Distribution records of Southern Indiana vascular plants.ABSTRACT. Collections in the Indiana University Southeast History The Indiana University Falls City Area Center was established by Floyd McMurray in 1941 as an extension center of Indiana University in New Albany, Indiana and Jeffersonville, Indiana. Herbarium herbarium, collection of dried and mounted plant specimens used in systematic botany. To preserve their form and color, plants collected in the field are spread flat in sheets of newsprint and dried, usually in a plant press, between blotters or absorbent paper. were reexamined and entered into a database. New and old collections yielded over 100 new vascular plant vascular plant or tracheophyte Any plant that has a specialized conducting system consisting mostly of phloem (food-conducting tissue) and xylem (water-conducting tissue), collectively called vascular tissue. county records for Clark, Crawford, Floyd, and Harrison Counties in southern Indiana Southern Indiana, in the United States, is notable because it is culturally distinct from the rest of the state. The area's geography has led to a blend of Northern and Southern culture that is not found in the rest of Indiana. . Indiana vascular species listed as extirpated (SX), endangered (SE), threatened (ST), or rare (SR) are noted. Others species which may become listed, or are otherwise noteworthy, are briefly discussed. Keywords: Vascular plants (Bot.) plants composed in part of vascular tissue, as all flowering plants and the higher cryptogamous plants, or those of the class See also: Vascular , Indiana, distribution records The Indiana University Southeast Herbarium serves as an educational and community resource as well as a research exchange facility. The herbarium contains vascular plant collections primarily from Clark, Crawford, Floyd, Harrison, and Jefferson Counties, with occasional collections from Washington, Scott, and Perry Counties. This area of southern Indiana includes the lowermost parts of the following regions: the Mitchell Karst Karst (kärst), Ital. Carso, Slovenian Kras, limestone plateau, W Slovenia, N of Istria and extending c.50 mi (80 km) SE from the lower Isonzo (Soča) valley between the Bay of Trieste and the Julian Alps. Plain and Knobstone Escarpment The Knobstone Escarpment is a rugged geologic region in Southern Indiana. Physically, the Knobstone Escarpment is the most rugged terrain in Indiana. The highest hill in the area is Weed Patch Hill, with an elevation of 1,056 feet above sea level. Sections of the Highland Rim The Highland Rim is a geographic term for the area in Tennessee surrounding the Central Basin. Nashville is largely surrounded by higher terrain in all directions. Geologicially, the Central Basin is actually opposite, being a dome. Natural Region, the Scottsburg Lowland and Muscatatuck Flats and Canyon Sections of the Bluegrass bluegrass, any species of the large and widely distributed genus Poa, chiefly range and pasture grasses of economic importance in temperate and cool regions. In general, bluegrasses are perennial with fine-leaved foliage that is bluish green in some species. Natural Region, and the Shawnee Hills The Shawnee Hills is a region of Western Kentucky and Southern Illinois. The Kentucky part consists of many low, rolling hills, primarily underlain by rocks of Pennsylvanian age, including coal. Natural Region (Homoya et al. 1985). The Indiana University Southeast Herbarium is in the process of creating an online database of its vascular plant collection at www.ius.edu/ biology/herbarium/homepage.stm. Ongoing collecting and review of existing collections in the herbarium have yielded over 100 new vascular plant county records since the publication of the last distribution list from the Indiana University Southeast Herbarium (Maxwell & Emmert 1995). Possible state records and noteworthy collections are listed alphabetically by their scientific names and briefly discussed, followed by a listing of state endangered, threatened, and rare species. Table 1 includes distribution records for Clark, Crawford, Floyd, and Harrison Counties with their collection vouchers. Nomenclature conforms primarily to the Field Guide to Indiana Wildflowers (Yatskievych 2000), with authors of plant names, following Brummitt & Powell (1992). Some nomenclature, primarily ferns, grasses, and common names cited, is from Mohlenbrock (1986), Gleason & Cronquist (1991), Homoya (1993) and Yatskievych (2000). Determination of species status follows the Indiana Natural Heritage Data Center Endangered, Threatened and Rare Vascular Plants list, Indiana, from the Department of Natural Resources Many sub-national governments have a Department of Natural Resources or similarly-named organization:
Former and present collectors include Indiana University Southeast students, the authors, several volunteers, Ray Weatherholt and several of his biology students from Floyd Central High School Floyd Central High School is a public high school of the New Albany-Floyd County Consolidated School Corporation and is located in Floyds Knobs, Indiana. History The school was built in 1967 and has had many expansions since then. . METHODS Sites of listed species are reported to Indiana DNR, Nature Preserves. Vouchers are currently held at the Indiana University Southeast Herbarium. Some collections are sent to the DNR, Nature Preserves for confirmation by Michael A. Homoya, DNR Botanist, as well as by George and Kay Yatskievych at the Missouri Botanical Garden The Missouri Botanical Garden is a botanical garden located in St. Louis, Missouri, and is also known informally as "Shaw's Garden" (named for founder Henry Shaw, a botanist and philanthropist). . Methods and criteria for listing Indiana's extirpated, endangered, threatened, and rare vascular plant species have been discussed by Aldrich et al. (1986). POSSIBLE STATE RECORDS AND NOTEWORTHY COLLECTIONS Achyranthes japonica japonica (jəpŏn`əkə): see quince; camellia. (Miq.) Nakai (Amaranthaceae). Japanese Chaff chaff 1. chaffed hay; called also chop. 2. the winnowings from a threshing, consisting of awns, husks, glumes and other relatively indigestible materials. Flower. A native of Asia which was introduced into Kentucky (Gleason & Cronquist 1991). Small, shrubby shrub·by adj. shrub·bi·er, shrub·bi·est 1. Consisting of, planted with, or covered with shrubs. 2. Of or resembling a shrub. plants, common along the upper floodplain floodplain, level land along the course of a river formed by the deposition of sediment during periodic floods. Floodplains contain such features as levees, backswamps, delta plains, and oxbow lakes. slopes and banks of the Ohio "'Banks of the Ohio'" is a nineteenth century murder ballad, in which Willie invites his young lover for a walk during which she rejects his marriage proposal. Once they are alone on the river bank, he murders her. River in the Indiana counties across from Louisville, Kentucky  “Louisville” redirects here. For other uses, see Louisville (disambiguation). . Colonies extend back from the river and down river to the west. We consider "W.E. Thomas 470" from Crawford County Crawford County is the name of eleven counties in the United States:
Egeria densa Planch. (Hydrocharitaceae). Brazilian Water-weed. The state record, "W.E. Thomas 294," was collected in shallow water See:
v. nat·u·ral·ized, nat·u·ral·iz·ing, nat·u·ral·iz·es v.tr. 1. To grant full citizenship to (one of foreign birth). 2. To adopt (something foreign) into general use. in various localities in the United States United States, officially United States of America, republic (2005 est. pop. 295,734,000), 3,539,227 sq mi (9,166,598 sq km), North America. The United States is the world's third largest country in population and the fourth largest country in area. , primarily in the southeast" (Beal & Thieret 1986). Clematis clematis (klĕm`ətĭs, kləmăt`ĭs), any plant of the large genus Clematis (sometimes subdivided into three or four genera), widely distributed herbs or vines of the family Ranunculaceae (buttercup family), many of them terniflora DC. (= C. dioscoreifolia Levi. & Vaniot) (Ranunculaceae). A native of Japan which appears occasionally with other vines along the Ohio River Ohio River Major river, eastern central U.S. Formed by the confluence of the Allegheny and Monongahela rivers, it flows northwest out of Pennsylvania, and west and southwest to form the state boundaries of Ohio–West Virginia, Ohio-Kentucky, Indiana-Kentucky, and . Fatoua villosa (Thunb.) Nakai (Moraceae). Hairy Crabweed, Mulberry Weed. Reported in Flora of North America The Flora of North America (FNA) is a multivolume work describing the native plants of North America. These days much of the Flora is available online. The work is expected to fill 30 volumes when completed. (3):390.1997, but neither the voucher nor the author of the Indiana report was identified (K. Yatskievych pers. commun.). The Floyd County Floyd County is the name of six counties in the United States:
Humulus japonicus Noun 1. Humulus japonicus - ornamental vine native to eastern Asia; cultivated for its variegated foliage Japanese hop hops, hop - twining perennials having cordate leaves and flowers arranged in conelike spikes; the dried flowers of this plant are used in Siebold & Zucc. (Moraceae). Japanese Hops. An introduced native of East Asia. The vine is fairly common along roadsides next to the Ohio River. Lespedeza bicolor Turcz. (Fabaceae). Bicolor bicolor a coat color of two colors. In dogs, usually black with tan markings but may be other combinations such as ticking on a white background. In cats, more than two spots of color on the body, either white and one basic color, or white with one tabby color. Lezpedeza. Two small shrubs were found in the open field northeast of the old black powder plant at Charlestown State Park Charlestown State Park is an Indiana state park on 2,400 acres (0 km) in Clark County, Indiana, in the United States. . Reported for Indiana in the Proceedings of the Indiana Academy of Science, 1953. The state record voucher collected by "Buser & Ahles s.n." in Fountain County is probably in the University of Illinois University of Illinois may refer to:
Liparis loeselii (L.) Rich. (Orchidaceae). Loesel's Twayblade twayblade: see orchid. . Found by Homoya in Jackson County on the Muscatatuck National Wildlife Refuge  National Wildlife Refuge (1990). Thomas discovered a population of about 14 plants
distributed in the lower end of a seep in Charlestown State Park off
Trail 1. This Clark County population represents a further southern
extension as suggested by Homoya (1993).
Melothria pendula L. (Cucurbitaceae). Creeping Cucumber. This vine was considered state extirpated (SX) until three sites were discovered by Thomas in Charlestown State Park in 2000. The vines seemed to be thriving in spite of being in competitive habitats: a roadside ditch, an open disturbed area, and the edge of the escarpment escarpment or scarp, long cliff, bluff, or steep slope, caused usually by geologic faulting (see fault) or by erosion of tilted rock layers. An example of a fault scarp is the north face of the San Jacinto Mts. in California. over Fourteen Mile Creek Fourteen Mile Creek, also referred to as Fourteenmile Creek, is a creek in Clark County, Indiana, close to Charlestown, Indiana. It is so named because it is fourteen miles away from the Falls of the Ohio; similar to how Eighteen Mile Island, Twelve Mile Island, and Six . Since 2000 the Creeping Cucumber has been found in numerous sites throughout Charlestown State Park. It has also been found in roadside ditches in Floyd County along the lower edges of the knobs beside the Ohio River floodplain. Mentha x rotundifolia (L.) Huds. (Lamiaceae). Apple Mint. A possible hybrid for southern Indiana. Several other collections have been made of this mint (Yatskievych 2000). Its occurrence in the state may be more common than the current reports indicate. Orthodon dianthera (Buch.-Ham.) Hand. Mazz. (= Mosla dianthera (Buch.-Ham. ex Roxb.) Maxim) (Lamiaceae). A native of East Asia, established in moist places in the southeast (Gleason & Cronquist 1991). Recorded for Indiana in Kartesz & Meachan's 1999 Synthesis without a reference (K. Yatskievych pers. commun.). Homoya was aware of a site in the Harrison-Crawford State Forest and made a collection 26 September 1997 in Harrison County. The Homoya collection is the state record and "W.E. Thomas 36," a second collection. Thomas reports the population expanding in open areas left by logging at two different sites in Harrison-Crawford State Forest. This may be another species moving up from Kentucky and becoming common in southern Indiana. Packera anonyma (A.W. Wood) W.A. Weber & A. Love (= Senecio Senecio a widespread genus of the Asteraceae family. The genus contains more than 1200 species of which at least 25 are known to be poisonous. Some of them are listed here; the toxins are a group of pyrrolizidine alkaloids which cause seneciosis hepatic injury, and the dummy anonymous A. Wood, S. smallii Britt.) (Asteraceae). Occasional on thin soil over a limestone ledge in Charlestown State Park. Perhaps a recent range expansion phenomenon (Homoya & Hedge 1990). Indiana Watch List (Yatskievych 2000). Papaver dubium L. (Papaveraceae). Garden Poppy. The collection "Maxwell 3166," is the second report of the Garden Poppy along a different railroad track in Clark County. About 15 plants were noted along the tracks about 2 1/2 km. north of highway I-265 along State Road 62 in 1998, but this second colony did not persist probably because of herbicide herbicide (hr`bəsīd'), chemical compound that kills plants or inhibits their normal growth. A herbicide in a particular formulation and application can be described as selective or nonselective. spraying along the right-of-way. The collection "W.E. Thomas 271," along State Road 31 north of Speed, indicates the initial Clark County colony is persisting and perhaps expanding in the much wider strip between State Road 31 and the railroad track. Its occurrence in the state may be more common than reports indicate. Platanthera lacera (Michx.) G. Don. (Orchidaceae). Green Fringed Orchid. This orchid is the most common Platanthera in Indiana (Homoya 1993), but is primarily found in the northern part of the state. A single plant was discovered by Thomas in the edge of a wet depression off Jersey Avenue east of the current Charlestown State Park boundary in an area expected to become annexed to the park. A photo voucher was taken 6 July 2002. This report indicates a range extension to the south as predicted by Homoya (1993). Indiana Watch List (Yatskievych 2000). Pycnanthemum muticum (Michx.) Pers. (Lamiaceae). Mountain Mint. The state record collector, K.F. Duffy, reports colonies in full sun, in poor clay soil next to a pond in a Harrison County hardwood forest. Deam (1940) excludes the species, citing reports, but no specimens. Schneck's report (Deam 1940) for the Lower Wabash Valley is probably correct. Mohlenbrock & Ladd (1978) show distribution in Wabash County, Illinois Wabash County is a county located in the U.S. state of Illinois. As of 2000, the population is 12,937. Its county seat is Mt. Carmel6. Geography According to the U.S. Census Bureau, the county has a total area of 590 km² (228 mi²). , next to the Wabash River. Duffy sent a collection, "K.F. Duffy s.n., 21 September 2002," to Kay Yatskievych at the Missouri Botanical Garden for verification; and this becomes the Indiana voucher specimen for the species and the county record. Our specimen, "K.F. Duffy 1," is a later collection from the same population. Selaginella eclipes Buck (Sellaginellaceae). Small Spikemoss Noun 1. spikemoss - any of numerous fern allies of the genus Selaginella little club moss, spike moss club moss, club-moss, lycopod - primitive evergreen moss-like plant with spores in club-shaped strobiles . First thought to be S. apoda (L.) Fern.(SE), later determined by Homoya as S. eclipes, and also confirmed by G. Yatskievych. Occasional to locally common in Charlestown State Park. Solanum Solanum a widespread plant genus of the family Solanaceae which contains a number of valuable crop plants but also some poisonous ones. Poisoning may be due to (1) the presence in the plant of toxic glycoalkaloids which cause diarrhea, (2) alkamines, e.g. sarachoides Sendter (Solanaceae). Hairy Nightshade nightshade, common name for the Solanaceae, a family of herbs, shrubs, and a few trees of warm regions, chiefly tropical America. Many are climbing or creeping types, and rank-smelling foliage is typical of many species. . A Gastony collection, "Gastony s.n., 17 July 1985, Fee Lane and Highway 46 Bypass, Monroe County, Bloomington, was discovered in the Indiana University Herbarium. This may be the state record. The Gastony collection ... "was apparently an escape to a roadside that was some distance from the experimental field" (K. Yatskievych pers. commun.). The experimental field refers to where Dr. Charles B. Heiser, Jr., of Indiana University carried out some of his well-known hybridization hybridization /hy·brid·iza·tion/ (hi?brid-i-za´shun) 1. crossbreeding; the act or process of producing hybrids. 2. molecular hybridization 3. work with sunflowers, nightshades and domestic plants. Heiser (1969) refers to using Solanum sarachoides in his crosses to rediscover the "wonderberry." The "Maxwell 3367, 25 August 2000," collection from Charlestown State Park would be a second report. This introduced weed was found in a disturbed area of the park and is persisting. However, it does not show the invasive capacity of other introduced species in the park. INDIANA STATE ENDANGERED (SE) Asplenium bradleyi D.C. Eaton (Aspleniaceae). Bradley's Spleenwort. Noted in Dubois County, 23 October 1984, by Homoya & Abrell (1986). "W.E. Thomas 159" was collected from a population of 13-15 plants scattered on several sandstone boulders in an open area near Wyandotte Cave in the Harrison-Crawford State Forest. Gentiana villosa L. (Gentianaceae). Striped Gentian. Deam (1940) reports two collections from southeast Harrison County in black and white oak woods. A Harrison County site was located during a DNR conducted survey for the Indiana Natural Heritage Program. A fragment, "W.E. Thomas 37," was collected in the Harrison County part of the Harrison-Crawford State Forest in a cedar woods along Cold Friday Road. Thomas reports a scattered population of about 30 40 plants in the woods. These sites are in Harrison County, but we cannot be sure they are the same. INDIANA STATE THREATENED (ST) Acalypha deamii (Weath.) Ahles. (= A. virginica L. var. deamii Weath.) (Euphorbiaceae). Mercury. Michael Homoya identified this species in October 1994, in the Jenny Lind Run area of the Indiana Army Ammunition Plant The Indiana Army Ammunition Plant was an Army manufacturing plant built in 1941 between Charlestown and Jeffersonville, Indiana. It consisted of three areas within two separate but attached manufacturing plants:
Diodia virginiana L. (Rubiaceae). Buttonweed. Collected from large colonies along the Ohio River in Floyd and Crawford Counties, and also found along the wet perimeter of Deam Lake in Clark County. See Homoya et al. (1995) for additional locations. Now believed common along the Ohio River. INDIANA STATE RARE (SR) Gentiana flavida A. Gray (G. alba Muhl.) (Gentianaceae). Yellowish Gentian gentian (jĕn`shən), common name for some members of the Gentianaceae, a family of widely distributed herbs, chiefly perennial and fall blooming. , Pale Gentian. An Indiana University Southeast Herbarium collection, "C.W. Henson 43, 19 Sept. 1975," from the Harrison-Crawford State Forest in a dry, oak woods clearing, was determined Gentiana villosa. Reexamination re·ex·am·ine also re-ex·am·ine tr.v. re·ex·am·ined, re·ex·am·in·ing, re·ex·am·ines 1. To examine again or anew; review. 2. Law To question (a witness) again after cross-examination. , as suggested by Homoya, showed it to be Gentiana flavida. The exact county distribution is unknown. Northoscordum bivalve bivalve, aquatic mollusk of the class Pelecypoda ("hatchet-foot") or Bivalvia, with a laterally compressed body and a shell consisting of two valves, or movable pieces, hinged by an elastic ligament. (L.) Britton (Liliaceae). Crow Poison crow poison see amianthium muscaetoxicum. . A colony of perhaps 60 plants was found on a low rock bluff along State Road 62 within Harrison-Crawford State Forest, west of Wyandotte Lake. Oxalis oxalis (ŏk`səlĭs) or wood sorrel, any species of the plant genus Oxalis. Most of the cultivated kinds are tropical herbs used as window plants. illinoensis Schwegman (Oxalidaceae). Illinois Woodsorrel. Two large Clark County populations were pointed out by Homoya about 1994 on the wooded floor of Lick Creek Ravine, in what is now Charlestown State Park. At that time, one population contained about 100 individuals and the other 500-1000. Since then several additional clusters have been found in the park along with a population of about 2000 individuals spreading down from the upper bluff on the east side of Lick Creek towards Fourteen Mile Creek. Two colonies, one with perhaps 100 plants, the other smaller with perhaps 20 plants, were found by Thomas in Crawford County. Both these sites were within the Harrison-Crawford State Forest along the upper tributaries of Dry Run Creek. The Illinois Woodsorrel has leaflets shallowly lobed lobed adj. Having a lobe or lobes: lobed leaves. Adj. 1. lobed - having deeply indented margins but with lobes not entirely separate from each other lobate with green margins; the Big Yellow Woodsorrel (Oxalis grandis Small) has leaflets deeply lobed with purplish-brown margins (Yatskievych 2000). Other characteristics are similar except the Illinois Woodsorrel has a small tuberous root (Homoya pers. commun.). We believe the Illinois Woodsorrel is more common than the Big Yellow Woodsorrel in our south central Indiana area. Spiranthes vernalis Engelm. & A. Gray (Orchidaceae). Spring Ladies' Tresses, Grassleaved Ladies' tresses. Observed by Thomas & Howell Curtis, Charlestown State Park in a disturbed area north of Trail 1 parking. "W.E. Thomas 332" serves as a Clark County voucher. Tragia cordata Michx. (Euphorbiaceae). Heart-leaved Noseburn. Homoya identified this vine in the rocky woods, north of the west little bluestem glade in Charlestown State Park.
Table 1.--State and county records. Abbreviations for species status
are: SE = State Endangered, ST
= State Threatened, SR = State Rare. Plant records without a number
are indicated by s.n. = (sine numero)
without a number, followed by the collection date.
Species Family County, collector and number
Acalypha gracilens A. Euphorbiaceae Clark, Maxwell 3351
Gray
Acalypha deamii (Weath.) Euphorbiaceae Harrison, W.E. Thomas 38
Ahles, (ST)
Achyrantes japonica Amaranthaceae Clark, W.E. Thomas 511;
(Miq.) Nakai Crawford W.E. Thomas 470;
Floyd, W.E. Thomas 536
Aesculus Pavia L. Hippocastanaceae Floyd, Maxwell 3075
Albizia julibrissin Mimosaceae Floyd, Boha & Chanley 8
Duraz.
Alisma subcordatum Raf. Alismaceae Harrison, R. Schoen s.n.,
21 July 1972
Amaranthus retroflexus L. Amaranthaceae Floyd, Weatherholt 1526
Ambrosia trifida L. Asteraceae Harrison, R. Schoen 59
Ammania robusta Heer & Lythraceae Floyd, W.E. Thomas 28
Regel
Amorpha fruticosa L. Fabaceae Floyd, Schoen s.n.,
2 June 1972
Amsonia tabernaemontana Apocynaceae Harrison, R. Schoen s.n.,
Walter 11 May 1972
Aquilegia canadensis L. Ranunculaceae Floyd, Weatherholt 880
Artmisia annua L. Asteraceae Clark, W.E. Thomas 30;
Floyd,
W.E.
Thomas 635
Artemisia ludoviciana Asteraceae Harrison, R. Schoen s.n.,
24
Nutt. August
1972
Artemisia vulgaris L. Asteraceae Floyd, Maxwell 1280
Asclepias purpurascens L. Asclepiadaceae Floyd, S. Newman 113
Asplenium bradleyi D.C. Aspleniaceae Crawford, W.E. Thomas 159
Eaton (SE)
Avena Sativa L. Poaceae Floyd, Weatherholt 1561
Buglossoides arvense Boraginaceae Floyd, Maxwell 3484
(L.) I.M. Johnst.
(= Lithospermum arvense
L.)
Capsella bursa-pastoris Brassicaceae Floyd, J.A. Kruer Jr. 90
(L.) Medik.
Cardamine hirsuta L. Brassicaceae Floyd, Maxwell 2543
Cardamine angustata O.E. Brassicaceae Harrison, W.E. Thomas 157
Schultz (=
Dentaria heterophylaa
Nutt.)
Clematis terniflora D.C. Ranunculaceae Floyd, W.E. Thomas 17
Diodia virginiana L. Rubiaceae Floyd, W.E. Thomas 1; Clark
(ST) W.E.
Thomas 335; Crawford, W.E.
Thomas 435
Draba brachycarpa Nutt. Brassicaceae Clark, W.E. Thomas 156
Gray
Egeria densa Planch. Hydrocharitaceae Crawford, W.E. Thomas 294
Enemion biternatum Raf. Ranunculaceae Floyd, Weatherholt 799
(= Isopyrum
biternatum (Raf.) Torr.
& Gray)
Equisetum arvense L. Equisetaceae Floyd, Weatherhold 845
Erianthus alopecuroides Poaceae Harrison, R. Schoen 47
(L.) Ell.
Erythronium albidum Nutt. Liliaceae Floyd, Weatherholt 794
Erythronium americanum Liliaceae Floyd, Gohmann &
Weatherholt
Ker Gawl. 32
Fatoua villosa (Thunb.) Moraceae Floyd, Maxwell s.n.,
Nakai 9/20/1995
Gymnocladus dioica (L.) Caesalpiniaceae Floyd, N.A. Myers 11
K. Koch.
Holosteum umbellatum L. Caryophyllaceae Clark, Maxwell & Thomas
3197
Houstonia pusilla Rubiaceae Clark, W.E. Thomas 62;
Schoepf Floyd, Maxwell
3483; Harrison, W.E. Thomas
58
Humulus japonicus Moraceae Clark, Reported by the Adams
Siebold & Zucc. (1993);
Crawford, W.E. Thomas 23;
Floyd,
W.E. Thomas 285
Hypochaeris radicata L. Asteraceae Clark, W.E. Thomas 340
Iva annua L. (= I. Asteraceae Clark, W.E. Thomas 18;
ciliata Willd. in Crawford,
Deam (1940) W.E. Thomas 125; Harrison,
W.E.
Thomas 495
Kickxia elatine (L.) Scropulariaceae Clark, Gilbert 105
Dumort
Koelreuteria paniculata Sapindaceae Floyd, F. Bierman 119
Laxm.
Table 1.--Continued.
Species Family County, collector and
number
Lespedeza bicolor Turcz. Fabaceae Clark, Maxwell 3373
Linaria vulgaris Mill. Scropulariaceae Clark, K.D. Bledsoe 80
Lindernia dubia (L.) Scropulariaceae Floyd, W. E. Thomas 291;
Pennell var. Harrison,
anagallidea (Michx.) W. E. Thomas 295
Cooperr.
Lindernia dubia (L.) Scropulariaceae Crawford, W. E. Thomas
Pennell var. dubia 24; Harrison,
W.E. Thomas 131
Linum usitatissimum L. Linaceae Floyd, Maxwell 3183
Liparis loeselii (L.) Orchidaceae Clark, Maxwell & Thomas
Rich. 3498
Melothria pendula L. Cucurbitaceae Clark, Maxwell 3355; Floyd,
Maxwell 3347
Mentha x rotundifoia (L.) Lamiaceae Crawford, W.E. Thomas 112;
Huds. Floyd,
W.E. Thomas 371
Muscari botryoides (L.) Liliaceae Harrison, R. Schoen s.n.,
Mill. 28 April 1972
Nasturtium officinale R. Brasiccaceae Harrison, R. Schoen s.n.,
Br. 11 June 1972
Nothoscordum bivale (L.) Liliaceae Crawford, W.E. Thomas 182
Britton (SR)
Oenothera biennis L. Onagraceae Harrison, R. Schoen 36
(= O. pycnocarpa
Atkinson & Bartlett
Oenothera laciniata Hill Onagraceae Floyd, J.A.M.(? Illegible) 8
Oenothera pillosella Raf. Onagraceae Floyd, K. Heavrin 103
Osmorhiza longistylis Apiaceae Harrison, R. Schoen s.n.,
(Torr.) D.C. 15 May 1972
Oxalis illinoensis Oxalidaceae Clark, Maxwell 3432
Schwegman (SR)
Packera anonyma (A.W. Asteraceae Clark, Maxwell & Thomas
Wood) W.A. 3492
Weber &
A. Love
Panicum nitidum Lam. Poaceae Floyd, R. Schoen 14
Pellaea glabella Mett. Adiantaceae Crawford, W.E. Thomas 185
ex Kuhn
Penstemon digitalis Scropulariaceae Floyd, Hobson 96
Nutt. ex Sims
Phleum pratense L. Poaceae Floyd, Barger & Weatherholt
72
Phlox divaricata L. Polemoniaceae Floyd, F. Gohmann 23
Platanthera lacera G. Orchidaceae Clark, Maxwell & Thomas
Don observed
Platanus occidentalis L. Platanaceae Clark, Maxwell 1706
Polygonum arenastrum Polygonaceae Harrison, W.E. Thomas 42
Jord. ex Boreau
Polygonum lapathifolium Polygonaceae Floyd, G. Doyle 7
L.
Pyrrhoppapus carolinianus Asteraceae Clark, W.E. Thomas 5;
(Walter) Harrison, W.E. Thomas
D.C. 272
Rananculus repens L. Ranunculaceae Floyd, Weatherholt 1012
Ratibida columnifera Asteraceae Floyd, Maxwell 3337
(Nutt.) Woot. &
Standl.
Robinia hispida L. Fabaceae Clark, S. Newman 61
Rubus occidentalis L. Rosaceae Harrison, Conrad 71
Rumex obtusifolius L. Polygonaceae Clark, Megraw 40
Salix babylonica L. Salicaceae Floyd, Chambers 51
Salix exigua Nutt. Salicaceae Floyd, Bennett 2
(= S. interior
Rowlee)
Selaginella eclipes Buck Selaginellaceae Clark, Maxwell 3420;
Floyd, Maxwell
3476
Sibara virginica (L.) Brassicaceae Clark, Maxwell 3478
Rollins
Sida spinosa L. Malvaceae Harrison, S. Lawton 88
Solanum sarachoides Solanaceae Clark, Maxwell 3367
Sendtner
Spermacoce glabra Michx. Rubiaceae Floyd, W.E. Thomas 3
Spiranthes vernalis Orchidaceae Clark, W.E. Thomas 332
Engelm. & A. Gray
(SE)
Thlaspi arvense L. Brassicaceae Harrison, W. E. Thomas 160
Tragia cordata Michx. Euphorbiaceae Clark, observed by Homoya
(SR)
Typha angustifolia L. Typhaceae Floyd, Weatherholt 1573
Viola lanceolata L. Violaceae Harrison, R. Schoen s.n.,
19 May, 1972
Woodsia obtusa (Spreng.) Aspleniaceae Clark, Maxwell 1534
Torrey
ACKNOWLEDGMENTS We wish to thank Dr. David W. Taylor Rear Admiral David Watson Taylor, USN (4 March 1864 – 28 July 1940) was a naval architect and engineer of the United States Navy. He served during World War I as Chief Constructor of the Navy, and Chief of the Bureau of Construction and Repair. , Curator of the Indiana University Southeast Herbarium, for his cooperation, use of the facilities, and continuing encouragement that collections and natural history have a place in current academia. We thank Michael A. Homoya and others in the Indiana DNR for their continual encouragement and assistance in the search for listed species in southern Indiana, and their concern for the preservation of our natural heritage. Thanks also to Mr. Gerald J. Pagac, Director of the Division of State Parks and Reservoirs, for providing a collection permit to Maxwell for Charlestown State Park. We also greatly appreciate the cooperation of Larry Gray, property .manager of Charlestown Park. We also thank Pete Thorn, property manager of Harrison-Crawford State Forest, for providing a collection permit to William E. Thomas to extend our survey. We also thank Kay and George Yatskievych of the Missouri Botanical Garden for their help in determining and confirming identifications of Indiana flora and sorting out the vouchers. We also thank our many former students for their collections as well as current collectors, students and volunteers. Although many student collections were mandatory and such collecting may now be considered an obsolete teaching method, they form an important segment of the Indiana University Southeast Herbarium. Once again, we thank Ray Weatherholt, long-time biology teacher at Floyd Central High School, now retired, for donating his collection to the herbarium. Thanks also to the reviewers and the editor for their helpful comments in revising our original manuscript. LITERATURE CITED Adams, W.R. & M.R. Adams. 1993. Flora of the Falls of the Ohio State Park Falls of the Ohio State Park is a state park in Indiana. It is located on the banks of the Ohio River at Clarksville, Indiana, across from Louisville, Kentucky. The park is part of the Falls of the Ohio National Wildlife Conservation Area. and National Wildlife Conservation Area (A preliminary report). Unpublished. Aldrich, J.R., J.A. Bacone & M.A. Homoya. 1986. List of extirpated, endangered, threatened and rare vascular plants in Indiana: An update. Proceedings of the Indiana Academy of Science 95: 413-419. Beal, E.O. & J.W. Thieret. 1986. Aquatic and Wetland Plants of Kentucky. Kentucky Nature Preserves Commission, Scientific and Technical Series Number 5. Frankfort, Kentucky. 315 pp. Brummitt, R.K. & C.E. Powell. 1992. Authors of Plant Names. Royal Botanic Gardens Royal Botanic Gardens may refer to:
Deam, C.C. 1940. Flora of Indiana. Department of Conservation, Indianapolis, Indiana. 1236 pp. Gleason, H.A. & A. Cronquist. 1991. Manual of the Vascular Plants of Northeastern United States and Adjacent Canada. 2nd ed. New York New York, state, United States New York, Middle Atlantic state of the United States. It is bordered by Vermont, Massachusetts, Connecticut, and the Atlantic Ocean (E), New Jersey and Pennsylvania (S), Lakes Erie and Ontario and the Canadian province of Botanic Gardens. Bronx, New York. 910 pp. Heiser, C.B. Jr. 1969. Nightshades, The Paradoxical Plants. W.H. Freeman & Company. San Francisco, California “San Francisco” redirects here. For other uses, see San Francisco (disambiguation). The City and County of San Francisco (EN IPA: [sænfrənˈsɪskoʊ] . 200 pp. Homoya, M.A., D.B. Abrell, J.R. Aldrich & T.W. Post. 1985. The natural regions of Indiana. Proceedings of the Indiana Academy of Science 94: 245-268. Homoya, M.A. & D.B. Abrell. 1986. Recent additions to the flora of southern Indiana. Proceedings of the Indiana Academy of Science 95:429-432. Homoya, M.A. & C.L. Hedge. 1990. Additions to the flora of southern Indiana, IV. Proceedings of the Indiana Academy of Science 99:67-71. Homoya, M.A. 1993. Orchids of Indiana. Indiana Academy of Science. Distributed by Indiana University Press Indiana University Press, also known as IU Press, is a publishing house at Indiana University that engages in academic publishing, specializing in the humanities and social sciences. It was founded in 1950. Its headquarters are located in Bloomington, Indiana. , Bloomington and Indianapolis. 276 pp. Homoya, M.A., D.B. Abrell, C.L. Hedge & R.L. Hedge. 1995. Additions to the flora of southern Indiana, V and VI. Proceedings of the Indiana Academy of Science 104:213-221. Kartesz, J.T. & C.A. Meachan. 1999. Synthesis of the North American North American named after North America. North American blastomycosis see North American blastomycosis. North American cattle tick see boophilusannulatus. Flora. Version 1.0 CD. North Carolina Botanical Garden The North Carolina Botanical Garden (about 700 acres, plus 210 acres of nature preserves) is a botanical garden operated by the University of North Carolina at Chapel Hill, Old Mason Farm Road, Chapel Hill, North Carolina. It is open daily without charge. . Chapel Hill. Maxwell, R.H. & Gall A. Emmert. 1995. Southern Indiana plant distribution records and notes on the endangered, threatened, and rare vascular plant species in the proposed state park areas of the Indiana Army Ammunition Plant. Proceedings of the Indiana Academy of Science 104(34):223-232. Mohlenbrock, R.H. 1986. Guide to the Vascular Flora of Illinois. Southern Illinois University Press Southern Illinois University Press (or SIU Press), founded in 1956, is a publisher and part of Southern Illinois University. External link
Mohlenbrock, R.H. & D.M. Ladd. 1978. Distribution of Illinois Vascular Plants. Southern Illinois University Press, Carbondale & Edwardsville, Illinois. 282 pp. Yatskievych, K. 2000. Field Guide to Indiana Wildflowers, Indiana University Press, Bloomington and Indianapolis. 357 pp. Manuscript received 3 December 2002, revised 16 March 2003. Richard H. Maxwell and William E. Thomas: Herbarium, Indiana University Southeast, New Albany, Indiana New Albany (IPA: [nu ˈɑl.bə.ni]) is a city in Floyd County, Indiana, situated along the Ohio River opposite Louisville, Ky. 47150 USA |
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