage and storage along the entire watercourse. Row-crop agriculture dominates its borderlands.


Our 12-year sampling period, 1996-2007, documents species within the Platte and Loup River bottoms after 150 years of settlement by Euro-Americans in these watersheds. For completeness and comparison, we list all species ever collected in Platte County, as documented by specimens deposited in various university herbaria, especially that of the University of Nebraska-Lincoln, which houses many specimens we collected. The list was assembled from Atlas of the Flora of the Great Plains (Great Plains Flora Association 1977), The Flora of Nebraska (Kaul et al. 2006), and our own observations and collecting. It consists almost entirely of plants collected in the bottomlands of Platte County, because very little collecting has been done in the much-disturbed, heavily agricultural uplands north of those bottomlands, most of whose species also occur in the bottomlands. This congregation of plants of river bottoms appears to be common in the lower reaches of the Platte River.



The focus for observations and collection were two primary sites on the Platte and Loup Rivers, where two of the largest watersheds in Nebraska unite (Fig. 1). The Platte River site is Witchey's Island (Fig. 2), a heavily wooded area (but not now a true island) on the north bank of the Platte River, equidistant between Duncan and Columbus on a parcel of property almost a mile in length. Witchey's Island has been grazed only minimally by livestock and natural deer populations, has not had fire for over 100 years, and has never been cultivated or farmed. It was homesteaded in 1864 by John Witchey, and according to historical record he maintained only a small garden and orchard. The Loup River site (Fig. 2) is on the south bank of the river and includes dense bottomland forest with open prairies, all on sandy soil.

The coordinates for most plants collected are as follows:

Witchey's Island-Platte River Site (Fig. 2 Top)

NW corner of site

Latitude 41.3809542

Longitude -97.445221

NE corner of site

Latitude 41.3834823

Longitude -97.430938

SW corner of site

Latitude 41.37656772

Longitude -97.444935

SE corner of site

Latitude 41.3779800

Longitude -97.430747

Loup River Site (Fix. 2 Bottom)

NW corner of the site

Latitude 41.4162131

Longitude -97.367379

NE corner of site

Latitude 41.4139801

Longitude -97.354334

SW corner of the site

Latitude 41.4079510

Longitude -97.366522

SE corner of site

Latitude 41.4083231

Longitude -97.354334

Lake Babcock, three miles northwest of Columbus, was another sampling location, as was Buck Island on the Loup River southeast of the city. But most collecting was at the established Loup and Platte River study sites because of their ease of access and the intact nature of their flora. The sampling locations were walked during the spring, summer and fall growing seasons, typically on a weekly basis, to identify species not previously vouchered.


The native and naturalized species known so far are summarized in Table 1 above, and the full list of those species follows the text, as Table 2.

The Witchey's Island site is representative of an eastern Nebraska riparian forest with a mixed-hardwood community, where the tree canopy is Celtis occidentalis, hackberry; Fraxinus pennslyvanica, green ash; Morus alba, white mulberry; Quercus macrocarpa, bur oak; and Ulmus americana, American elm. Cottonwoods, Populus deltoides, are present, but not to the large size of those in the Loup River site. The representative shrubs for this community are Cornus drummondii, rough-leaf dogwood; Ribes missouriense, Missouri gooseberry; Symphoricarpos occidentalis and S. orbiculatus, wolfberry and coralberry; and Zanthoxylum americanum, prickly-ash. Abundant woody vines (lianas) form jungle-like tangles: Parthenocissus quinquefolia, Virginia creeper, and P. vitacea, woodbine; Smilax hispida, green briar; Toxicodendron radicans, poison ivy; Menispermum canadense, moonseed; and Vitis riparia, riverbank grape. The herbaceous plants in this community include Carex spp., sedges; Elymus canadensis, Canada wild rye; Ageratina altissima, white snakeroot; Muhlenbergia spp., muhly grasses; Sanicula canadensis, Canada sanicle; and Viola sororia, sister violet. This community occurs in the floodplains of rivers and streams in the eastern fourth of the state and extends only slightly westward into central Nebraska, along the Loup and Platte River systems. Western wild rose, Rosa woodsii, is more typical of central and western Nebraska, but it is abundant and robust here.

The Loup River site is also representative of eastern-Nebraska riparian woodland, where the tree canopy is represented by cottonwood, many of them 10-30 m in height. The subcanopy contains Acer negundo, boxelder; Gleditsia triacanthos, honey locust; Fraxinus pennslyvanica, green ash; Ulmus americana, American elm; and Morus alba, white mulberry--all common at this site. Shrubs include Cornus drummondii, Ribes missouriense, Sambucus canadensis (elderberry), and Symphoricarpos orbiculatus. Parthenocissus quinquefolia, Toxicodendron radicans and Vitis riparia are also common. Representative herbaceous plants include Ageratina altissima, white snakeroot; Galium aparine, bedstraw: Geum canadense, white avens; Poa pratensis, Kentucky bluegrass; Rudbeckia laciniata, cutleaf coneflower; and Viola sororia, sister violet--all common to abundant.

Recent intrusions of exotic, invasive species are evident: Lythrum salicaria, purple loosestrife; Euphorbia esula, leafy spurge; and introduced genotypes of Phragmites australis, common reed. We witnessed their rapid proliferation, but Tamarix ramosissima (salt-cedar) is not yet known in the study area, although it is rampant to the west, along the Platte. Some species apparently are moving upriver, east to west, such as Robinia pseudoacacia, black locust; Catalpa speciosa, northern catalpa; and Lonicera tatarica, Tatarian honeysuckle. The invasive exotics Amur honeysuckle (Lonicera maackii) and autumn-olive (Elaeagnus umbellata) were not present in or near our study sites, but their currently aggressive westward spread across Nebraska assures eventual arrival in Platte County.

The Loup River site has naturalized species such as Rhamnus cathartica, buckthorn; Convallaria majalis, lily-of-the-valley; and Lonicera tatarica, which are common there but not at the Witchey's Island site. The latter site harbors such natives as Hibiscus laevis, halberd-leaf rosemallow; Mimulus glabratus, roundleaf monkey flower; Quercus macrocarpa, bur oak, and the introduced Lysimachia nummularia, moneywort--all rare or absent at the Loup River site. Prickly-ash, Zanthoxylum americanum, and rough-leaf dogwood, Cornus drummondii, both native to the area, are increasingly abundant in the understory at Witchey's Island, especially in heavily grazed places because livestock do not eat them. Black locust, Robinia pseudoacacia, and northern catalpa, Catalpa speciosa, both native to southeastern North America, are invading the forests near the confluence of the rivers. Siberian elm, Ulmus pumila, and white mulberry, Morus alba, both introduced to North America from Asia long ago, are established and abundant. A native invasive tree, red-cedar (Juniperus virginiana), is abundant in fields and even in deep forests, and here as almost everywhere across the state, it overwhelms native ecosystems.

Some species reach their western limits in or near these sites, e.g., Arisaema triphyllum (jack-in-the-pulpit), Zanthoxylum americanum (prickly-ash), Erythronium albidum (prairie fawnlily), Viola pubescens (smooth yellow violet), and Parthenocissus quinquefolia (Virginia creeper). Others reach their eastern limits in the same area, e.g., Opuntia fragilis, little prickly-pear.


Aughey, Samuel. 1875. Catalogue of the Flora of Nebraska. University of Nebraska-Lincoln: 37 pp.

Carleton, James H. 1844-1846 (1983). The Prairie Logbooks: Dragoon Campaigns to the Pawnee Villages in 1844, and to the Rocky Mountains in 1845. Lincoln, Nebraska: University of Nebraska Press: 295 pp.

Curry M. 1950. The History of Platte County, Nebraska. Murray and Gee, Inc. Culver City, CA: 1011p.

Ducey, J. E. 2000. Birds of the Untamed West. Making History, Omaha, Nebraska: 299 pp.

Goodman G.J., and C.E. Lawson. 1995. Retracing Major Stephen H. Long's 1820 Expedition- The Itinerary and Botany. University of Oklahoma Press, Norman: 366pp.

Great Plains Flora Association. 1977. Atlas of the Flora of the Great Plains. Iowa State University Press, Ames: 777 pp.

Kantak, G.E., and S.P. Churchill. 1986. A bibliography of the ecological and taxonomic literature of the Nebraska vascular plants. Transactions of the Nebraska Academy of Sciences 14: 61-78.

Kaul, R.B., and S.B. Rolfsmeier. 1993. Native Vegetation of Nebraska. Map, 1:1000,000. University of Nebraska-Lincoln, Conservation and Survey Division.

Kaul, R. B., D. M. Sutherland, and S. B. Rolfsmeier. 2006. The Flora of Nebraska. University of Nebraska-Lincoln, Conservation and Survey Division: 968 pp.

Nebraska Gap-Analysis Project. 1993. Land Cover Classification of Nebraska. Map 1:1,500,000. Center for Advanced Land Management Information Technologies, University of Nebraska-Lincoln.

Warren, Gouveneur Kemble. 1875. Preliminary Report of Explorations in Nebraska and Dakota, in the Years 1855-'56-'57. Washington D.C.: U.S. Army Corps of Topographical Engineers.

Webber, Herbert J. 1890. Catalogue of the Flora of Nebraska. Nebraska State Board of Agriculture, Report for 1889: 36-163.

Michael P. Gutzmer

New Century Environmental LLC

Columbus, Nebraska 68601-6335


Robert B. Kaul

University of Nebraska State Museum

Lincoln, Nebraska 68588-0514
Table 1. Species of native and naturalized trees, shrubs,
woody vines, and herbs

 Trees Shrubs Lianas Herbs totals

Native 13 20 7 385 425
Introduced 8 10 1 98 117
totals 21 30 8 483 542

Table 2. Platte County families and species of vascular plants
documented. New records (boldface) are as compared to mapped
records in Atlas of the Flora of the Great Plains (Great Plains
Flora Assn. 1977). Some of these new records are mapped in The
Flora of Nebraska (Kaul et al. 2006), whose nomenclature is used
below. Introduced, naturalized species are indicated by an asterisk
(*). Most of the vouchering specimens are in the Bessey Herbarium
of the University of Nebraska State Museum, Lincoln.

Acer ginnala,# Amur maple*
Acer negundo,# boxelder
Acer saccharinum,# silver maple, soft maple

Viburnum opulus,# highbush-cranberry*

Yucca glauca,# yucca

Sagittaria cuneata,# arrowhead, duck-potato
Sagittaria latifolia,# arrowhead

Amaranthus albus,# pale amaranth
Amaranthus retroflexus, redroot pigweed
Amaranthus tuberculatus,# water-hemp
Froelichia floridana,# snake-cotton
Froelichia gracilis,# slender snake-cotton

Rhus aromatica,# fragrant sumac*
Rhus glabra, smooth sumac
Rhus typhina "Laciniata",# staghorn sumac*
Toxicodendron radicans negundo, poison ivy
Toxicodendron radicans rydbergii,# poison ivy

Berula erecta,# water-parsnip
Cicuta maculata,# common water-hemlock
Conium maculatum,# poison-hemlock*
Osmorhiza longistylis,# long-styled sweet cicely
Polytaenia nuttallii, prairie-parsley
Sanicula canadensis, Canada sanicle
Sanicula odorata,# cluster sanicle
Spermolepis inermis, scaleseed

Apocynum cannabinum,# hemp dogbane
Asclepius arenaria, sand milkweed
Asclepius engelmanniana, slender-leaf milkweed
Asclepius incarnata,# swamp milkweed
Asclepius speciosa, showy milkweed
Asclepius speciosa x syriaca, hybrid milkweed
Asclepius syriaca, common milkweed
Asclepius verticillata, whorled milkweed
Asclepius viridiflora,# green milkweed

Arisamea triphyllum,# jack-in-the-pulpit

Achillea millefolium, yarrow
Ageratina altissima, white snakeroot
Ambrosia artemisiifolia,# common ragweed
Ambrosia psilostachya,# western ragweed
Ambrosia trifida, giant ragweed
Antennaria neglecta, pussytoes
Antennaria parvifolia,# pussytoes
Arctium minus,# burdock*
Artemisia dracunculus,# silky wormwood
Artemisia ludoviciana,# Louisiana sage
Aster novae-angliae, New England aster
Aster oblongifolius, aromatic aster
Aster praealtus neb raskensis,# willowleaf aster
Bidens bipinnatus, Spanish needles
Bidens cernuus, bur-marigold
Bidens comosus, beggarticks
Bidens connatus (both varieties), beggarticks
Bidens frondosus, beggarticks
Bidens trichosperma,# tickseed sunflower
Bidens vulgatus, beggarticks
Brickellia eupatorioides corymbulosa, false boneset
Carduus nutans, musk thistle*
Cirsium altissimum,# tall thistle
Cirsium canescens, Platte thistle
Conyza canadensis, horseweed, marestail
Coreopsis lanceolata,# lanceleaf tickseed*
Coreopsis tinctoria, plains coreopsis
Dyssodia papposa, fetid marigold
Echinacea angustifolia, narrow-leaf purple coneflower
Eclipta prostrata,# yerba de tajo
Erechtites hieraciifolia,# fireweed
Erigeron annuus, annual fleabane
Erigeron philadelphicus, marsh fleabane
Erigeron strigosus, daisy fleabane
Eupatorium altissimum,# tall boneset
Eupatorium maculatum v. bruneri,# spotted Joe Pye weed
Eupatorium perfoliatum,# boneset
Euthamia gymnospermoides, goldentop
Gnaphalium obtusifolium,# fragrant cudweed
Grindelia squarrosa,# curly-cup gumweed
Helenium autumnale,# sneezeweed
Helianthus annuus,# common sunflower
Helianthus grosseserratus, sawtooth sunflower
Helianthus petiolaris,# plains sunflower
Helianthus tuberosus, Jerusalem artichoke
Heliopsis helianthoides,# false sunflower
Heterotheca latifolia,# camphor-weed
Heterotheca villosa, golden-aster
Hymenopappus tenuifolius
Iva annua,# marsh-elder
Lactuca serriola,# prickly lettuce*
Leucanthemum vulgare,# oxeye daisy*
Liatris punctata, gayfeather
Liatris squarrosa glabrata, gayfeather
Matricaria matricarioides, pineapple weed*
Nothocalais cuspidata, false dandelion
Ratibida columnifera, prairie coneflower
Ratibida pinnata,# grayhead coneflower
Rudbeckia hirta,# black-eyed Susan
Rudbeckia laciniata,# cutleaf coneflower
Senecio plattensis, prairie ragwort
Silphium integrifolium v. laeve,# rosinweed
Silphium laciniatum,# compass plant
Silphium perfoliatum,# cup plant
Solidago canadensis, Canada goldenrod
Solidago gigantea,# tall goldenrod
Solidago mollis, soft goldenrod
Solidago nemoralis,# gray goldenrod
Taraxacum officinale, dandelion*
Thelesperma megapotamicum, greenthread
Tragopogon dubius, goat's-beard*
Vernonia baldwinii, interior, western ironweed
Xanthium strumarium,# spiny cocklebur

Impatiens capensis, touch-me-not

Berberis vulgaris, European barberry*

Catalpa speciosa,# northern catalpa*

Hackelia virginiana, stickseed
Lithospermum caroliniense, plains puccoon
Lithospermum incisum, fringed puccoon
Onosmodium molle occidentale,# false gromwell

Arabis hirsuta pycnocarpa,# rock cress
Barbarea vulgaris, winter cress*
Capsella bursa-pastoris, shepherd's-purse*
Cardamine pensylvanica,# bitter cress
Chorispora tenella,# blue mustard*
Descurainia pinnata,# tansy mustard
Descurainia sophia,# tansy mustard*
Erysimum repandum, bushy wallflower*
Hesperis matronalis,# dame's rocket*
Lepidium densiflorum, pepper-grass
Physaria ludoviciana, bladderpod
Rorippa palustris glabra, bog yellow-cress
Rorippa sessiliflora,# yellow-cress
Sisymbrium loeselii, tall hedge-mustard*
Thlaspi arvense, field penny-cress*

Opuntia fragilis,# little prickly-pear
Opuntia humifusa, eastern prickly-pear

Chamaecrista fasciculata, showy partidge-pea
Gleditsia triacanthos,# honey-locust
Gymnocladus dioica,# Kentucky coffee tree

Lobelia siphilitica,# great blue lobelia
Lobelia spicata, palespike lobelia
Triodanis perfoliata, Venus's looking-glass

Cannabis sativa,# marijuana*
Humulus lupulus,# hop

Lonicera japonica,# Japanese honeysuckle*
Lonicera morrowii,# Morrow's honeysuckle*
Lonicera tatarica, Tatarian honeysuckle*
Sambucus canadensis, elderberry
Symphoricarpos occidentalis, wolfberry
Symphoricarpos orbiculatus,# coralberry

Agrostemma githago, corn-cockle*
Arenaria serpyllifolia,# thyme-leaf sandwort*
Cerastium brachypodum,# chickweed
Cerastium fontanum vulgare,# chickweed*
Dianthus armeria,# Deptford pink*
Holosteum umbellatum,# jagged chickweed*
Saponaria officinalis,# soapweed, bouncing Bet*
Stellaria media,# common chickweed*

Celastrus scandens,# American bittersweet
Euonymus atropurpureus,# wahoo

Celtis occidentalis, hackberry

Ceratophyllum demersum,# coontail

Chenopodium glaucum,# oakleaf goosefoot*
Chenopodium simplex,# maple-leaf goosefoot
Chenopodium standleyanum,# Standley's goosefoot
Chenopodium stricture, goosefoot
Corispermum americanum,# American bugseed
Cycloloma atriplicifolium, winged pigweed
Kochia scoparia kochia, summer cypress*
Salsola tragus=S. iberica, Russian thistle*

Lechea mucronata, pinweed

Polanfsia jamesii, James's clammyweed

Hypericum perforatum,# common

Commelina communis,# dayflower*
Tradescantia occidentalis,# spiderwort

Calystegia macounii,# Macoun's bindweed
Calystegia sepium angulata, hedge bindweed
Convolvulus arvensis, field bindweed*
Ipomoea hederacea,# ivyleaf morning-glory*
Ipomoea purpurea,# common morning-glory*

Cornus drummondii, rough-leaf dogwood
Cornus sericea, red-osier dogwood

Penthorum sedoides,# ditch stonecrop

Echinocystis lobata, wild-cucumber
Sicyos angulatus,# bur-cucumber

Juniperus virginiana,# eastern red-cedar

Bolboschoenus fluviatilis,# river bulrush
Carex bebbii,# sedge
Carex blanda,# sedge
Carex brevior, sedge
Carex comosa,# sedge
Carex crawei,# sedge
Carex davisii,# sedge
Carex eleocharis, sedge
Carex emoryi,# sedge
Carex granularis haleana,# sedge
Carex gravida,# sedge
Carex heliophila, sunsedge
Carex hystericina, sedge
Carex laeuiconica, sedge
Carex meadii,# sedge
Carex molesta,# sedge
Carex pellita,# sedge
Ca rex praegracills,# sedge
Carex sartwellii,# sedge
Carex scoparia,# sedge
Carex stipata, sedge
Carex tetanica,# sedge
Carex vulpinoidea,# sedge
Cyperus acuminatus,# flatsedge
Cyperus bipartitus,# brook flatsedge
Cyperus erythrorhizos, redroot flatsedge
Cyperus lupulinus lupulinus,# flatsedge
Cyperus lupulinus x schweinitzii,# flatsedge
Cyperus odoratus,# rusty flatsedge
Cyperus schweinitzii,# Schweinitz's flatsedge
Cyperus squarrosus,# flatsedge
Cyperus strigosus, false nutsedge, umbrella sedge
Eleocharis compressa, spikerush
Eleocharis erythropoda, redstem spikerush
Schoenoplectus pungens, common threesquare
Schoenoplectus tabernaemontani,# softstem bulrush
Scirpus pallidus,# bulrush
Scirpus pendulus,# bulrush

Onoclea sensibilis,# sensitive fern

Elaeagnus angustifolia,# Russian olive*

Equisetum arvense, field horsetail
Equisetum hyemale v. affine, scouring-rush
Equisetum laevigatum,# scouring-rush

Acalypha rhomboidea,# three-seeded mercury
Croton texensis,# Texas croton
Euphorbia davidii, western toothed spurge
Euphorbia esula esula,# leafy spurge*
Euphorbia geyeri, Geyer's spurge
Euphorbia glyptosperma, spurge
Euphorbia hexagona,# six-angle spurge
Euphorbia marginata, snow-on-the-mountain
Euphorbia missurica intermedia, Missouri spurge
Euphorbia nutans, eyebane
Euphorbia stictospora, spurge

Amorpha canescens, leadplant
Amorpha fruticosa, wild-indigo
Apios americana, ground-nut
Astragalus canadensis,# Canadian milkvetch
Astragalus crassicarpus, ground-plum
Crotalaria sagittalis, rattlebox
Dalea leporina, hare's-foot dalea
Dalea purpurea purpurea, purple prairie-clover
Dalea villosa, silky prairie-clover
Desmodium glutinosum,# large-flowered tick-clover
Glycyrrhiza lepidota, wild licorice
Lathyrus polymorphus, hoary vetch
Lespedeza capitata,# round-head lespedeza
Lotus corniculatus,# bird's-foot trefoil*
Lotus purshianus,# prairie trefoil
Medicago lupulina, black medick*
Medicago sativa sativa,# alfalfa*
Melilotus albus,# white sweet-clover*
Melilotus officinalis, yellow sweet-clover*
Oxytropis lambertii, purple locoweed
Pediomelum argophyllum, silver-leaf scurfpea
Pediomelum digitatum, palmleaf scurfpea
Psoralidium lanceolatum, lemon scurfpea
Robinia pseudoacacia,# black-locust*
Strophostyles helvula,# wild bean
Strophostyles leiosperma,# slickseed wild bean
Trifolium hybridum,# Alsike clover*
Trifolium pratense, red clover*
Trifolium repens,# white clover*
Vicia villosa, hairy vetch*

Quercus macrocarpa,# bur oak

Gentiana andrewsii dakotica,# bottle gentian
Gentiana puberulenta, downy gentian

Ribes missouriense,# Missouri gooseberry

Elodea nuttallii, elodea
Naias guadalupensis,# naiad

Ellisia nyctelea, waterpod

Belamcanda chinensis,# blackberry lily*
Iris germanica,# bearded iris*
Iris pseudacorus,# yellow iris, yellow flag*
Sisyrinchium campestre, blue-eyed grass
Sisyrinchium montanum,# blue-eyed grass

Juglans nigra,# black walnut

Juncus arcticus balticus,# Baltic rush
Juncus brachyphyllus,# shortleaf rush
Juncus dudleyi,# Dudley's rush
Juncus interior, inland rush
Juncus torreyi,# Torrey's rush

Hedeoma hispida, rough false pennyroyal
Leonurus cardiaca,# motherwort*
Lycopus americanus, American water-horehound
Lycopus asper,# western water-horehound
Mentha arvensis, field mint
Monarda fistulosa,# wild-bergamot
Monarda punetata oceidentalis,# horsemint*
Nepeta cataria, catnip*
Prunella vulgaris lanceolata,# American heal-all
Prunella vulgaris vulgaris,# European heal-all
Pycnanthemum virginianum, Virginia mountain-mint
Salvia reflexa,# Rocky Mountain salvia
Scutellaria galericulata, marsh skullcap
Scutellaria lateriflora,# mad-dog skullcap
Scutellaria parvula missouriensis, little skullcap
Teucrium canadense canadense, American germander
Teucrium canadense occidentale,# American germander

Lemna spp.,# duckweed
Spirodela polyrrhiza,# greater duckweed
Wolffia columbiana,# Columbian watermeal

Allium canadense canadense,# wild onion
Allium canadense fraseri, wild onion
Allium canadense lavendulare, wild onion
Allium perdulce, fragrant onion
Asparagus officinalis,# asparagus*
Convallaria majalis,# lily-of-the-valley*
Erythronium albidum,# prairie fawnlily
Hernerocallis fulva,# daylily*
Hypoxis hirsuta,# yellow star-grass
Polygonatum biflorum,# Solomon's seal
Smilacina stellata,# starry false Solomon's seal

Linum rigidum simulans, stiff flax
Linum sulcatum,# grooved flax

Ammannia robusta,# toothcup
Lythrum alarum,# winged loosestrife
Lythrum salicaria,# purple loosestrife*

Abutilon theophrasti, velvet leaf*
Callirhoe alcaeoides,# pink poppy-mallow
Callirhoe involucrata, purple poppy-mallow
Hibiscus laevis,# halberd-leaf rose-mallow

Marsilea vestita,# pepperwort, water-clover

Menispermum canadense,# moonseed

Desmanthus illinoensis,# Illinois tick-clover

Mollugo verticillata,# carpet-weed

Morus alba,# white mulberry*

Nelumbo lutea,# American lotus, chinkapin

Mirabilis hirsuta,# hairy four-o'clock
Mirabilis nyctaginea,# wild four-o'clock

Nymphaea odorata s.l., white waterlily

Fraxinus pennsylvanica,# green ash

Calylophus serrulatus, plains evening-primrose
Gaura coccinea,# scarlet gaura
Oenothera biennis,# common evening-primrose
Oenothera rhombipetala,# fourpoint evening-primrose

Cypripedium candidum,# white lady's-slipper

Oxalis dillenii, gray oxalis

Argemone polyanthemos,# prickly poppy

Phryma leptostachya,# lopseed

Plantago lanceolata,# English plantain*
Plantago patagonica patagonica, woolly plantain
Plantago rugelii,# American plantain
Plantago virginica,# plantain

Agropyron cristatum, crested wheatgrass*
Agrostis hyemalis, tickle-grass
Agrostis stolonifera, redtop*
Andropogon gerardii gerardii, big bluestem
Andropogon gerardii hallii, sand bluestem
Aristida basiramea, forktip three-awn
Aristida oligantha, oldfield three-awn
Bouteloua curtipendula, sideoats grama
Bouteloua gracilis, blue grama
Bouteloua hirsuta, hairy grama
Bromus inermis, smooth brome*
Bromus japonicus, hairy chess, Japanese brome*
Bromus pubescens,# Canada brome
Bromus tectorum, cheatgrass*
Buchloe dactyloides,# buffalo grass
Calamovilfa longifolia,# prairie sandreed
Cenchrus longispinus, sandbur
Chloris verticillata, windmill grass
Dactylis glomerata, orchard grass*
Digitaria cognata, fall witchgrass
Digitaria ischaemum, smooth crabgrass*
Digitaria sanguinalis, hairy crabrass*
Echinochloa crusgalli,# barnyard grass*
Echinochloa muricata microstachya,# barnyard grass
Eleusine indica, goosegrass*
Elymus canadensis, Canada wild rye
Elymus elongatus, tall wheatgrass*
Elymus hispidus, intermediate wheatgrass
Elymus repens, quackgrass*
Elymus smithii, western wheatgrass
Elymus trachycaulus trachycaulus, slendeer wheatgrass
Elymus villosus, hairy wild-rye
Elymus virginicus, Virginia wild-rye
Eragrostis cilianensis,# stinkgrass*
Eragrostis pectinacea,# Carolina lovegrass
Eragrostis spectabilis, purple lovegrass
Eragrostis trichodes, sand lovegrass
Eriochloa contracta,# prairie cupgrass
Glyceria grandis, tall manna-grass
Glyceria striata, fowl manna-grass
Hordeum jubatum, foxtail barley
Hordeum pusillum, little barley
Koeleria macrantha, Junegrass
Leersia virginica, Virginian cutgrass
Lolium arundinaceum, tall-fescue
Lolium perenne, perennial ryegrass*
Miscanthus saccharifolius, silver grass, miscanthus*
Muhlenbergia frondosa, wirestem muhly
Muhlenbergia glomerata, muhly
Muhlenbergia mexicana,# wirestem muhly
Muhlenbergia pungens, blowout
Muhlenbergia racemosa, marsh muhly
Panicum acuminatum, panic grass
Panicum capillare, common witchgrass
Panicum dichotomiflorum, fall panicum
Panicum leibergii, Leiberg panicum
Panicum oligosanthes scribnerianum, Scribner's panicum
Panicum virgatum, switchgrass
Paspalum setaceum stramineum, paspalum
Phalaris arundinacea, reed canary-grass*
Phleum pratense, timothy*
Phragmites australis,# common reed
Poa annua,# annual bluegrass*
Poa compressa,# Canada bluegrass
Poa pratensis,# Kentucky bluegrass (*?)
Polypogon monspeliensis,# rabbitfoot grass*
Redfieldia flexuosa,# blowout grass
Schedonnardus paniculatus, tumblegrass
Schizachyrium scoparium, little bluestem
Secale cereale, rye*
Setaria italica, foxtail millet*
Setaria pumila, yellow foxtail*
Setaria verticillata, bristly foxtail*
Setaria viridis, green foxtail*
Sorghastrum nutans,# switchgrass
Sorghum bicolor, sorghum, milo*
Spartina pectinata, prairie cordgrass
Sphenopholis obtusata major, wedgegrass
Sphenopholis obtusata obtusata, wedgegrass
Sporobolus cryptandrus, sand dropseed
Sporobolus vaginiflorus, poverty grass
Stipa spartea, porcupine grass
Tridens flavus,# purpletop grass
Triplasis purpurea, purple sandgrass
Tripsacum dactyloides,# eastern gamagrass
Triticum aestivum, wheat*
Vulpia octoflora, six-weeks fescue
Zizania palustris interior,# interior wild-rice

Phlox paniculata,# summer phlox*

Eriogonum annuum,# annual wild-buckwheat
Polygonum achoreum, knotweed
Polygonum aviculare,# knotweed, wireweed*
Polygonum bicorne, pink smartweed
Polygonum coccineum,# scarlet smartweed
Polygonum convolvulus, black-bindweed*
Polygonum lapathifolium, nodding smartweed
Polygonum pensylvanicum,# Pennsylvania smartweed
Polygonum persicaria,# smartweed*
Polygonum punctatum,# water smartweed
Polygonum ramosissimum (all vars.), bushy knotweed
Polygonum scandens, climbing false-buckwheat
Polygonum tenue,# slender knotweed
Rumex acetosella,# sheep sorrel*
Rumex altissimus, pale dock, tall dock
Rumex patientia orientalis,# patience dock*
Rumex stenophyllus,# narrowleaf dock*

Heteranthera limosa,# mud-plantain

Phemeranthus parviflorus, prairie fame-flower
Phemeranthus rugospermus, sand fame-flower

Potamogeton foliosus,# leafy pondweed
Potamogeton illinoensis,# Illinois pondweed
Potamogeton nodosus, floating-leaf pondweed
Potamogeton pectinatus,# sago pondweed
Potamogeton pusillus,# small pondweed

Androsace occidentalis, western rock-jasmine
Lysimachia nummularia,# moneywort*

Anemone canadensis, meadow anemone
Anemone caroliniana, Carolina anemone
Aquilegia canadensis,# columbine
Delphinium virescens, prairie larkspur
Ranunculus abortivus,# kidney-leaf buttercup
Ranunculus flabellaris,# yellow water-crowfoot
Ranunculus longirostris,# white water-crowfoot
Ranunculus sceleratus, cursed crowfoot
Thalictrum dasycarpum,# purple meadow-rue

Rhamnus cathartica,# buckthorn*

Agrimonia gryposepala,# agrimony
Geum canadense,# white avens
Potentilla recta,# sulfur cinquefoil
Prunus americana,# American plum
Prunus pumila besseyi, sand cherry
Prunus tomentosa,# Nanking cherry, bush cherry*
Prunus virginiana,# chokecherry
Rosa arkansana, dwarf prairie-rose
Rosa multiflora,# multiflora rose*
Rosa woodsii,# western wild rose
Rubus occidentalis,# black raspberry

Galium aparine, cleavers
Galium circaezans,# woods bedstraw
Galium obtusum,# bluntleaf bedstraw

Zanthoxylum americanum,# prickly-ash

Populus alba,# silver poplar*
Populus deltoides occidentalis,# Plains cottonwood
Salix amygdaloides,# peach-leaf willow
Salix eriocephala famelica,# diamond willow
Salix exigua interior, sandbar willow

Comandra umbellata umbellata, comandra, bastard toadflax

Agalinis tenuifolia parvifolia,# gerardia
Bacopa rotundifolia,# water-hyssop
Gratiola neglecta, hedge-hyssop
Leucospora multifida,# leucospora
Lindernia dubia, false pimpernel
Mimulus glabratus,# roundleaf monkey-flower
Mimulus ringens,# Allegheny monkey-flower
Penstemon albidus, white beardtongue
Penstemon angustifolius angustifolius, narrowleaf beardtongue
Penstemon buckleyi,# Buckley's penstemon
Penstemon grandiflorus, large beardtongue
Scrophularia marilandica,# eastern figwort
Verbascum blattaria,# moth mullein*
Verbascum thapsus,# common mullein*
Veronica anagallis-aquatica,# water speedwell*
Veronica catenata,# water speedwell*
Veronica peregrina peregrina,# purslane speedwell*
Veronica peregrina xalapensis, purslane speedwell*
Veronica polita,# veronica*

Ailanthus altissima,# tree-of-heaven*

Smilax hispida,# greenbriar, prickly catbriar

Datura wrightii,# angel's trumpet*
Physalis heterophylla, ivyleaf ground-cherry
Physalis longifolia, common ground-cherry
Solanum carolinense,# horse-nettle
Solanum interius, Plains black nightshade
Solanum ptycanthum, black nightshade
Solanum rostratum, buffalo bur

Sparganium eurycarpum,# bur-reed

Typha angustifolia,# narrowleaf cattail
Typha latifolia,# broadleaf cattail

Ulmus americana,# American elm
Ulmus pumila,# Siberian elm*
Ulmus rubra,# red elm, slippery elm

Boehmeria cylindrica, false nettle
Parietaria pensylvanica,# pellitory
Urtica dioica, stinging nettle

Lippia lanceolata,# northern fogfruit
Verbena bracteata, prostrate vervain
Verbena hastata, common vervain
Verbena stricta, hoary vervain
Verbena urticifolia,# white vervain

Viola pubescens eriocarpa,# smooth yellow violet
Viola sororia, sister violet

Parthenocissus quinquefolia, Virginia creeper
Parthenocissus vitacea,# woodbine
Vitis riparia,# riverbank grape

Zannichellia palustris, horned pondweed

Tribulus terrestris, puncture vine*

Note: New records (boldface) are as compared to mapped
records in Atlas of the Flora of the Great Plains (Great Plains
Flora Assn. 1977) is indicated with #.
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Author:Gutzmer, Michael P.; Kaul, Robert B.
Publication:Transactions of the Nebraska Academy of Sciences
Date:Aug 1, 2008
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