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An inventory of the herbaceous plants of the twin swamps nature preserve, Posey County, Indiana.



ABSTRACT. Twin Swamps is a 242 ha nature preserve located in Posey County in the extreme southwestern corner of Indiana. This area, containing one of the few remaining swamp cottonwood, bald cypress, and overcup oak swamps in Indiana, is owned and maintained by the Division of Nature Preserves of the Indiana Department of Natural Resources The Indiana Department of Natural Resources is the agency of the U.S. state of Indiana charged with maintaining natural areas such as state parks, state forests, recreation areas, etc.  (IDNR). Twin Swamps was set aside by the state in 1987 to preserve one of Indiana's few remaining bald cypress swamps. This study presents an initial inventory of herbaceous flowering plants round in the preserve. A total of 139 species of plants is listed in this study. The study is also presented on a web site that includes photographs of most species listed. The web site indexes the species by habitat, flowering period, and flower color. The following is the URL URL
 in full Uniform Resource Locator

Address of a resource on the Internet. The resource can be any type of file stored on a server, such as a Web page, a text file, a graphics file, or an application program.
: (www.usi.edu/science/biology/Twin Swamps/Wildflowers-of-Twin-Swamps.htm)

Keywords: Twin Swamps Nature Preserve, Indiana, Posey County, flora, bald cypress

**********

Twin Swamps Nature Preserve is a 242 ha wooded area very close to the confluence of the Wabash and Ohio rivers in the extreme southwest corner of Indiana (Figs. 1, 2). A trail leads visitors to two separate swamp areas, one dominated by bald cypress (Taxodium distichum) and the other dominated by overcup oak (Quercus lyrata).

[FIGURE 1-2 OMITTED]

The Twin Swamps Nature Preserve is owned and maintained by the Indiana Department of Natural Resources. The IDNR set the land aside in 1987 to ensure the survival of bald cypress trees in the state. These trees were once abundant along the river bottoms of southwestern Indiana; but their population has decreased in part due to overgrazing overgrazing

see overstocking.
, logging, farming, and the damming of the Ohio River for navigational purposes (Ridgway 1872; Lindsey 1962; Abrell 1997). The remains of another large stand of Taxodium can be seen at Hovey Lake, which is about 3.2 km southeast of Twin Swamps Nature Preserve. However, the nearby river dam has raised the level of Hovey lake, permanently inundating the Taxodium and threatening their survival.

Twin Swamps represents an ecological region known as the Southern Bottomlands of Indiana (Homoya et al. 1985). This region, which hugs the lower Wabash and Ohio rivers, has one of the state's longest growing periods. Temperatures, rainfall and humidity are high and winters are mild. As a result, Twin Swamps vegetation has a close affinity with the lower Mississippi Valley and Gulf Coast Plain (Homoya et al. 1985; Abrell 1997; Deam 1940). The climatic conditions and the environmental pressures listed above have resulted in many plants located in this area to be considered as endangered or threatened (IDNR 1993).

While several studies have included plants from the southwestern Indiana region and Posey County (Deam 1940; Homoya 1983, 1987; Homoya & Abrell 1986; Homoya & Hedge 1990), no studies have concentrated specifically on the Twin Swamps Nature Preserve. This study consists of an initial inventory of herbaceous plants found in the Twin Swamps Nature Preserve.

METHODS

During the 1997, 1998, and part of the 1999 growing seasons, several forays per week were made into the study area. Collections were preserved following standard protocol (Waiters & Keil 1996). Collections are stored in the University of Southern Indiana The University of Southern Indiana (USI) is a public university in Evansville, Indiana. This publicly-funded institution is rapidly growing and is the fastest growing comprehensive state university in Indiana.  Herbarium herbarium, collection of dried and mounted plant specimens used in systematic botany. To preserve their form and color, plants collected in the field are spread flat in sheets of newsprint and dried, usually in a plant press, between blotters or absorbent paper.  (USIH). Notes on vegetation consisted of estimates of abundance, seasonal presence, and habitat location. For purposes of this study, the following terminology is used to illustrate abundance: sparse--fewer than five plants seen at preserve; infrequent--seen occasionally, but not widespread; frequent--widespread and easily noticed, may be locally abundant in certain habitats; abundant--seen in great numbers, though may be restricted to specific habitats.

RESULTS

The attached list of plants includes 139 species identified for this study representing 115 genera and 53 families. Gleason and Cronquist (1991) was used as the primary taxonomic and nomenclatural reference. Several standard reference books were used as taxonomic references and aids for identification (Britton & Brown 1915; Deam 1940: Gleason & Cronquist 1991; Mohlenbrock 1986; Mohlenbrock & Voigt 1959; Venning 1984; and Wharton 1971, 1973).

Habitat Descriptions.--Twin Swamps Nature Preserve is transected by a trail that passes through a variety of habitats. The trail begins in a parking area and passes through an open field that gradually slopes down to a wooded area. The trail passes one end of the overcup oak slough and over 0.8 km of low-lying, poorly-drained flatwoods, primarily of oak (Quercus spp.) and hickory (Carya spp.) trees. The trail then rises gradually about a meter to a higher area of better-drained soil. The trail then drops sharply to the bald cy press swamp. IDNR has provided a boardwalk that extends into the swamp to a raised platform among the cypress trees. A secondary trail continues through the "high, dry woods" to the overcup oak slough, gradually descending approximately a meter. This trail passes the oak slough and loops back to the main trail. Land elevation in this part of Indiana is approximately 107 m above sea level.

For the purposes of this inventory, five habitats have been designated: field; low, wet woods; low, dry woods; high, dry woods; cypress swamp.

The habitats are distinguished by varying degrees of shade and by the amount of soil drainage. Very subtle changes in elevation seem to have a profound effect on how much moisture the soil contains in different areas of the preserve. Some areas tend to dry out quickly, while others often have standing water or mud that dries slowly.

The plant communities tend to be distinctive in each zone. With a few notable exceptions (such as Oxalis oxalis (ŏk`səlĭs) or wood sorrel, any species of the plant genus Oxalis. Most of the cultivated kinds are tropical herbs used as window plants.  sp. and Toxicodendron radicans), plants in one habitat tend not to be found in the other habitats.

It has been observed that herbaceous plants in the "low, wet woods" tend to grow no more than 1 m high (Phlox phlox, common name for plants of the genus Phlox and for members of the Polemoniaceae, a family of herbs (and some shrubs and vines) found chiefly in the W United States.  glaberrima, smooth phlox, is typical); those in the "low, dry woods" may reach slightly over 1 m (such as Verbesina helianthoides, yellow crownbeard); those in the "high, dry woods" may reach 2 m or more (Phytolacca americana is dominant and this area has been dubbed the "pokeweed pokeweed or pokeberry, tall, bushy perennial herb (Phytolacca americana) native to North America but cultivated and naturalized in Europe.  forest").

This height variation is seasonal. Water levels at the preserve are typically higher in the spring. In the early spring, the only plants in flower at the preserve are found in the high, dry woods and these tend to be no more than 0.3 m tall (e.g., Thalictrum thalictroides, rue anemone anemone (ənĕm`ənē) or windflower, any of the perennial herbs, wild or cultivated, of the genus Anemone of the family Ranunculaceae (buttercup family). ). Toward the end of summer, when the soil has had a chance to dry out, taller plants may be found in the low, wet woods (such as Lobelia lobelia (lōbēl`yə), any plant of the genus Lobelia, annual and perennial herbs of tropical and temperate woodlands and moist places. Most lobelias have blue or purple flowers on a long (1–4 ft/30–122 cm), leafy stem.  cardinalis, cardinal flower, and Vernonia fasciculata, ironweed ironweed

Any of about 500 species of perennial plants constituting the genus Vernonia (family Asteraceae). Small herbaceous (nonwoody) species are found throughout the world; shrubs and trees are found primarily in tropical regions.
).

Field.--(This habitat is generally open and dry, with some low-lying, wet areas.)

The open area between the parking lot and the woods is dominated by Andropogon virginicus (broom sedge), which gives the field a golden tan appearance most of the year. Other notable flowering plants are those which are seen in most fallow fields of southwestern Indiana especially the asters (Aster spp.) and goldenrods (Solidago Solidago

North American plant genus in the family Asteraceae; contain an unidentified toxin. In some outbreaks there is suspicion that the poisoning is caused by a fungus growing on the plant but tests with the plant alone have proved its toxicity.
 spp.) of late summer. Some areas of the field are lower and more poorly-drained than others. These areas have some plants, such as Aselepias incarnata (swamp milkweed milkweed, common name for members of the Asclepiadaceae, a family of mostly perennial herbs and shrubs characterized by milky sap, a tuft of silky hairs attached to the seed (for wind distribution), and (usually) a climbing habit. ) and Lycopus rubellus (stalked water horehound) that do not seem to appear in other areas of the preserve.

A stand of young sweet gum trees (Liquidambar styraciflua) is conspicuous in the middle of the field. At the edge of the parking lot is a large shingle oak tree (Quereus imbricaria). At the parking lot and at the entrance to the woods, black locust trees (Robinia pseudoacacia) are dominant.

The habitats of Twin Swamps Nature Preserve are relatively free in invasive or exotic species, with the notable exception of the open field area. The open field contains several small populations of Alliaria petiolata (garlic mustard) and abundant populations of Lonicera japonica (Japanese honeysuckle) at the edge of the low, wet woods.

Low, wet woods.--(This habitat is well-shaded, poorly drained, and is generally muddy with areas of standing water.)

The environment changes dramatically at the entry way to the woods. Open areas give way to deep shade and the trail, which slopes downward through the field, bottoms out in an area that is usually muddy and often has standing water of approximately 2-10 cm in depth.

Trees are plentiful in this area, despite the water. Carya ovata (shagbark hickory), Ulmus rubra (slippery elm), and Quercus spp. (oaks) are most common. Most of the trees carry Toxicodendron radicans (poison ivy) vines, which thrive throughout the woods.

In early spring (through April), very few plants are be in flower in the low, wet woods. Only three plants have been recorded: Viola papilionacea (common violet), Ranunculus septentrionalis (swamp buttercup buttercup or crowfoot, common name for the Ranunculaceae, a family of chiefly annual or perennial herbs of cool regions of the Northern Hemisphere. , which does well even in standing water), and Cardamine bulbosa (spring cress).

By mid-May, however, spots of color can be found throughout the low, wet area. The purple, white, and yellow flowers of Phlox glaberrima (smooth phlox), Penstemon Penstemon

a North American genus of plants in the family Scrophulariaceae which act as facultative selenium converters; the selenocompounds produced by the plant cause alopecia, lameness, laminitis; called also beard tongue.
 alluviorum (lowland beardstongue), and Krigia biflora (false or dwarf dandelion) are conspicuous (though not abundant). These predominant plants tend to be no more than 1--2 m tall.

In June, the plant inventory includes Hedyotis purpurea var. calycosum (a white flower in the Rubiaceae family, difficult to identify). Wharton (1971) calls it Houstonia lanceolata and Deam (1940) does not separate var. calycosa from H. purpurea. Also notable are Arisaema dracontium (green dragon), Pycnanthemum tenuifolium (slender mountain mint), Potentilla simplex (common cinquefoil cinquefoil (sĭngk`foil) [O.Fr.,=five leaves], name for any plant of the widely distributed genus Potentilla of the family Rosaceae (rose family), chiefly herbs of north temperate and subarctic regions. ), and early specimens of Impatiens impatiens (ĭmpā`shēĕnz'): see jewelweed.
impatiens

Any of about 900 species of herbaceous plants in the genus Impatiens (balsam family), so named because the seedpod bursts when slightly touched. Garden balsam (I.
 capensis (spotted jewelweed jewelweed, common name for the Balsaminaceae, a family of widely distributed annual and perennial herbs. The principal genus is Impatiens, so named because of the sudden bursting of the mature seed capsules when touched. ).

By the end of July, this area supports an abundance of Laportea canadensis (wood nettle) and Boehmeria cylindrica (false nettle). Also flowering at this time is the poisonous Cicuta maculata (water hemlock) and Hackelia virginiana (stickseed stick·seed  
n.
Any of various plants of the genera Hackelia or Lappula, having small barbed fruits that cling to clothing or fur.
). More unusual are a few specimens of Ludwigia alternifolia (seedbox, with its cube-shaped seedpod, that rattles when shaken).

In August, some taller plants flower. Prominent are Mimulus
For the crab genus, see Mimulus (crab).
Mimulus is also an OpenWetWare community for Mimulus biology .
''Monkey-flower and variants redirect here.
 alatus (the southern species of monkey flower; both purple and more uncommonly white-flowered forms), Lobelia cardinalis (red cardinal flower), and Vernonia fasciculata (ironweed). Also notable are Desmodium paniculatum (panicled pan·i·cle  
n.
A branched cluster of flowers in which the branches are racemes.



[Latin p
 tick trefoil trefoil (trē`foil) [O.Fr.,=three-leaf], in botany, name for several plants, chiefly of the pulse family, having trifoliate leaves. Best known of the trefoils is clover. ), Polygonum Polygonum

genus of toxic plants in the family Polygonaceae, called collectively smartweeds. Some cause nitrate-nitrite poisoning, some cause photosensitization; includes P. aviculare (wireweed), P. convolvulus (Fallopia convolvulus), P. esculentum, P.
 hydropiperoides (mild water pep per), and Euphorbia euphorbia (yfôr`bēə): see spurge.  corollata (flowering spurge spurge (spûrj), common name for members of the Euphorbiaceae, a family of herbs, shrubs, and trees of greatly varied structure and almost cosmopolitan distribution, although most species are tropical. , which was seen in 1997 and 1999, but not in 1998).

Low, dry woods.--(This habitat is well-shaded and often wet but dries out more readily than the low, wet woods habitat.)

As the trail rises out of the moist flood-prone portion of the preserve, it branches in two directions. The main trail continues west toward the cypress swamp. A secondary trail heads southwest to skirt the overcup oak slough. This secondary trail rises slightly into an area of somewhat better-drained soil. Though the slope is nearly imperceptible, the plant community changes significantly.

In the early spring (through April), few species bloom in the low, dry area. The only plants noted in this study are Thalictrum thalictroides, Viola eriocarpa (smooth yellow violet), and V. papilionacea.

Through May, however, a number of new species come into bloom. These plants, which tend to reach 0.5-1.5 m in height, include Asclepias purpurascens (purple milkweed), Phryma leptostachya (lopseed), Porteranthus stipulatus (American ipecac or Indian physic), Ruellia strepens (wild petunia petunia, any plant of the genus Petunia, South American herbs of the family Solanaceae (nightshade family). The common garden petunias, planted also in window boxes, are all considered hybrids of white-flowered and violet-flowered species from Argentina. ), Smilacina racemosa (false Solomon's seal false Solomon's seal
n.
Any of several plants of the genus Smilacina, native to North America and Asia and having a plumelike cluster of small greenish-white flowers with a persistent perianth. Also called Solomon's plume.
), and Verbesina helianthoides (yellow crownbeard). Also seen are several species noted in the low, wet woods.

By the end of July, species in this habitat include Agrimonia rostellata (woodland agrimony agrimony (ăg`rĭmō'nē), any plant of the genus Agrimonia, perennials of the family Rosaceae (rose family) native to north temperate zones, to Brazil, and to Africa. They are found wild in the N and central United States. ), Chasmanthium latifolium (spangle span·gle  
n.
1. A small, often circular piece of sparkling metal or plastic sewn especially on garments for decoration.

2. A small sparkling object, drop, or spot: spangles of sunlight.
 grass), Helianthus Helianthus (hē'lēăn`thəs): see sunflower.  divaricatus (woodland sunflower), Lobelia inflata (Indian tobacco), Oenothera biennis (evening primrose), Polygonum virginianum (Virginia knotweed knotweed

polygonumaviculare.
), Silene stellata (starry campion), and Verbena verbena, common name for some members of the Verbenaceae, a family of herbs, shrubs, and trees (often climbing forms) of warmer regions of the world. Well-known wild and cultivated members of the family include species of the shrubby Lantana and of  urticifolia (white vervain vervain: see verbena.

vervain

indicates bewitching powers. [Flower Symbolism: Flora S ymbo lica, 178]

See : Enchantment
), to name the most conspicuous.

Through August, the selection of flowering species changes little. Many of the plants found in July continue to bloom. A notable addition is Hymenocallis caroliniana (spider lily), a showy species of southern swamplands that may reach its northern limit in Posey County.

High, dry woods.--(This habitat is wellshaded and well-drained without standing water.)

The main trail continues through low-lying woods for about 0.8 km, when it rises noticeably. The rise is not great, only about 1 m, but it creates an area of much drier soil. The plant community changes significantly. The secondary trail, which passed through the "low, dry woods" as it skirted the overcup oak slough, turns north and eventually also rises into the high, dry woods.

Throughout the growing season, the variety of flowering plants is greatest in this section of the preserve. Plants here tend to grow tall.

Through April, this is the area of the preserve where one is most likely to find plants in flower. The first to bloom are Cloytonia virginica (spring beauty), Dentaria laciniata (cut-leaved toothwort toothwort, any species of the genus Dentaria [Lat. dens=tooth, for the toothed rhizomes of some species], slender perennials of the family Cruciferae (mustard family), native to north temperate regions. ), and Thalictrum thalictroides. They will be followed by Corydalis flavula (yellow corydalis), Cynoglossum virginianum (wild comfrey or hound's-tongue), Geranium maculatum (wild geranium), and Uvularia sessilifolia (sessile-leaved bellwort). In mid-April, an area between the secondary trail and the cypress swamp supports a large number of Mertensia virginica (Virginia bluebells). Along the secondary trail, several specimens of Lithospermum latifolium (American gromwell) may be seen. Also notable as they bloom in the spring are the woody Lindera benzoin benzoin (bĕn`zoin, –zōĭn) or benzoinum (bĕnzoin`əm), balsamic resin, the dried exudation from the pierced bark of various species of the benzoin tree (Styrax  (spicebush spicebush: see laurel.
spicebush

Deciduous, dense shrub (Lindera benzoin, or Benzoin aestivale) of the laurel family, native to eastern North America. Found most often in damp woods, it grows 5–20 ft (1.5–6 m) tall.
) and Asimina triloba (pawpaw pawpaw: see custard-apple; papaya.  tree).

Several other species start to flower by mid-June. These include Arisaema dracontium, Arisaema triphyllum (Jack-in-the-pulpit), Blephilia hirsuta (hairy wood mint Noun 1. hairy wood mint - a variety of wood mint
Blephilia hirsuta

Blephilia, genus Blephilia - small genus of North American herbs: wood mints
), Desmodium glutinosum (pointed-leaf tick trefoil), Phytolacca americana (the abundant pokeweed), Podophyllum peltatum (mayapple mayapple (māˈ·aˑ·p ), and Stachys tenuifolia (smooth hedge-nettle).

At the end of July, a new selection of species is in flower. These plants, which grow 1-2 m tall, include Aralia racemosa (American spikenard spikenard (spīk`närd), name for several plants. The biblical spikenard, or nard, was a costly aromatic ointment, preserved in alabaster boxes, whose chief ingredient is believed to have been derived from Nardostachys grandiflora (or ), Campanula americana (tall bellflower), and Eupatorium purpureum (purple Joe-pye weed).

In August Eupatorium rugosum (white snakeroot white snakeroot, North American woods perennial (Eupatorium urticifolium) of the family Asteraceae (aster family), having a flat-topped cluster of small white flowers. It is of the same genus as the boneset and joe-pye weed. ), Helianthus microcephalus (small wood sunflower), Pilea pumila (clearweed clear·weed  
n.
Either of two eastern North American annual plants (Pilea pumila or P. fontana) having short drooping clusters of tiny greenish-white flowers and translucent stems and leaves. Also called richweed.
), Scrophularia marilandica (late figwort figwort, common name for some members of the Scrophulariaceae, a family comprising chiefly herbs and small shrubs and distributed widely over all continents. The family includes a few climbing types and some parasitic and saprophytic forms. ), Solidago caesia (blue-stemmed goldenrod goldenrod, any species of the large genus Solidago of the family Asteraceae (aster family), chiefly North American weedy herbs. They have small yellow flowers clustered, often in panicles, along a wandlike stem. ), and Verbesina alternifolia (yellow ironweed) are added.

By the end of September, most species at the preserve have gone to seed. A notable exception is the abundant Eupatorium rugosum, whose white flowers are conspicuous in the high, dry woods well into October.

Cypress swamp.--(This habitat is well-shaded, usually with standing water that varies from a few centimeters to an estimated depth of 0.5 m as judged by traces of a waterline on the cypress trees.)

The variety of flowering plants is much smaller in the cypress swamp than in the rest of the preserve. Only plants that can thrive in standing water are found in this habitat.

The dominant tree in the swamp is bald cypress tree (Taxodium distichum). According to the IDNR list (1993), Taxodium distichum is listed as threatened in Indiana. Another prominent tree is Populus heterophylla (swamp cottonwood), which in May drops cotton-like clusters of seeds that float on the dark waters of the swamp.

The only flowering species noted in April is Ranunculus Ranunculus

a very large plant genus of family Ranunculaceae; the buttercups. All of them should be regarded as potentially poisonous. The species listed below have been reported as causing poisoning in animals.
 flabellaris (yellow water buttercup). Also in April, the surface of the swamp is often covered with duckweed duckweed, any plant of the genus Lemna and sometimes of related genera. Duckweeds are tiny floating or submerged aquatic plants with reduced or obsolete roots. They flower only rarely, and their flowers are small and inconspicuous.  (Lemna sp.).

In May, the unusual Hottonia inflata (featherfoil) might be seen. This aquatic plant is sparse in some years and abundant in others. According to the IDNR list (1993), Hottonia inflata, is listed as a threatened species in Indiana.

Cephalanthus occidentalis (buttonbush but·ton·bush  
n.
A deciduous North American shrub (Cephalanthus occidentalis) having opposite leaves and spherical clusters of small white flowers.
) and Rosa palustris (swamp rose) come into bloom in June. At the edge of the swamp, in very shallow water, is a large stand of Saururus cernuus (lizard's tail).

In September, Bidens discoidea (swamp beggar-tick) comes into flower. The inconspicuous orange flowers may be seen in large numbers as summer ends.

APPENDIX 1

Catalog of Vascular Plants of the Twin Swamps Nature Preserve (arranged alphabetically by family)

DIVISION POLYPODIOPHYTA

Thelypteridaceae

Phegopteris hexagonoptera (Michx.) Fee: Broad beech fern Noun 1. broad beech fern - beech fern of North American woodlands having straw-colored stripes
Dryopteris hexagonoptera, Phegopteris hexagonoptera, southern beech fern, Thelypteris hexagonoptera
; high, dry woods; common, but in only a few locations; USIH 963.

DIVISION MAGNOLIOPHYTA

Acanthaceae (Acanthus Family)

Ruellia strepens L.: Smooth ruellia or wild petunia; May (blooming); low, dry woods; infrequent; narrow range; USIH 945.

Anacardiaceae (Cashew Family)

Toxicodendron radicans (L.) Kuntze: Common poison ivy; low, wet to high, dry woods; abundant in all habitats; USIH 914.

Apiaceae (Carrot Family)

Cicuta maculata L.: Water hemlock; July-August; low, wet woods; frequent; USIH 968.

Cryptotaenia canadensis (L.) DC.: Honewort hone·wort  
n.
An eastern North American perennial plant (Cryptotaenia canadensis) having umbels of small whitish flowers and compound leaves with larger, double serrate leaflets.
; June; low, dry to high woods; abundant in dry, shady woods; USIH 940.

Daucus carota L.: Queen Anne's lace Queen Anne's lace or wild carrot, herb (Daucus carota) of the family Umbelliferae (carrot family), native to the Old World but naturalized and often weedy throughout North America. ; July-August; fields; infrequent in open field; USIH 973.

Sanicula canadensis L.: Canadian black snakeroot; May; low, dry woods; infrequent; USIH 985.

Sanicula gregaria Bickn.: Common snakeroot snakeroot, name for several plants, among them black snakeroot (see bugbane), button snakeroot or blazing star, senega snakeroot (see milkwort), and white snakeroot. ; May; low, dry woods; frequent; USIH 948.

Apocynaceae (Dogbane Family)

Apocynum cannabinum L.: Indian hemp; June; field; common; USIH 961.

Araceae (Arum Family)

Arisaema dracontium (L.) Schott.: Green dragon; May; low wet; low dry; high dry woods; frequency varies--hard to find some years, frequent in others; thrives in poorly-drained soil; USIH 929.

Arisaema triphyllum (L.) Schott.: Jack-in-the-pulpit; May; high, dry woods; infrequent; prefers well-drained soil; USIH 954.

Araliaceae (Ginseng Family)

Aralia racemosa L.: American spikenard; July; high, dry woods; infrequent and sparse; USIH 986.

Asclepiadaceae (Milkweed Family)

Asclepias incarnata L.: Swamp milkweed; August; field; infrequent; in poorly-drained area of field; USIH 988.

Asclepias purpurascens L.: Purple milkweed; May-June; low, dry woods; infrequent, but conspicuous; limited habitat; USIH 938.

Asteraceae (Aster Family)

Achillea millefolium L.: Common yarrow: May; field; infrequent; USIH 994.

Ambrosia artemisiifolia L.: Common ragweed ragweed, any plant of the genus Ambrosia, coarse, weedy herbs belonging to the family Asteraceae (aster family), most of which are native to America. They have inconspicuous greenish flowers and soft subdivided leaves. ; August; field; abundant; USIH 975.

Aster ericoides L.: Heath aster; August-September; field; frequent; USIH 974.

Aster pilosus Willd.: Frost-weed aster; September; low, wet woods; infrequent; USIH 976.

Bidens aristosa (Michx.) Britt.: Western tickseed sunflower; August; field; infrequent, though very conspicuous; USIH 989.

Bidens discoidea (Torr. & Gray) Britt.: Swamp beggar-tick; August-September; cypress swamp; abundant; USIH 990.

Boltonia asteroides (L.) L'Her.: False aster; August; field; sparse; USIH 927.

Cirsium discolor dis·col·or  
v. dis·col·ored, dis·col·or·ing, dis·col·ors

v.tr.
To alter or spoil the color of; stain.

v.intr.
To become altered or spoiled in color.
 (Muhl.) Spreng.: Field thistle; August-September; field; frequent: USIH 1018.

Conyza canadensis (L.) Cronq.: Horseweed; August; field; frequent; USIH 1019.

Erigeron annuus (L.) Pers.: Daisy fleabane fleabane, any plant of the genus Erigeron, widely distributed herbs of the family Asteraceae (aster family), especially abundant in temperate and mountainous regions of North America. ; June-August; field; frequent: USIH 1020.

Eupatorium coelestinum L.: Mistflower mist·flow·er  
n.
A perennial plant (Eupatorium coelestinum) of the southeast and central United States, having corymbs of small blue flowers.

Noun 1.
; August; low, wet woods; sparse; USIH 971.

Eupatorium perfoliatum L.: True boneset boneset (bōn`sĕt) or thoroughwort (thûr`əwûrt', –wôrt'), perennial North American herb (Eupatorium perfoliatum ; August; field; infrequent; USIH 1027.

Eupatorium purpureum L.: Purple Joe-pye weed; July-August; high, dry woods; infrequent; USIH 993.

Eupatorium rugosum Houtt.: White snakeroot; August-September: high, dry woods; abundant; conspicuous; USIH 987.

Eupatorium serotinum Michx.: Late or false boneset; August; field; infrequent; USIH 1024.

Euthamia graminifolia (L.) Salisb.: Grass-leaved goldenrod; August; low, moist areas of field: abundant; USIH 1025.

Gnaphalium obtusifolium L.: Old-field balsam balsam (bôl`səm), fragrant resin obtained from various trees. The true balsams are semisolid and insoluble in water, but they are soluble in alcohol and partly so in hydrocarbons.  (rabbit-tobacco); August; field; infrequent; USIH 1023.

Helianthus divaricatus L.: Woodland sunflower; July-August; low, dry woods; frequent and conspicuous; USIH 1006.

Helianthus microcephalus Torr. & Gray: Small wood sunflower; August; high, dry woods; sparse; USIH 1007.

Helianthus sp.: Species unknown; July-August; field; infrequent; USIH 1008.

Krigia biflora (Walt.) Blake: Dwarf or false dandelion; May; low, wet woods; infrequent, conspicuous; USIH 932.

Lactuca biennis (Moench) Fern.: Tall blue lettuce; August-September; low, dry woods; infrequent; USIH 1022.

Prenanthes altissima L.: Rattlesnake root or tall white lettuce; August-September; high, dry woods; sparse; USIH 970.

Senecio glabellus (Poir.): Butterweed but·ter·weed  
n.
1. A succulent annual or biennial plant (Senecio glabellus), native to the eastern United States and having pinnately divided leaves and bright yellow, radiate flower heads.

2. The horseweed.
; April-May; low, dry woods; common; USIH 946.

Solidago caesia L.: Blue-stemmed or woodland goldenrod; September-October; high, dry woods; infrequent; USIH 1002.

Solidago canadensis L.: Tall goldenrod; August-September; field; abundant; USIH 1003.

Solidago juncea Ait.: Early goldenrod; July; field; abundant; USIH 1004.

Solidago nemoralis Ait.: Gray goldenrod; August-September; field; frequent; USIH 1005.

Taraxacum officinale Weber: Common dandelion; field; frequent; USIH 1021.

Verbesina alternifolia (L.) Britt.: Yellow ironweed or wingstem; August-September; high, dry woods; common; USIH 977.

Verbesina helianthoides Michx.: Yellow crownbeard; June; low, dry woods; infrequent; USIH 947.

Vernonia fasciculata Michx.: Ironweed; August; low, wet woods; infrequent; USIH 965.

Balsaminaceae (Touch-Me-Not Family)

Impatiens capensis Meerb.: Spotted (orange) jewelweed; June-August; low, wet to low, dry woods; abundant; USIH 931.

Berberidaceae (Barberry Family)

Podophyllum peltatum L.: Mayapple; April-May; high, dry woods; abundant; USIH 950.

Bignoniaceae (Trumpet Creeper Family)

Campsis radicans (L.) Seem.: Trumpet creeper; June-July; field; sparse, although abundant along nearby roads; USIH 962.

Boraginaceae (Borage Family)

Cynoglossum virginianum L.: Hound's tongue or wild comfrey; April; high, dry woods; infrequent; USIH 921.

Hackelia virginiana L.: Stickseed; July; low, wet to high, dry woods; abundant; USIH 964.

Lithospermum latifolium Michx.: American gromwell; April-May; high, dry woods; Infrequent; USIH 925.

Mertensia virginica (L.) Pers.: Virginia bluebell; April; high, dry woods; abundant in certain locations; USIH 918.

Myosotis verua Nutt.: Scorpion grass; May; field; infrequent; USIH 1028.

Brassicaceae (Mustard Family)

Alliaria petiolata (Bieb.) Cavara & Grande: Garlic mustard; April; field; frequent; USIH 1032.

Cardamine bulbosa (Schreb.) BSR: Spring cress; April; low, wet woods; frequent; USIH 915.

Dentaria laciniata Muhl: Cut-leaved toothwort; March-April; high, dry woods; frequent; USIH 922.

Caesalpiniaceae (Caesalpinia Family)

Cassia cassia (kăsh`ə): see cinnamon; senna.
cassia

Spice, also called Chinese cinnamon, consisting of the aromatic bark of the Cinnamomum cassia plant, of the laurel family.
 fasciculate fas·cic·u·late   also fas·cic·u·lat·ed
adj.
Arranged in or formed of fascicles; fascicular.



fas·cicu·late·ly adv.
 Michx.: Partridge pea; July-August; field; frequent; USIH 1031.

Campanulaceae (Bellflower Family)

Campanula americana L.: Tall bellflower; July-August; high, dry woods; infrequent; USIH 967.

Triodanis perfoliata (L.) Nieuwl: Venus's looking-glass; May June; field; infrequent; USIH 966.

Caprifoliaceae (Honeysuckle Family)

Lonicera japonica Thunb.: Japanese honeysuckle; May; field; abundant at edge of woods; USIH 1029.

Caryophyllaceae (Pink Family)

Dianthus armeria L.: Deptford pink; June; field; infrequent; USIH 991.

Silene stellota (L.) Ait.: Starry campion; June-August; low, dry woods; infrequent; USIH 992.

Convolvulaceae (Morning Glory Family)

Ipomoea Ipomoea

widespread genus of poisonous vines of the family Convolvulaceae; may contain various toxins including the indole alkaloid lysergic acid, furanoterpenes, indolizidine alkaloids (swainsonine). Includes I. asarifolia (salsa), I.
 pandurata (L.) G.F.W. Mey: Wild potato vine; July-August; field; frequent and conspicuous; USIH 1034.

Cuscutaceae (Dodder dodder: see morning glory.
dodder

Any of the leafless, twining, parasitic vines (see parasitism) that make up the genus Cuscuta (family Cuscutaceae), containing more than 150 species found throughout temperate and tropical regions.
 Family)

Cuseuta gronovii Willd.: Dodder; August-September; field; infrequent; USIH 1041.

Dioscoreaceae (Yam Family)

Dioscorea villosa L.: Wild yam; June; high, dry woods; sparse; USIH 1043.

Euphorbiaceae (Spurge Family)

Euphorbia corollata L.: Flowering spurge; August; low, wet woods; sparse; USIH 1045.

Fabaceae (Pea Family)

Desmodium glabellum (Michx.) DC.: Tick trefoil; August; low, wet woods; frequent; USIH 1036.

Desmodium glutinosum (Muhl.) Wood: Pointed tick trefoil; June-July; high, dry woods; Infrequent; USIH 949. Desmodium paniculatum (L.) DC.: Panicled tick trefoil; August; field and low, wet woods; infrequent; USIH 1035.

Lespedeza lespedeza (lĕs'pədē`zə) or bush clover, any plant of the genus Lespedeza, leguminous herbs or undershrubs of the family Leguminosae (pulse family); native to North America, Asia, and Australia.  virginica (L.) Britt.: Slender bush clover; August; field; frequent and conspicuous; USIH 1030.

Trillium pratense L.: Red clover; June; field; infrequent; USIH 995.

Trifolium repens L.: White clover; July-August; field; frequent; USIH 996.

Geraniaceae (Geranium Family)

Geranium carolinianum L.: Wild cranesbill; May; field; infrequent; USIH 997.

Geranium maculatum L Geranium maculatum L,
n See tormentil.
.: Wild geranium; April; high, dry woods; infrequent; USIH 920.

Hypericaceae (St. John's Wart Family)

Hypericum Hypericum /Hy·per·i·cum/ (hi-per´i-kum) a genus of herbs, including several types of St. John's wort.

Hypericum perfora´tum  the species of St.
 punctatum Lam.: Spotted St. John's wart; July; field; infrequent; USIH 1037.

Lamiaceae (Mint Family)

Blephilio hirsuta (Pursh) Benth.: Hairy wood-mint; June-August; high, dry woods; frequent: USIH 953.

Lamium sp.: Dead nettle; March-April; field; USIH 1009.

Lycopus rubellus Moench.: Stalked water horehound; August; field; infrequent in poorly-drained areas of field; USIH 1010.

Prunella pru·nel·la   also pru·nelle
n.
A strong heavy fabric of worsted twill, used chiefly for shoe uppers, clerical robes, and academic gowns.
 vulgaris L.: Self-heal or heal-all; July-August; field to low, dry woods; frequent; USIH 1033.

Pycnanthemum tenuifolium Schrad.: Slender mountain mint; June-July; low, wet to low, dry woods; frequent; USIH 936.

Stachys tenuifolia Willd.: Smooth hedge-nettle; June-August; high, dry woods; frequent; USIH 952.

Lauraceae (Laurel Family)

Lindera benzoin (L.) Blume: Spicebush; March-April; high, dry woods; frequent; USIH 923.

Lemnaceae (Duckweed Family)

Lemna sp.: Duckweed; cypress swamp, overcup oak slough; abundant; USIH 956.

Liliaceae (Lily Family)

Hymenocallis caroliniana (L.) Herb.: Spider-lily; August; low, dry woods; sparse; USIH 979.

Smilacina racemosa (L.) Desf.: False Solomon's seal; April-May; low, dry to high, dry woods; infrequent; USIH 1038.

Uvularia sessilifolia L.: Sessile-leaved bellwort; April; high, dry woods; infrequent; USIH 926.

Lobeliaceae (Lobelia Family)

Lobelia cardinalis L.: Cardinal flower; August; low, wet woods; infrequent; USIH 1012.

Lobelia inflata L.: Indian tobacco; July; low, dry woods; infrequent; USIH 1011.

Onagraceae (Evening Primrose Family)

Circaea lutetiana Aschers. & Magnus: Enchanter's nightshade; May-June; high, dry woods; infrequent; USIH 978.

Ludwigia alternifolia L.: Seedbox; July; low, wet woods; infrequent; USIH 1040.

Oenothera biennis L.: Evening primrose; June-August; low, dry woods and field; infrequent; USIH 1039.

Oxalidaceae (Wood Sorrel Family)

Oxalis dillenii Jacq.: Yellow wood sorrel; May-August; field to high, dry woods; abundant; USIH 933.

Papaveraceae (Poppy Family)

Corydalis flavula (Raf.) DC.: Yellow corydalis; April; high, dry woods; frequent; USIH 924.

Passifloraceae (Passion-Flower Family)

Passiflora lutea L.: Yellow passionflower passionflower, any plant of the genus Passiflora, mostly tropical American vines having pulpy fruits. Some species are grown in greenhouses for their large, unusual flowers of various colors; those seen by early Spanish settlers were interpreted as symbolic of ; July; field; sparse; USIH 1042.

Phrymaceae (Lopseed Family)

Phryma leptostachya L.: Lopseed; June-July; low, dry to high, dry woods; abundant; USIH 943.

Phytolaccaceae (Pokeweed Family)

Phytolacca americana L.: Pokeweed; June-July; high, dry woods; abundant; USIH 1017.

Plantaginaceae (Plantain plantain (plăn`tĭn), any plant of the genus Plantago, chiefly annual or perennial weeds of wide distribution. Many species are lawn pests and the pollen is often a hay fever irritant. P.  Family)

Plantago major L.: Common plantain; July; field; abundant; USIH 1044.

Poaceae (Grass Family)

Agrostis sp.: August; low, wet woods; frequent; USIH 1026.

Andropogon virginicus L.: Broom sedge; August; field; abundant; USIH 1056.

Chasmanthium latifolium (Michx.) Yates: Spangle grass or sea oats; June-October; low, dry woods; infrequent; USIH 939.

Leersia lenticularis Michx.: Catchfly catchfly: see pink.  grass; August; low, wet woods; infrequent; USIH 1046.

Setaria glauca (L.) Beauv.: Foxtail; August-September; field to low, dry woods; abundant; USIH 981.

Polemoniaceae (Phlox Family)

Phlox glaberrima L.: Smooth phlox; May-August; low, wet woods; abundant; USIH 934.

Polygonaceae (Smartweed smartweed: see buckwheat.  Family)

Polygonum hydropiperoides Michx.: Mild water pepper; August-September; low, wet woods; infrequent; USIH 1047.

Polygonum pensylvanicum L.: Pennsylvania smartweed; August; field (wet areas); infrequent; USIH 980.

Polygonum virginianum L.: Virginia knotweed; July-August; low, dry to high, dry woods; abundant; USIH 1049.

Portulacaceae (Purslane Family)

Claytonia virginica L.: Spring beauty; March-April; high, dry woods; abundant; USIH 919.

Primulaceae (Primrose Family)

Hottonia inflata Ell.: Featherfoil; May; cypress swamp and overcup oak slough; abundant some years, sparse in others; USIH 1058.

Ranunculaceae (Buttercup Family)

Ranunculus flabellaris Raf.: Yellow water buttercup; April; cypress swamp; infrequent; USIH 927.

Ranunculus septentrionalis Poir.: Swamp buttercup; April; low, wet woods; infrequent; USIH 913.

Thalictrum thalictroides (L.) Eaves & Boivin: Rue anemone; April; high, dry woods; frequent; USIH 916.

Rosaceae (Rose Family)

Agrimonia rostellata Wallr.: Woodland agrimony; July-August; low, dry woods; infrequent; USIH 1013.

Geum canadense Jacq.: White avens avens

Any of the low-growing, perennial flowering plants (approximately 50 species) of the genus Geum, in the rose family. Most occur in the northern or southern temperate zones or in the Arctic.
; June-August; low, dry woods; infrequent; USIH 942.

Porteranthus stipulatus (Muhl.) Britt.: Indian physic or American ipecac; May-June; low, dry woods; infrequent; USIH 944.

Potentilla simplex Michx.: Common cinquefoil; May; low, wet woods; infrequent; USIH 935.

Rosa multiflora Thunb.: Multiflora rose; May; field; frequent; USIH 983.

Rosa palustris Marsh.: Swamp rose; June; cypress swamp; sparse; USIH 958.

Rosa setigera Michx.: Prairie rose; June; field; infrequent; USIH 984.

Rubus sp.: Blackberry; May; field; abundant; USIH 982.

Rubiaceae (Madder Family)

Cephalanthus occidentalis L.: Buttonbush; June; cypress swamp; abundant; USIH 955.

Galium oparine L.: Cleavers; May-June; low, wet to low, dry woods; frequent; USIH 998.

Galium triflorum Michx.: Sweet-scented bedstraw bedstraw: see madder.
bedstraw

Any low perennial herbaceous plant of the genus Galium, in the madder family, found in damp woods and swamps and along stream banks and shores.
; June; low, wet woods; infrequent; USIH 1015.

Galium sp.: Bedstraw; May-June; low, dry woods; frequent; USIH 941.

Hedyotis purpurea var. calycosum (Gray) Fosberg:Houstonia; May; low, wet woods; frequent; USIH 930.

Saxifragaceae (Saxifrage saxifrage (săk`sĭfrĭj), common name for several members of the Saxifragaceae, a family of widely varying herbs, shrubs, and small trees of cosmopolitan distribution.  Family)

Penthorum sedoides L.: Ditch stonecrop stonecrop, common name for members of the Crassulaceae (also called orpine, or hen-and-chickens, family), a family of succulent, fleshy herbs and shrubs mostly inhabiting arid regions in many parts of the world. ; August; low, wet woods; sparse; USIH 1001.

Saururaceae (Lizard's-Tail Family)

Saururus cernuus L.: Lizard's tail; June; cypress swamp; abundant; USIH 959.

Scrophulariaceae (Figwort Family)

Mimulus alotus Ait.: Monkey flower; August; low, wet woods; infrequent; USIH 1050.

Penstemon alluviorum Pennell: Lowland beardstongue; May; low, wet woods; frequent; USIH 937.

Scrophularia marilandica L.: Maryland figwort; July-August; high, dry woods; infrequent; USIH 1048.

Solanaceae (Nightshade Family)

Solanum Solanum

a widespread plant genus of the family Solanaceae which contains a number of valuable crop plants but also some poisonous ones. Poisoning may be due to (1) the presence in the plant of toxic glycoalkaloids which cause diarrhea, (2) alkamines, e.g.
 carolinense L.: Horse nettle; July; field; infrequent; USIH 1016.

Urticaceae (Nettle Family)

Boehmeria cylindrica (L.) Sw.: False nettle; July-August; low, wet to low, dry woods; frequent; USIH 1014.

Laportea canadensis (L.) Wedd.: Wood nettle; July-August; low, wet to low, dry woods; infrequent; USIH 1053.

Pilea pumila (L.) Gray: Clearweed: August; high, dry woods; abundant; USIH 1054.

Verbenaceae (Vervain Family)

Verbena urticifolia L.: White vervain; June-August; low wet to high, dry woods; abundant; USIH 1052.

Violaceae (Violet Family)

Viola eriocarpa (Schwein.) Russell: Smooth yellow violet; April; plentiful in high, dry woods; USIH 917.

Viola papilionacea Pursh: Common violet; April; low, wet to low, dry woods; abundant; USIH 912.

Vitaceae (Grape Family)

Parthenocissus quinquefolia (L.) Planch.: Virginia creeper; throughout woods; abundant; USIH 1051.

LITERATURE CITED

Abrell, D.B. 1997. A taste of the south: The southern bottomlands natural region. 582 pp. In The Natural Heritage of Indiana. (M.T. Jackson, ed.) Indiana University Press Indiana University Press, also known as IU Press, is a publishing house at Indiana University that engages in academic publishing, specializing in the humanities and social sciences. It was founded in 1950. Its headquarters are located in Bloomington, Indiana. . Bloomington, Indiana.

Deam, C. 1940. Flora of Indiana. Dept. of Conserv., Wm. B. Burford Printing Co. Indianapolis, Indiana. 1236 pp.

Gleason, H.A. & A. Cronquist. 1991. Manual of the Vascular Plants of Northeastern United States and Adjacent Canada. New York Botanical Garden For the botanical garden in Queens, see .
The New York Botanical Garden is a prestigious botanical garden in New York City. One of the premier botanical gardens in the United States, it spans some 240 acres of Bronx Park in the borough of The Bronx and is home to some of the
. Bronx, New York. 910 pp.

Homoya, M.A. 1983. Additions to the flora of southern Indiana. Proceedings of the Indiana Academy of Science 92:379-382.

Homoya, M.A. 1987. Additions to the flora of southern Indiana III. Proceedings of the Indiana Academy of Science 97:449-452.

Homoya, M.A., D.B. Abrell, & J.R. Aldrich, & T.W. Post. 1985. The natural regions of Indiana. Proceedings of the Indiana Academy of Science 94:245-268.

Homoya, M.A. & D.B. Abrell. 1986. Recent additions to the flora of southern Indiana. Proceedings of the Indiana Academy of Science 95:429-432.

Homoya, M.A. & C.L. Hedge. 1990. Additions to the flora of southern Indiana. Proceedings of the Indiana Academy of Science 99:67-72.

Indiana Department of Natural Resources. 1993. Indiana's rare plants and animals Plants and Animals are a Canadian indie-rock band from Montreal, comprised of guitarist-vocalists Warren Spicer and Nic Basque, and drummer-vocalist Matthew Woodley.[1] They are signed to Secret City Records. : A checklist of endangered and threatened species. Indiana Department of Natural Resources. Indianapolis, Indiana. 26 pp.

Lindsey, A.A. 1962. Analysis of an original forest of the lower Wabash floodplain and upland. Proceedings of the Indiana Academy of Science 72: 282-287.

Mohlenbrock, R.H. 1986. Guide to the Vascular Flora of Illinois. Southern Illinois Univ. Press. Carbondale, Illinois. 507 pp.

Mohlenbrock, R.H. & J.W. Voigt. 1959. A Flora of Southern Illinois. Southern Illinois Univ. Press. Carbondale, Illinois. 390 pp.

Ridgway, R. 1872. Notes on the vegetation of the Lower Wabash Valley. American Naturalist 6: 658-665.

Venning, ED. 1984. A Guide to Field Identification: Wildflowers of North America. Golden Press. New York, New York. 340 pp.

Walters, D.R. & D.J. Keil. 1996. Vascular Plant Taxonomy. 4th ed. Kendall/Hunt Publishing Co.. Dubuque, Iowa. 608 pp.

Wharton, M. E. 1971. Guide to the Wildflowers and Ferns of Kentucky. Univ. of Kentucky Press. Lexington, Kentucky. 328 pp.

Wharton, M.E. 1973. Trees and Shrubs of Kentucky. Univ. of Kentucky Press. Lexington, Kentucky. 582 pp.

Manuscript received 19 August 2003, revised 3 October 2003.

Rick Mark and Scott A. Gordon: Biology Department, University of Southern Indiana, Evansville, Indiana 47712 USA
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